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<document id="A581318273F5B7D0565C1FE58AF3E914" ID-DOI="10.11646/zootaxa.4999.2.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5089395" ID-ZooBank="0A01B4BE-166F-4CDB-9CFC-261B9D5A17E4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625899582046" checkinUser="plazi" docAuthor="Solovyev, Alexey V. &amp; Saldaitis, Aidas" docDate="2021" docId="5F6A453BA120AD50F987D863AC53CE5B" docLanguage="en" docName="zootaxa.4999.2.1.pdf" docOrigin="Zootaxa 4999 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Iragoides dudai Solovyev &amp; Saldaitis 2021, sp. nov." docType="treatment" docUuid="9965E432-8A88-41DC-8090-EC4282B67AE8" docUuidSource="ZooBank" docVersion="3" lastPageNumber="103" masterDocId="A3533D43A122AD52F910D90DAD03C972" masterDocTitle="Five new species of Limacodidae (Lepidoptera: Zygaenoidea) from South-East Asia" masterLastPageNumber="116" masterPageNumber="101" pageNumber="103" updateTime="1699000525101" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="2F854FC4497B9FD389460D38A18816EF" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="245F8340C96BACAF4A25F6C8B2A49C5F">
<mods:title id="C7EDCBB9AD31311C9BDB10FDB367C74A">Five new species of Limacodidae (Lepidoptera: Zygaenoidea) from South-East Asia</mods:title>
</mods:titleInfo>
<mods:name id="67B20DB9C258A5C8FF9DE5EBAE6724DC" type="personal">
<mods:role id="A639AD68FC678F1DE7891051AAA03CDD">
<mods:roleTerm id="298C654553DA34AF45C27ABB55C9F80E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1A7093D65E0D72287876CBEA017C647E">Solovyev, Alexey V.</mods:namePart>
<mods:nameIdentifier id="117DD9896B3451B6B67DEBE537F43710" type="ORCID">0000-0002-4837-2554</mods:nameIdentifier>
<mods:affiliation id="E658063FC0BB505C2B6ED75D4A555D9A">Ulyanovsk State Pedagogical University, Department of Biology and Chemistry, sq. Lenina 4 / 5, RUS- 432072, Ulyanovsk, Russia. solovyev _ alexey @ mail. ru; https: // orcid. org / 0000 - 0002 - 4837 - 2554</mods:affiliation>
<mods:nameIdentifier id="549D045437A524B3256B4935BFBC3156" type="email">solovyev_alexey@mail.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="913CA51B1FC9A0E911F91C5C798F5D98" type="personal">
<mods:role id="BA58F10ECA05D1138A96AA5EB6C55AF3">
<mods:roleTerm id="82F1E44384BEC7255CF8D545DD257346">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AE34DD127469E244E1FDDA914B49F63">Saldaitis, Aidas</mods:namePart>
<mods:affiliation id="1B02DF4182111E5A52A3A0264EC7333E">Nature Research Centre, Akademijos str. 2, LT- 08412, Vilnius- 21, Lithuania.</mods:affiliation>
</mods:name>
<mods:typeOfResource id="AC65572F2AB914AD19E3577E477C39D8">text</mods:typeOfResource>
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<mods:title id="EB9D3618964AD8EE3E073B1CEC459325">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="6EB99321E607D6F7B5A6DAADDC2D2C96">
<mods:date id="39770816F64C33CD95968E1041443B54">2021</mods:date>
<mods:detail id="4A0D19B9DAC0A3927806D802D72E65C3" type="pubDate">
<mods:number id="3C1C71C717B5BBA667B9597144E06640">2021-07-09</mods:number>
</mods:detail>
<mods:detail id="BCA676E9B2CD1B5D4452393409571DF6" type="volume">
<mods:number id="FEC3FE832F9C65B03EBB26348B6E316A">4999</mods:number>
</mods:detail>
<mods:detail id="58ABEA0E13EBA87B43D9DFE2C82BC286" type="issue">
<mods:number id="D7CC380DA8800E1F06F40B1C87DCDFF7">2</mods:number>
