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<document id="17C22DEF3357F9FE5A862ED6769E88BA" ID-CLB-Dataset="4740" ID-DOI="10.11646/zootaxa.4500.2.3" ID-GBIF-Dataset="7392ae8e-f0ef-4ad2-af7a-03e3a3f32bdb" ID-ISSN="1175-5326" ID-Zenodo-Dep="5297537" ID-ZooBank="4E758BE2-90DA-471A-86E6-FF5D318FE98C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630083675860" checkinUser="felipe" docAuthor="Hrivniak, Ľuboš, Sroka, Pavel, Godunko, Roman J., Palatov, Dmitry, Polášek, Marek, Manko, Peter &amp; Oboňa, Jozef" docDate="2018" docId="6103A142FF966B24FF26FEFBFE3B067B" docLanguage="en" docName="zootaxa.4500.2.3.pdf" docOrigin="Zootaxa 4500 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ecdyonurus (Ecdyonurus) eurycephalus Hrivniak &amp; Sroka &amp; Godunko &amp; Palatov &amp; Polášek &amp; Manko &amp; Oboňa 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="198" masterDocId="9D3AD93AFF956B27FFB1FF9CFFE50208" masterDocTitle="Diversity of Armenian mayflies (Ephemeroptera) with the description of a new species of the genus Ecdyonurus (Heptageniidae)" masterLastPageNumber="221" masterPageNumber="195" pageNumber="198" updateTime="1699146250564" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="29BAFA09D3458BBE2F04A1FDF6CC23A7">Diversity of Armenian mayflies (Ephemeroptera) with the description of a new species of the genus Ecdyonurus (Heptageniidae)</mods:title>
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<mods:namePart id="616595808433D1B21FD7F00962FD37A7">Hrivniak, Ľuboš</mods:namePart>
<mods:affiliation id="9D0A308F035EF201BDFAA427C75BCFC1">The Czech Academy of Sciences, Biology Centre, Institute of Entomology, Branišovská 31, CZ- 37005 České Budějovice, Czech Republic. E-mails: lubos. hrivniak @ gmail. com, pavel. sroka @ centrum. cz &amp; Faculty of Sciences, University of South Bohemia, Branišovská 31, CZ- 37005 České Budějovice, Czech Republic</mods:affiliation>
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<mods:namePart id="34D50F6AB72DC2891B9A1B442E5CF7A2">Sroka, Pavel</mods:namePart>
<mods:affiliation id="ABB4DD3D9A5F560AB0C0F2DBB695C83E">The Czech Academy of Sciences, Biology Centre, Institute of Entomology, Branišovská 31, CZ- 37005 České Budějovice, Czech Republic. E-mails: lubos. hrivniak @ gmail. com, pavel. sroka @ centrum. cz</mods:affiliation>
</mods:name>
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<mods:roleTerm id="87376A1699C28B0873FE99D26024447E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7F6F2C0DE084971A8D5F1C377F64F2B9">Godunko, Roman J.</mods:namePart>
<mods:affiliation id="922F52EDD6E667EBAEB8460454BFBB56">The Czech Academy of Sciences, Biology Centre, Institute of Entomology, Branišovská 31, CZ- 37005 České Budějovice, Czech Republic. E-mails: lubos. hrivniak @ gmail. com, pavel. sroka @ centrum. cz &amp; State Museum of Natural History, National Academy of Sciences of Ukraine, Teatralna 18, UA- 79008 Lviv, Ukraine.</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="7DC7D85D40864DC361FE15E633E4B249">Palatov, Dmitry</mods:namePart>
<mods:affiliation id="FC1CB5EDA7406112156159AB4CC092B9">Department of Hydrobiology, Biological Faculty, Moscow State University, Leninskie Gory, 1 / 12, RU- 119234 Moscow, Russian Federation and A. N. Severtzov Institute of Ecology and Evolution, Leninsky prospect, 33, RU- 117071 Moscow, Russian Federation. E-mail: triops @ yandex. ru</mods:affiliation>
</mods:name>
<mods:name id="D78C6A8903DEA1F33E2CBDAF139FD257" type="personal">
<mods:role id="264265B1CD43E6FD125B23D76144EC24">
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</mods:role>
<mods:namePart id="22228ED5B70C099C1C7357083034D8F5">Polášek, Marek</mods:namePart>
<mods:affiliation id="939207233FC39996857B71E0422888EC">Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlářská 2, CZ- 61137 Brno, Czech Republic.</mods:affiliation>
</mods:name>
<mods:name id="44B96E77C5D9A6AB195A79442932789F" type="personal">
<mods:role id="00BCF3F81E0B198FB35C26850ACFFFA5">
<mods:roleTerm id="E9520F4BCD433E03E978B062EA32A0C4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB69CEBA105E62BC54EE210BB197A50F">Manko, Peter</mods:namePart>
<mods:affiliation id="2F50361293E58AA19BA705D7B3987076">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Prešov, 17. novembra 1, SK- 08116 Prešov, Slovakia. E-mails: peter. manko @ unipo. sk; obonaj @ centrum. sk</mods:affiliation>
<mods:nameIdentifier id="45A636B8C71640AD463FC74CAB71243E" type="email">peter.manko@unipo.sk</mods:nameIdentifier>
</mods:name>
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</mods:role>
<mods:namePart id="054F3E46685F8AEED03B42366B031E69">Oboňa, Jozef</mods:namePart>
<mods:affiliation id="4977A03BA6F322E28C7B4535CBC10C8F">Department of Ecology, Faculty of Humanities and Natural Sciences, University of Prešov, 17. novembra 1, SK- 08116 Prešov, Slovakia. E-mails: peter. manko @ unipo. sk; obonaj @ centrum. sk</mods:affiliation>
<mods:nameIdentifier id="8D90588C2280BAB18CC394E5CE3D7F03" type="email">peter.manko@unipo.sk</mods:nameIdentifier>
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<mods:title id="16F1A8317D7D9DF23805FD5F824DE14B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="3220CDD6769D5F5BC33DAB5E85B18903">
<mods:date id="BA1AD75E354F41AA5F1A48C70F410103">2018</mods:date>
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<mods:number id="075E472D45520ABCBB7E462DC9674F71">2018-10-16</mods:number>
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<mods:number id="69415E2B5F6C9661111884FA6684C7D7">4500</mods:number>
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<mods:number id="D22121F65F42DEFC153AB16EFB079B85">2</mods:number>
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<mods:classification id="7214EF5D7EF5199E5D839A0815CE08AF">journal article</mods:classification>
<mods:identifier id="1837542C0CFE3E4082ABF88BE9A5A547" type="CLB-Dataset">4740</mods:identifier>
<mods:identifier id="4881B3606AD81B945B97C072500D1738" type="DOI">10.11646/zootaxa.4500.2.3</mods:identifier>
<mods:identifier id="5B4FE14E31B543DF7ACC4D45C784EFFE" type="GBIF-Dataset">7392ae8e-f0ef-4ad2-af7a-03e3a3f32bdb</mods:identifier>
<mods:identifier id="7DAFB4D3C292ADA2D188BA08C337FD5A" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
<treatment id="6103A142FF966B24FF26FEFBFE3B067B" ID-DOI="http://doi.org/10.5281/zenodo.5304125" ID-GBIF-Taxon="185118369" ID-Zenodo-Dep="5304125" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B" httpUri="http://treatment.plazi.org/id/6103A142FF966B24FF26FEFBFE3B067B" lastPageNumber="198" pageId="3" pageNumber="198">
<subSubSection id="A1B043DFFF966B24FF26FEFBFBE4038A" box="[151,1025,359,386]" pageId="3" pageNumber="198" type="nomenclature">
<paragraph id="E9151054FF966B24FF26FEFBFBE4038A" blockId="3.[151,1025,359,419]" box="[151,1025,359,386]" pageId="3" pageNumber="198">
<heading id="B25DA738FF966B24FF26FEFBFBE4038A" bold="true" box="[151,1025,359,386]" fontSize="11" level="1" pageId="3" pageNumber="198" reason="1">
<emphasis id="DBDECC46FF966B24FF26FEFBFBE4038A" bold="true" box="[151,1025,359,386]" pageId="3" pageNumber="198">
<taxonomicName id="2EAA6BD7FF966B24FF26FEFBFC73038A" authority="Hrivniak &amp; Godunko" authorityName="Hrivniak &amp; Sroka &amp; Godunko &amp; Palatov &amp; Polášek &amp; Manko &amp; Oboňa" authorityYear="2018" box="[151,918,359,386]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="198" phylum="Arthropoda" rank="species" species="eurycephalus" status="sp. nov." subGenus="Ecdyonurus">
<emphasis id="DBDECC46FF966B24FF26FEFBFECC0389" bold="true" box="[151,297,359,385]" italics="true" pageId="3" pageNumber="198">Ecdyonurus</emphasis>
(
<emphasis id="DBDECC46FF966B24FE8BFEFBFE280389" bold="true" box="[314,461,359,385]" italics="true" pageId="3" pageNumber="198">Ecdyonurus</emphasis>
)
<emphasis id="DBDECC46FF966B24FE6FFEFBFD9A0389" bold="true" box="[478,639,359,385]" italics="true" pageId="3" pageNumber="198">eurycephalus</emphasis>
Hrivniak &amp; Godunko
</taxonomicName>
,
<taxonomicNameLabel id="C0ED713DFF966B24FC15FEF4FBE4038A" box="[932,1025,360,386]" pageId="3" pageNumber="198" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="A1B043DFFF966B24FF26FE16FEFC03AB" box="[151,281,394,419]" pageId="3" pageNumber="198" type="description">
<paragraph id="E9151054FF966B24FF26FE16FEFC03AB" blockId="3.[151,1025,359,419]" box="[151,281,394,419]" pageId="3" pageNumber="198">
(
<figureCitation id="71910CD1FF966B24FF2EFE16FEF503AB" box="[159,272,394,419]" captionStart-0="FIGURES 2–12" captionStart-1="FIGURES 13–14" captionStart-2="FIGURES 15–23" captionStart-3="FIGURES 24–26" captionStart-4="FIGURES 27–28" captionStart-5="FIGURES 29–30" captionStart-6="FIGURES 31–45" captionStartId-0="4.[151,264,1049,1072]" captionStartId-1="4.[151,264,1968,1991]" captionStartId-2="5.[151,264,1612,1635]" captionStartId-3="6.[151,264,749,772]" captionStartId-4="6.[151,264,1410,1433]" captionStartId-5="7.[151,264,725,748]" captionStartId-6="7.