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<mods:start id="315F0DDB9A54A54AE0F90676C3DCE23F">101</mods:start>
<mods:end id="36857FA6F047694CCBCDDCF7EF1683D4">116</mods:end>
</mods:extent>
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<mods:classification id="F641DE84A3E545A527477838BAA50E53">journal article</mods:classification>
<mods:identifier id="348FFA278E1697EF34A40DA11BFA3735" type="DOI">10.11646/zootaxa.4999.2.1</mods:identifier>
<mods:identifier id="17D7F44E228EEB756C5A9D7B23455290" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CDBEB81638C8FF720F01B793770ECB9C" type="Zenodo-Dep">5089395</mods:identifier>
<mods:identifier id="E26285E30545C748763F2E69B78A6B33" type="ZooBank">0A01B4BE-166F-4CDB-9CFC-261B9D5A17E4</mods:identifier>
</mods:mods>
<treatment id="5F6A453BA120AD50F987D863AC53CE5B" LSID="urn:lsid:zoobank.org:act:9965E432-8A88-41DC-8090-EC4282B67AE8" httpUri="http://treatment.plazi.org/id/5F6A453BA120AD50F987D863AC53CE5B" lastPageNumber="103" pageId="2" pageNumber="103">
<subSubSection id="9FD9A7A6A120AD50F987D863ACB9C8FB" box="[151,442,366,393]" pageId="2" pageNumber="103" type="nomenclature">
<paragraph id="D77CF42DA120AD50F987D863ACB9C8FB" blockId="2.[151,442,366,430]" box="[151,442,366,393]" pageId="2" pageNumber="103">
<heading id="8C344341A120AD50F987D863ACB9C8FB" bold="true" box="[151,442,366,393]" fontSize="11" level="1" pageId="2" pageNumber="103" reason="1">
<emphasis id="E5B7283FA120AD50F987D863ACB9C8FB" bold="true" box="[151,442,366,393]" pageId="2" pageNumber="103">
<taxonomicName id="10C38FAEA120AD50F987D863AC56C8FB" ID-CoL="9BYZJ" authority="Solovyev &amp; Saldaitis, 2021" authorityName="Solovyev &amp; Saldaitis" authorityYear="2021" box="[151,341,366,393]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="dudai" status="sp. nov.">
<emphasis id="E5B7283FA120AD50F987D863AC56C8FB" bold="true" box="[151,341,366,393]" italics="true" pageId="2" pageNumber="103">Iragoides dudai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FE849544A120AD50F84CD862ACB9C8FB" box="[348,442,367,393]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9FD9A7A6A120AD50F987D89EAC9CCAD7" pageId="2" pageNumber="103" type="description">
<paragraph id="D77CF42DA120AD50F987D89EAC1AC8DC" blockId="2.[151,442,366,430]" box="[151,281,403,430]" pageId="2" pageNumber="103">
(
<figureCitation id="4FF8E8A8A120AD50F98FD89EADE5C8DF" box="[159,230,403,429]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Figs 1</figureCitation>
,
<figureCitation id="4FF8E8A8A120AD50F9E4D89EAC12C8DC" box="[244,273,403,430]" captionStart="FIGURES 16–21" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[170,1419,190,1728]" captionTargetId="figure-17@9.[156,1432,181,1741]" captionTargetPageId="9" captionText="FIGURES 16–21. Male terminalia of Iragoides spp., Euphlyctina spp. and Sansarea spp. (genitalia in ventral view, aedeagus detached in lateral view, sternite VIII (only for Sansarea spp.): 16—Iragoides dudai sp. nov., holotype, the apical spine on the aedeagus is shown by arrow (genital slide CAS-20-06, WIGJ); 17—I. crispa, India (KalimpongAlgarah), the apical spine on the aedeagus is shown by arrow (genital slide 14178, MWM/ZSM); 18—Euphlyctina butvilai sp. nov., holotype (genital slide 10- 41, ZFMK); 19—E. phaeopasta, India (Darjeeling) (genital slide SAV-08-43, MfN); 20—Sansarea alenae sp. nov., holotype, papilla-shaped medial process of gnathos is shown by arrow, enlarged medial part of gnathos is shown in upper left side (genital slide CAS-20-03, WIGJ); 21—S. circulifera, China (Sichuan) (genital slide 17718, MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089399" httpUri="https://zenodo.org/record/5089399/files/figure.png" pageId="2" pageNumber="103">16</figureCitation>