[151,264,1798,1821]" captionTargetBox-0="[162,1428,307,1025]" captionTargetBox-1="[154,1435,1184,1942]" captionTargetBox-2="[157,1430,952,1589]" captionTargetBox-3="[154,1435,412,727]" captionTargetBox-4="[154,1422,860,1384]" captionTargetBox-5="[151,1436,193,703]" captionTargetBox-6="[154,1434,831,1774]" captionTargetId-0="figure-139@4.[156,1432,301,1028]" captionTargetId-1="figure-148@4.[151,1436,1182,1946]" captionTargetId-2="figure-468@5.[155,1432,949,1592]" captionTargetId-3="figure-414@6.[150,1436,412,728]" captionTargetId-4="figure-423@6.[151,1436,852,1389]" captionTargetId-5="figure-139@7.[151,1436,193,704]" captionTargetId-6="figure-148@7.[151,1436,828,1777]" captionTargetPageId-0="4" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="6" captionTargetPageId-5="7" captionTargetPageId-6="7" captionText-0="FIGURES 2–12. Ecdyonurus (Ecdyonurus) spp., male imagoes: 2–5, 8–12, E. (E.) eurycephalus sp. nov.: 2, 4, head lateral; 3, 5, head dorsal; 8, abdomen lateral; 9, abdomen ventral; 10, abdomen dorsal; 11, forewing; 12, hind wing; 6–7, E. (E.) ornatipennis: 6, head lateral; 7, head dorsal." captionText-1="FIGURES 13–14. Ecdyonurus (Ecdyonurus) eurycephalus sp. nov., genitalia of male imago: 13, styliger, forceps and penis ventral; 14, penis dorsal." captionText-2="FIGURES 15–23. Ecdyonurus (Ecdyonurus) eurycephalus sp. nov., larva: 15, habitus dorsal; 16, habitus ventral; 17, habitus lateral; 18, head dorsal; 19–21, semipronotum of mature larva; 22–23, semipronotum of early-instar larva." captionText-3="FIGURES 24–26. Ecdyonurus (Ecdyonurus) eurycephalus sp. nov., mouthparts: 24, labrum dorsal (left) and ventral (right); 25, labrum contour; 26, labial glossae and paraglossae: left part ventral view, right part contour." captionText-4="FIGURES 27–28. Ecdyonurus (Ecdyonurus) eurycephalus sp. nov., leg: 27, hind leg dorsal; 28a–h, setae on dorsal surface of femora." captionText-5="FIGURES 29–30. Ecdyonurus (Ecdyonurus) eurycephalus sp. nov., larval abdomen and leg: 29, hind margin of middle femora; 30, posterior margin of tergum III." captionText-6="FIGURES 31–45. Ecdyonurus (Ecdyonurus) eurycephalus sp. nov., gills (flattened on slide) and larval abdomen: 31–36, gill plate I: 31–34 mature larva; 35–36, early-instar larva; 37–39, gill plate IV: 37–38, mature larva; 39, early instar larva; 40–43, gill plate VII: 40–42, mature larva; 43, early-instar larva; 44, abdominal sterna V–IX (male); 45, sternum IX (female)." figureDoi-0="http://doi.org/10.5281/zenodo.5297541" figureDoi-1="http://doi.org/10.5281/zenodo.5297543" figureDoi-2="http://doi.org/10.5281/zenodo.5297545" figureDoi-3="http://doi.org/10.5281/zenodo.5297547" figureDoi-4="http://doi.org/10.5281/zenodo.5297549" figureDoi-5="http://doi.org/10.5281/zenodo.5297551" figureDoi-6="http://doi.org/10.5281/zenodo.5297553" httpUri-0="https://zenodo.org/record/5297541/files/figure.png" httpUri-1="https://zenodo.org/record/5297543/files/figure.png" httpUri-2="https://zenodo.org/record/5297545/files/figure.png" httpUri-3="https://zenodo.org/record/5297547/files/figure.png" httpUri-4="https://zenodo.org/record/5297549/files/figure.png" httpUri-5="https://zenodo.org/record/5297551/files/figure.png" httpUri-6="https://zenodo.org/record/5297553/files/figure.png" pageId="3" pageNumber="198">Figs 2–45</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="A1B043DFFF966B24FF26FE4EFE3B067B" pageId="3" pageNumber="198" type="materials_examined">
<paragraph id="E9151054FF966B24FF26FE4EFB7303E3" blockId="3.[151,1437,466,1139]" box="[151,1174,466,491]" pageId="3" pageNumber="198">
<emphasis id="DBDECC46FF966B24FF26FE4EFB7303E3" box="[151,1174,466,491]" inLineHeading="true" pageId="3" pageNumber="198" reason="1">
<emphasis id="DBDECC46FF966B24FF26FE4EFED803E3" bold="true" box="[151,317,466,491]" pageId="3" pageNumber="198">Type material</emphasis>
(numbers in parentheses correspond to localities on
<figureCitation id="71910CD1FF966B24FC3AFE4EFC2903E3" box="[907,972,466,491]" captionStart="FIGURE 1" captionStartId="2.[151,250,1863,1885]" captionTargetBox="[165,1429,667,1825]" captionTargetId="figure-281@2.[151,1436,663,1842]" captionTargetPageId="2" captionText="FIGURE 1. Distribution of sampling sites in Armenia. Occurrence of E. (E.) eurycephalus sp. nov. shown by red triangles. Sites sampled by D. Palatov marked with asterisk, other sites sampled by Ľ. Hrivniak, P. Manko &amp; J. Oboňa. The numbers of sampling sites correspond to Appendix 1 and the codes on labels and field protocols of the respective collectors." figureDoi="http://doi.org/10.5281/zenodo.5297539" httpUri="https://zenodo.org/record/5297539/files/figure.png" pageId="3" pageNumber="198">Fig. 1</figureCitation>
and Appendix 1).