)
</paragraph>
<paragraph id="D77CF42DA120AD50F987D8D7AEA3CB4F" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F987D8D7AC44C886" bold="true" box="[151,327,474,500]" pageId="2" pageNumber="103">Type-material.</emphasis>
<materialsCitation id="67ABFE70A120AD50F85BD8D6A9A9CB68" collectingDate="2019-06-27" collectionCode="WIGJ" collectorName="Saldaitis" country="China" county="Yaan" elevation="2000" location="Erlang Shan Mt." municipality="Kangding" pageId="2" pageNumber="103" specimenCount="1" specimenCount-male="1" stateProvince="Sichuan" typeStatus="holotype">
<emphasis id="E5B7283FA120AD50F85BD8D6ACBBC887" bold="true" box="[331,440,475,501]" pageId="2" pageNumber="103">
<typeStatus id="08784A8FA120AD50F85BD8D6ACBBC887" box="[331,440,475,501]" pageId="2" pageNumber="103" type="holotype">Holotype</typeStatus>
</emphasis>
:
<specimenCount id="C1C53FA4A120AD50F8D4D8D6ACD8C887" box="[452,475,475,501]" pageId="2" pageNumber="103" type="male">♂</specimenCount>
, “
<collectingCountry id="AFD4B4BDA120AD50F8FED8D6AF35C887" box="[494,566,475,501]" name="China" pageId="2" pageNumber="103">China</collectingCountry>
, W.
<collectingRegion id="15073ACFA120AD50FB71D8D6AFBDC887" box="[609,702,475,501]" country="China" name="Sichuan" pageId="2" pageNumber="103">Sichuan</collectingRegion>
, | road
<collectingCounty id="3E1D8CA1A120AD50FA15D8D6AE3CC887" box="[773,831,475,501]" pageId="2" pageNumber="103">Yaan</collectingCounty>
/
<collectingMunicipality id="37186E57A120AD50FA57D8D6AEB0C887" box="[839,947,475,501]" pageId="2" pageNumber="103">Kangding</collectingMunicipality>
|
<location id="D21CA2F6A120AD50FAD1D8D6A977C887" LSID="urn:lsid:plazi:treatment:5F6A453BA120AD50F987D863AC53CE5B:D21CA2F6A120AD50FAD1D8D6A977C887" box="[961,1140,475,501]" country="China" county="Yaan" municipality="Kangding" name="Erlang Shan Mt." pageId="2" pageNumber="103" stateProvince="Sichuan">Erlang Shan Mt.</location>
| H-
<quantity id="103B59C8A120AD50FD8ED8D6A9F2C887" box="[1182,1265,475,501]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="103" unit="m" value="2000.0">
<elevation id="5CEE131EA120AD50FD8ED8D6A9F2C887" box="[1182,1265,475,501]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="103" unit="m" value="2000.0">2000 m</elevation>
</quantity>
| N 29˚87.340`` | E102˚30.970`` |
<date id="A37DD2EDA120AD50F84DD8F2ACDACB6B" box="[349,473,511,537]" pageId="2" pageNumber="103" value="2019-06-27">
<collectingDate id="B3392B05A120AD50F84DD8F2ACDACB6B" box="[349,473,511,537]" pageId="2" pageNumber="103" value="2019-06-27">27.vi.2019</collectingDate>
</date>
. | Butvila &amp;
<collectorName id="7A3691FBA120AD50FB78D8F2AFCBCB6B" box="[616,712,511,537]" pageId="2" pageNumber="103">Saldaitis</collectorName>
leg.” (
<collectionCode id="B1D26CE8A120AD50FA05D8F2AE5ECB6B" box="[789,861,511,537]" pageId="2" pageNumber="103">WIGJ</collectionCode>
; genital slide № CAS-20-06)
</materialsCitation>
.