</emphasis>
</paragraph>
<paragraph id="E9151054FF966B24FF76FE6AFBA6003A" blockId="3.[151,1437,466,1139]" pageId="3" pageNumber="198">
<materialsCitation id="59C21A09FF966B24FF76FE6AFBA6003A" ID-GBIF-Occurrence="3339717303" collectingDate="2015-09-02" country="Armenia" elevation="2195" latitude="41.003654" location="Akhurian River" longLatPrecision="1" longitude="43.95691" pageId="3" pageNumber="198" specimenCount="1" stateProvince="Shirak" typeStatus="holotype">
<emphasis id="DBDECC46FF966B24FF76FE6AFED30007" box="[199,310,502,527]" italics="true" pageId="3" pageNumber="198">
<typeStatus id="3611AEF6FF966B24FF76FE6AFED40007" box="[199,305,502,527]" pageId="3" pageNumber="198" type="holotype">Holotype</typeStatus>
:
</emphasis>
male imago:
<collectingCountry id="91BD50C4FF966B24FE65FE6AFDBF0007" box="[468,602,502,527]" name="Armenia" pageId="3" pageNumber="198">ARMENIA</collectingCountry>
,
<collectingRegion id="2B6EDEB6FF966B24FDD7FE6AFCF90007" box="[614,796,502,527]" country="Armenia" name="Shirak" pageId="3" pageNumber="198">Shirak province</collectingRegion>
,
<locationDeviation id="F8A70976FF966B24FC98FE6BFBB20007" box="[809,1111,502,527]" location="Musayelyan village" pageId="3" pageNumber="198">NE of Musayelyan village</locationDeviation>
; small steppe river, tributary of
<location id="EC75468FFF966B24FF04FD86FE8D003B" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B:EC75468FFF966B24FF04FD86FE8D003B" box="[181,360,538,563]" country="Armenia" latitude="41.003654" longLatPrecision="1" longitude="43.95691" name="Akhurian River" pageId="3" pageNumber="198" stateProvince="Shirak">Akhurian River</location>
;
<geoCoordinate id="8C9E7693FF966B24FEC2FD86FDC1003A" box="[371,548,538,563]" degrees="41" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="13.15" value="41.003654">41°00'13.15&quot; N</geoCoordinate>
,
<geoCoordinate id="8C9E7693FF966B24FD9EFD86FD33003B" box="[559,726,538,563]" degrees="43" direction="east" minutes="57" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="24.87" value="43.95691">43°57'24.87&quot;E</geoCoordinate>
;
<elevation id="6287F767FF966B24FD50FD86FC96003B" box="[737,883,538,563]" metricMagnitude="3" metricUnit="m" metricValue="2.195" pageId="3" pageNumber="198" unit="m" value="2195.0">
<quantity id="2E52BDB1FF966B24FD50FD86FCD3003A" box="[737,822,538,563]" metricMagnitude="3" metricUnit="m" metricValue="2.195" pageId="3" pageNumber="198" unit="m" value="2195.0">2195 m</quantity>
a.s.l.
</elevation>
;
<date id="9D143694FF966B24FC31FD86FBE6003B" box="[896,1027,538,563]" pageId="3" pageNumber="198" value="2015-09-02">
<collectingDate id="8D50CF7CFF966B24FC31FD86FBE6003B" box="[896,1027,538,563]" pageId="3" pageNumber="198" value="2015-09-02">02/09/2015</collectingDate>
</date>
,
<emphasis id="DBDECC46FF966B24FBBCFD86FBD9003B" bold="true" box="[1037,1084,538,563]" pageId="3" pageNumber="198">(36)</emphasis>
.
</materialsCitation>
</paragraph>
<paragraph id="E9151054FF966B24FF76FDA3FE780192" blockId="3.[151,1437,466,1139]" pageId="3" pageNumber="198">
<emphasis id="DBDECC46FF966B24FF76FDA3FEDC005E" box="[199,313,575,598]" italics="true" pageId="3" pageNumber="198">
<typeStatus id="3611AEF6FF966B24FF76FDA3FEDC005E" box="[199,313,575,598]" pageId="3" pageNumber="198" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="FFACDBDDFF966B24FEF8FDA2FE73005F" box="[329,406,574,599]" pageId="3" pageNumber="198" type="male">2 male</specimenCount>
imagoes (genitalia and wings of 1 individual mounted on slide),
<specimenCount id="FFACDBDDFF966B24FB39FDA2FB0E005F" box="[1160,1259,574,599]" pageId="3" pageNumber="198" type="female">9 female</specimenCount>
(1 mounted on slide) and
<materialsCitation id="59C21A09FF966B24FEBFFDFEFC3B0073" ID-GBIF-Occurrence="3339717305" box="[270,990,610,635]" collectingDate="2015-09-02" country="Armenia" elevation="2195" latitude="41.003654" location="Akhurian River" longLatPrecision="1" longitude="43.95691" pageId="3" pageNumber="198" specimenCount="6" specimenCount-male="6" stateProvince="Shirak" typeStatus="paratype">
<specimenCount id="FFACDBDDFF966B24FEBFFDFEFEBE0073" box="[270,347,610,635]" pageId="3" pageNumber="198" type="male">6 male</specimenCount>
larvae (4 mounted on slide), same data as holotype
<emphasis id="DBDECC46FF966B24FC01FDFEFC3B0073" bold="true" box="[944,990,610,635]" pageId="3" pageNumber="198">(36)</emphasis>
</materialsCitation>
;
<specimenCount id="FFACDBDDFF966B24FC5EFDFEFBBA0073" box="[1007,1119,610,635]" pageId="3" pageNumber="198" type="female">10 female</specimenCount>
(2 mounted on slide) and
<materialsCitation id="59C21A09FF966B24FA3FFDFEFBAC00CB" ID-GBIF-Occurrence="3339717304" collectingDate="2015-09-02" country="Armenia" elevation="2286" latitude="41.00559" location="Akhurian River" longLatPrecision="1" longitude="43.987232" pageId="3" pageNumber="198" specimenCount="8" specimenCount-male="8" stateProvince="Shirak" typeStatus="paratype">
<specimenCount id="FFACDBDDFF966B24FA3FFDFEFF280097" pageId="3" pageNumber="198" type="male">8 male</specimenCount>
larvae (1 mounted on slide):
<collectingCountry id="91BD50C4FF966B24FDA5FD1AFD7F0097" box="[532,666,646,671]" name="Armenia" pageId="3" pageNumber="198">ARMENIA</collectingCountry>
,
<collectingRegion id="2B6EDEB6FF966B24FD15FD1AFCBD0097" box="[676,856,646,671]" country="Armenia" name="Shirak" pageId="3" pageNumber="198">Shirak province</collectingRegion>
,
<locationDeviation id="F8A70976FF966B24FCD3FD1BFB6E0097" box="[866,1163,646,671]" location="Musayleyan village" pageId="3" pageNumber="198">NE of Musayleyan village</locationDeviation>
; steppe stream, tributary of
<location id="EC75468FFF966B24FF06FD36FE8900CB" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B:EC75468FFF966B24FF06FD36FE8900CB" box="[183,364,682,707]" country="Armenia" latitude="41.00559" longLatPrecision="1" longitude="43.987232" name="Akhurian River" pageId="3" pageNumber="198" stateProvince="Shirak">Akhurian River</location>
;
<geoCoordinate id="8C9E7693FF966B24FECBFD36FDC600CB" box="[378,547,682,707]" degrees="41" direction="north" minutes="00" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="20.12" value="41.00559">41°00'20.12&quot;N</geoCoordinate>
,
<geoCoordinate id="8C9E7693FF966B24FD80FD36FD3D00CB" box="[561,728,682,707]" degrees="43" direction="east" minutes="59" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="14.04" value="43.987232">43°59'14.04&quot;E</geoCoordinate>
;
<elevation id="6287F767FF966B24FD57FD36FC9900CB" box="[742,892,682,707]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="198" unit="m" value="2286.0">
<quantity id="2E52BDB1FF966B24FD57FD36FCD800CA" box="[742,829,682,707]" metricMagnitude="3" metricUnit="m" metricValue="2.286" pageId="3" pageNumber="198" unit="m" value="2286.0">2286 m</quantity>
a.s.l.