<materialsCitation id="67ABFE70A120AD50FDA5D8F2AE9FCB4F" collectingDate="2019-06-27" collectionCode="CAFM, CASV, CASU" collectorName="Saldaitis" country="China" county="Yaan" elevation="2000" location="Erlang Shan Mt." municipality="Kangding" pageId="2" pageNumber="103" specimenCount="3" specimenCount-male="3" stateProvince="Sichuan" typeStatus="paratype">
<emphasis id="E5B7283FA120AD50FDA5D8F2A82ECB6B" bold="true" box="[1205,1325,511,537]" pageId="2" pageNumber="103">
<typeStatus id="08784A8FA120AD50FDA5D8F2A82ECB6B" box="[1205,1325,511,537]" pageId="2" pageNumber="103" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="C1C53FA4A120AD50FC2CD8F2A862CB6B" box="[1340,1377,511,537]" pageId="2" pageNumber="103" type="male">3♂</specimenCount>
, data as holotype (
<collectionCode id="B1D26CE8A120AD50F836DB2EAC79CB4F" box="[294,378,547,573]" pageId="2" pageNumber="103">CAFM</collectionCode>
,
<collectionCode id="B1D26CE8A120AD50F895DB2EACCCCB4F" box="[389,463,547,573]" pageId="2" pageNumber="103">CASV</collectionCode>
and
<collectionCode id="B1D26CE8A120AD50FB16DB2EAF57CB4F" box="[518,596,547,573]" pageId="2" pageNumber="103">CASU</collectionCode>
, genital slide № CAS-20-11)
</materialsCitation>
.
</paragraph>
<paragraph id="D77CF42DA120AD50F9D7DB4BA889CBBF" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DB4BAC52CB12" bold="true" box="[199,337,582,608]" pageId="2" pageNumber="103">Description</emphasis>
.
<emphasis id="E5B7283FA120AD50F871DB4AAC98CB13" box="[353,411,583,609]" italics="true" pageId="2" pageNumber="103">Male</emphasis>
. Forewing length 14.5–15.0 mm, wingspan 29.0–32.0 mm (
<figureCitation id="4FF8E8A8A120AD50FD55DB4AA98BCB13" box="[1093,1160,583,609]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Fig. 1</figureCitation>
). Antenna shortly pectinate. Ground color bronzy brown. Head, anterior part of thorax dorsally, and dorsal patches of abdomen red. White scales below base of antennae, and on distal parts of coxa, femur, tibia and tarsal segments. Forewings bronzy dark brown, with pale oblique medial fasciae running from apex to ca. 1/3 of inner margin. Hindwings greyish brown.
</paragraph>
<paragraph id="D77CF42DA120AD50F9D7DBDAADD1CAF3" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DBDAAC32CB83" box="[199,305,727,753]" italics="true" pageId="2" pageNumber="103">Genitalia</emphasis>
. Uncus triangular with slender tip and with strongly sclerotized apical spur (
<figureCitation id="4FF8E8A8A120AD50FD87DBDAA9E8CB80" box="[1175,1259,727,754]" captionStart="FIGURES 16–21" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[170,1419,190,1728]" captionTargetId="figure-17@9.[156,1432,181,1741]" captionTargetPageId="9" captionText="FIGURES 16–21. Male terminalia of Iragoides spp., Euphlyctina spp. and Sansarea spp. (genitalia in ventral view, aedeagus detached in lateral view, sternite VIII (only for Sansarea spp.): 16—Iragoides dudai sp. nov., holotype, the apical spine on the aedeagus is shown by arrow (genital slide CAS-20-06, WIGJ); 17—I. crispa, India (KalimpongAlgarah), the apical spine on the aedeagus is shown by arrow (genital slide 14178, MWM/ZSM); 18—Euphlyctina butvilai sp. nov., holotype (genital slide 10- 41, ZFMK); 19—E. phaeopasta, India (Darjeeling) (genital slide SAV-08-43, MfN); 20—Sansarea alenae sp. nov., holotype, papilla-shaped medial process of gnathos is shown by arrow, enlarged medial part of gnathos is shown in upper left side (genital slide CAS-20-03, WIGJ); 21—S. circulifera, China (Sichuan) (genital slide 17718, MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089399" httpUri="https://zenodo.org/record/5089399/files/figure.png" pageId="2" pageNumber="103">Fig. 16</figureCitation>
). Gnathos slen- der, somewhat flattened apically. Valva generally triangle-shaped, with slightly concave costa, acute apically. Juxta broad. Anellus with two lateral areas densely covered with strongly sclerotized piliform spurs. Aedeagus approximately as long as valva, tubular, strongly curved in middle part, with strongly sclerotized small apical spur dorsally.