</elevation>
;
<date id="9D143694FF966B24FC3AFD36FBEB00CB" box="[907,1038,682,707]" pageId="3" pageNumber="198" value="2015-09-02">
<collectingDate id="8D50CF7CFF966B24FC3AFD36FBEB00CB" box="[907,1038,682,707]" pageId="3" pageNumber="198" value="2015-09-02">02/09/2015</collectingDate>
</date>
,
<emphasis id="DBDECC46FF966B24FBAAFD36FBAC00CB" bold="true" box="[1051,1097,682,707]" pageId="3" pageNumber="198">(35)</emphasis>
</materialsCitation>
;
<materialsCitation id="59C21A09FF966B24FBEAFD36FD580103" ID-GBIF-Occurrence="3339717307" collectingDate="2015-09-02" country="Armenia" elevation="1987" latitude="40.97236" location="Akhurian River" longLatPrecision="1" longitude="43.768578" pageId="3" pageNumber="198" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Shirak" typeStatus="paratype">
<specimenCount id="FFACDBDDFF966B24FBEAFD36FB5B00CB" box="[1115,1214,682,707]" pageId="3" pageNumber="198" type="female">3 female</specimenCount>
and
<specimenCount id="FFACDBDDFF966B24FB48FD36FAA300CB" box="[1273,1350,682,707]" pageId="3" pageNumber="198" type="male">1 male</specimenCount>
larvae:
<collectingCountry id="91BD50C4FF966B24FF26FD52FEF800EF" box="[151,285,718,743]" name="Armenia" pageId="3" pageNumber="198">ARMENIA</collectingCountry>
,
<collectingRegion id="2B6EDEB6FF966B24FE9AFD52FE0600EF" box="[299,483,718,743]" country="Armenia" name="Shirak" pageId="3" pageNumber="198">Shirak province</collectingRegion>
,
<locationDeviation id="F8A70976FF966B24FE43FD53FD0400EF" box="[498,737,718,743]" location="Amasia village" pageId="3" pageNumber="198">N of Amasia village</locationDeviation>
, steppe stream, tributary of
<location id="EC75468FFF966B24FB9BFD52FB0500EF" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B:EC75468FFF966B24FB9BFD52FB0500EF" box="[1066,1248,718,743]" country="Armenia" latitude="40.97236" longLatPrecision="1" longitude="43.768578" name="Akhurian River" pageId="3" pageNumber="198" stateProvince="Shirak">Akhurian River</location>
;
<geoCoordinate id="8C9E7693FF966B24FB5FFD52FA7D00EF" box="[1262,1432,718,743]" degrees="40" direction="north" minutes="58" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="20.49" value="40.97236">40°58'20.49&quot;N</geoCoordinate>
,
<geoCoordinate id="8C9E7693FF966B24FF26FD6EFED80103" box="[151,317,754,779]" degrees="43" direction="east" minutes="46" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="06.88" value="43.768578">43°46'06.88&quot;E</geoCoordinate>
;
<elevation id="6287F767FF966B24FEFFFD6EFE0E0103" box="[334,491,754,779]" metricMagnitude="3" metricUnit="m" metricValue="1.9869999999999997" pageId="3" pageNumber="198" unit="m" value="1987.0">
<quantity id="2E52BDB1FF966B24FEFFFD6EFE4C0102" box="[334,425,754,779]" metricMagnitude="3" metricUnit="m" metricValue="1.9869999999999997" pageId="3" pageNumber="198" unit="m" value="1987.0">1987 m</quantity>
a.s.l.
</elevation>
;
<date id="9D143694FF966B24FE4CFD6EFD9A0103" box="[509,639,754,779]" pageId="3" pageNumber="198" value="2015-09-02">
<collectingDate id="8D50CF7CFF966B24FE4CFD6EFD9A0103" box="[509,639,754,779]" pageId="3" pageNumber="198" value="2015-09-02">02/09/2015</collectingDate>
</date>
,
<emphasis id="DBDECC46FF966B24FD3EFD6EFD580103" bold="true" box="[655,701,754,779]" pageId="3" pageNumber="198">(38)</emphasis>
</materialsCitation>
;
<materialsCitation id="59C21A09FF966B24FD63FD6EFF20015B" ID-GBIF-Occurrence="3339717302" collectingDate="2015-09-03" country="Armenia" elevation="2052" latitude="41.07255" location="Akhurian River" longLatPrecision="1" longitude="43.745785" pageId="3" pageNumber="198" specimenCount="1" specimenCount-female="1" stateProvince="Shirak" typeStatus="paratype">
<specimenCount id="FFACDBDDFF966B24FD63FD6EFCDD0103" box="[722,824,754,779]" pageId="3" pageNumber="198" type="female">1 female</specimenCount>
larva:
<collectingCountry id="91BD50C4FF966B24FC21FD6EFBF30103" box="[912,1046,754,779]" name="Armenia" pageId="3" pageNumber="198">ARMENIA</collectingCountry>
,
<collectingRegion id="2B6EDEB6FF966B24FB97FD6EFB050103" box="[1062,1248,754,779]" country="Armenia" name="Shirak" pageId="3" pageNumber="198">Shirak province</collectingRegion>
,
<locationDeviation id="F8A70976FF966B24FB41FD6FFF030127" location="Aghvorik village" pageId="3" pageNumber="198">E of Aghvorik village</locationDeviation>
, steppe stream, tributary of
<location id="EC75468FFF966B24FD85FC8AFD0F0127" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B:EC75468FFF966B24FD85FC8AFD0F0127" box="[564,746,790,815]" country="Armenia" latitude="41.07255" longLatPrecision="1" longitude="43.745785" name="Akhurian River" pageId="3" pageNumber="198" stateProvince="Shirak">Akhurian River</location>
;
<geoCoordinate id="8C9E7693FF966B24FD4BFC8AFC410127" box="[762,932,790,815]" degrees="41" direction="north" minutes="04" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="21.19" value="41.07255">41°04'21.19&quot;N</geoCoordinate>
,
<geoCoordinate id="8C9E7693FF966B24FC02FC8AFBBF0127" box="[947,1114,790,815]" degrees="43" direction="east" minutes="44" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="44.83" value="43.745785">43°44'44.83&quot;E</geoCoordinate>
;
<elevation id="6287F767FF966B24FBDBFC8AFAE00127" box="[1130,1285,790,815]" metricMagnitude="3" metricUnit="m" metricValue="2.052" pageId="3" pageNumber="198" unit="m" value="2052.0">
<quantity id="2E52BDB1FF966B24FBDBFC8AFB260126" box="[1130,1219,790,815]" metricMagnitude="3" metricUnit="m" metricValue="2.052" pageId="3" pageNumber="198" unit="m" value="2052.0">2052 m</quantity>
a.s.l.
</elevation>
;
<date id="9D143694FF966B24FAA7FC8AFA7D0127" box="[1302,1432,790,815]" pageId="3" pageNumber="198" value="2015-09-03">
<collectingDate id="8D50CF7CFF966B24FAA7FC8AFA7D0127" box="[1302,1432,790,815]" pageId="3" pageNumber="198" value="2015-09-03">03/09/2015</collectingDate>
</date>
,
<emphasis id="DBDECC46FF966B24FF26FCA6FF20015B" bold="true" box="[151,197,826,851]" pageId="3" pageNumber="198">(39)</emphasis>
</materialsCitation>
;
<specimenCount id="FFACDBDDFF966B24FF66FCA6FEDF015B" box="[215,314,826,851]" pageId="3" pageNumber="198" type="female">3 female</specimenCount>
(1 mounted on slide) and
<materialsCitation id="59C21A09FF966B24FDC1FCA6FEB90193" ID-GBIF-Occurrence="3339717306" collectingDate="2015-09-03" country="Armenia" elevation="1885" latitude="40.931942" location="Akhurian river" longLatPrecision="1" longitude="43.87924" pageId="3" pageNumber="198" specimenCount="5" specimenCount-male="5" stateProvince="Shirak" typeStatus="paratype">
<specimenCount id="FFACDBDDFF966B24FDC1FCA6FD5B015B" box="[624,702,826,851]" pageId="3" pageNumber="198" type="male">5 male</specimenCount>
larvae (l mounted on slide):
<collectingCountry id="91BD50C4FF966B24FBA0FCA6FB72015B" box="[1041,1175,826,851]" name="Armenia" pageId="3" pageNumber="198">ARMENIA</collectingCountry>
,
<collectingRegion id="2B6EDEB6FF966B24FB14FCA6FAB8015B" box="[1189,1373,826,851]" country="Armenia" name="Shirak" pageId="3" pageNumber="198">Shirak province</collectingRegion>
,
<locationDeviation id="F8A70976FF966B24FADBFCA7FE90017F" location="Torosgyugh village" pageId="3" pageNumber="198">E of Torosgyugh village</locationDeviation>
, small steppe river, tributary of the
<location id="EC75468FFF966B24FCAAFCC2FC2D017F" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B:EC75468FFF966B24FCAAFCC2FC2D017F" box="[795,968,862,887]" country="Armenia" latitude="40.931942" longLatPrecision="1" longitude="43.87924" name="Akhurian river" pageId="3" pageNumber="198" stateProvince="Shirak">Akhurian river</location>
;
<geoCoordinate id="8C9E7693FF966B24FC67FCC2FB65017F" box="[982,1152,862,887]" degrees="40" direction="north" minutes="55" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="54.99" value="40.931942">40°55'54.99&quot;N</geoCoordinate>
,
<geoCoordinate id="8C9E7693FF966B24FB3FFCC2FAD0017F" box="[1166,1333,862,887]" degrees="43" direction="east" minutes="52" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="45.26" value="43.87924">43°52'45.26&quot;E</geoCoordinate>
;
<elevation id="6287F767FF966B24FAF5FCC2FF280193" metricMagnitude="3" metricUnit="m" metricValue="1.885" pageId="3" pageNumber="198" unit="m" value="1885.0">
<quantity id="2E52BDB1FF966B24FAF5FCC2FA79017E" box="[1348,1436,862,887]" metricMagnitude="3" metricUnit="m" metricValue="1.885" pageId="3" pageNumber="198" unit="m" value="1885.0">1885 m</quantity>
a.s.l.
</elevation>
;
<date id="9D143694FF966B24FF6BFC1EFEB90193" box="[218,348,898,923]" pageId="3" pageNumber="198" value="2015-09-03">
<collectingDate id="8D50CF7CFF966B24FF6BFC1EFEB90193" box="[218,348,898,923]" pageId="3" pageNumber="198" value="2015-09-03">03/09/2015</collectingDate>
</date>
</materialsCitation>
,
<emphasis id="DBDECC46FF966B24FED6FC1EFE700193" bold="true" box="[359,405,898,923]" pageId="3" pageNumber="198">(40)</emphasis>
.
</paragraph>
<paragraph id="E9151054FF966B24FF76FC3AFC9D060E" blockId="3.[151,1437,466,1139]" pageId="3" pageNumber="198">
<emphasis id="DBDECC46FF966B24FF76FC3AFE2001B7" box="[199,453,934,959]" italics="true" pageId="3" pageNumber="198">Other material studied</emphasis>
(not
<typeStatus id="3611AEF6FF966B24FE4EFC3AFDD101B7" box="[511,564,934,959]" pageId="3" pageNumber="198">type</typeStatus>
):
<specimenCount id="FFACDBDDFF966B24FDF6FC3AFD4D01B7" box="[583,680,934,959]" pageId="3" pageNumber="198" type="female">3 female</specimenCount>
and
<specimenCount id="FFACDBDDFF966B24FD6FFC3AFCCD01B7" box="[734,808,934,959]" pageId="3" pageNumber="198" type="male">1 male</specimenCount>
larvae,
<emphasis id="DBDECC46FF966B24FC30FC3AFC4A01B7" bold="true" box="[897,943,934,959]" pageId="3" pageNumber="198">(36)</emphasis>
;
<materialsCitation id="59C21A09FF966B24FC0FFC3AFEEE060F" ID-GBIF-Occurrence="3339717301" collectingDate="2015-09-02" country="Armenia" elevation="1949" latitude="41.066624" location="Dzoraget River" longLatPrecision="1" longitude="44.09562" pageId="3" pageNumber="198" specimenCount="1" specimenCount-female="1" stateProvince="Lorri">
<specimenCount id="FFACDBDDFF966B24FC0FFC3AFBFA01B7" box="[958,1055,934,959]" pageId="3" pageNumber="198" type="female">1 female</specimenCount>
larva:
<collectingCountry id="91BD50C4FF966B24FBDDFC3AFB1701B7" box="[1132,1266,934,959]" name="Armenia" pageId="3" pageNumber="198">ARMENIA</collectingCountry>
,
<collectingRegion id="2B6EDEB6FF966B24FB4DFC3AFA7D01B7" box="[1276,1432,934,959]" country="Armenia" name="Lorri" pageId="3" pageNumber="198">Lori province</collectingRegion>
,
<locationDeviation id="F8A70976FF966B24FF26FC57FEDD01EA" box="[151,312,970,995]" location="Katnarat" pageId="3" pageNumber="198">W of Katnarat</locationDeviation>
, steppe stream, tributary of the
<location id="EC75468FFF966B24FD26FC57FCA301EB" LSID="urn:lsid:plazi:treatment:6103A142FF966B24FF26FEFBFE3B067B:EC75468FFF966B24FD26FC57FCA301EB" box="[663,838,970,995]" country="Armenia" latitude="41.066624" longLatPrecision="1" longitude="44.09562" name="Dzoraget River" pageId="3" pageNumber="198" stateProvince="Lorri">Dzoraget River</location>
;
<geoCoordinate id="8C9E7693FF966B24FCE0FC56FC1F01EB" box="[849,1018,970,995]" degrees="41" direction="north" minutes="03" orientation="latitude" pageId="3" pageNumber="198" precision="1" seconds="59.85" value="41.066624">41°03'59.85&quot;N</geoCoordinate>
,
<geoCoordinate id="8C9E7693FF966B24FBB4FC56FB4901EB" box="[1029,1196,970,995]" degrees="44" direction="east" minutes="05" orientation="longitude" pageId="3" pageNumber="198" precision="1" seconds="44.24" value="44.09562">44°05'44.24&quot;E</geoCoordinate>
;
<elevation id="6287F767FF966B24FB06FC56FAAD01EB" box="[1207,1352,970,995]" metricMagnitude="3" metricUnit="m" metricValue="1.949" pageId="3" pageNumber="198" unit="m" value="1949.0">
<quantity id="2E52BDB1FF966B24FB06FC56FAE901EA" box="[1207,1292,970,995]" metricMagnitude="3" metricUnit="m" metricValue="1.949" pageId="3" pageNumber="198" unit="m" value="1949.0">1949 m</quantity>
a.s.l.
</elevation>
;
<date id="9D143694FF966B24FAE5FC56FF37060F" pageId="3" pageNumber="198" value="2015-09-02">
<collectingDate id="8D50CF7CFF966B24FAE5FC56FF37060F" pageId="3" pageNumber="198" value="2015-09-02">02/09/ 2015</collectingDate>
</date>
,
<emphasis id="DBDECC46FF966B24FF6CFC72FEEE060F" bold="true" box="[221,267,1006,1031]" pageId="3" pageNumber="198">(33)</emphasis>
</materialsCitation>
;
<specimenCount id="FFACDBDDFF966B24FEABFC72FE9E060F" box="[282,379,1006,1031]" pageId="3" pageNumber="198" type="female">2 female</specimenCount>
and
<specimenCount id="FFACDBDDFF966B24FE03FC72FE18060F" box="[434,509,1006,1031]" pageId="3" pageNumber="198" type="male">1 male</specimenCount>
larvae,
<emphasis id="DBDECC46FF966B24FDE7FC72FD60060F" bold="true" box="[598,645,1006,1031]" pageId="3" pageNumber="198">(35)</emphasis>
;
<specimenCount id="FFACDBDDFF966B24FD25FC72FD10060F" box="[660,757,1006,1031]" pageId="3" pageNumber="198" type="female">1 female</specimenCount>
larva,
<emphasis id="DBDECC46FF966B24FCF3FC72FC95060F" bold="true" box="[834,880,1006,1031]" pageId="3" pageNumber="198">(40)</emphasis>
.
</paragraph>
<paragraph id="E9151054FF966B24FF76FB8EFE3B067B" blockId="3.[151,1437,466,1139]" pageId="3" pageNumber="198">
All types and other studied material were collected by P. Manko, J. Oboňa and Ľ. Hrivniak. Material is deposited in the collection of the Czech Academy of Sciences, Biology Centre, Institute of Entomology, České Budějovice,
<collectingCountry id="91BD50C4FF966B24FE95FBC6FE3F067B" box="[292,474,1114,1139]" name="Czech Republic" pageId="3" pageNumber="198">Czech Republic</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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