</paragraph>
<paragraph id="D77CF42DA120AD50F9D7DA86AC9CCAD7" blockId="2.[151,1437,474,1833]" box="[199,415,907,933]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DA86AC18CAD7" box="[199,283,907,933]" italics="true" pageId="2" pageNumber="103">Female</emphasis>
. Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9FD9A7A6A120AD50F9D7DAA3A854CDD3" pageId="2" pageNumber="103" type="diagnosis">
<paragraph id="D77CF42DA120AD50F9D7DAA3A854CDD3" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DAA3AC39CABA" bold="true" box="[199,314,942,968]" pageId="2" pageNumber="103">Diagnosis</emphasis>
. The male of
<taxonomicName id="10C38FAEA120AD50F8CFDAA2AF97CABB" authority="Solovyev &amp; Saldaitis, 2021" authorityName="Solovyev &amp; Saldaitis" authorityYear="2021" box="[479,660,943,969]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="dudai" status="sp. nov.">
<emphasis id="E5B7283FA120AD50F8CFDAA2AF97CABB" box="[479,660,943,969]" italics="true" pageId="2" pageNumber="103">Iragoides dudai</emphasis>
</taxonomicName>
<emphasis id="E5B7283FA120AD50FB8DDAA2AFFACABB" bold="true" box="[669,761,943,969]" pageId="2" pageNumber="103">
<taxonomicNameLabel id="FE849544A120AD50FB8DDAA2AFFACABB" box="[669,761,943,969]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguished externally from
<taxonomicName id="10C38FAEA120AD50FD66DAA2A89FCABB" authority="(Swinhoe, 1890)" baseAuthorityName="Swinhoe" baseAuthorityYear="1890" box="[1142,1436,943,969]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="crispa">
<emphasis id="E5B7283FA120AD50FD66DAA2A9D5CABA" box="[1142,1238,943,968]" italics="true" pageId="2" pageNumber="103">I. crispa</emphasis>
(
<bibRefCitation id="B35289DCA120AD50FDF7DAA2A897CABB" author="Swinhoe, C." box="[1255,1428,943,969]" pageId="2" pageNumber="103" pagination="396 - 432" refId="ref9262" refString="Swinhoe, C. (1890) On new Indian Lepidoptera, chiefly Heterocera. Proceedings of the Scientific Meetings of the Zoological Society of London, 1889, 396 - 432. https: // doi. org / 10.1111 / j. 1469 - 7998.1889. tb 06782. x" type="journal article" year="1890">Swinhoe, 1890</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="10C38FAEA120AD50F9D8DADEACE2CA9F" authority="Hering, 1931" authorityName="Hering" authorityYear="1931" box="[200,481,979,1005]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="elongata">
<emphasis id="E5B7283FA120AD50F9D8DADEAC40CA9F" box="[200,323,979,1005]" italics="true" pageId="2" pageNumber="103">I. elongata</emphasis>
<bibRefCitation id="B35289DCA120AD50F85CDADEACE2CA9F" author="Hering, M." box="[332,481,979,1005]" pageId="2" pageNumber="103" refId="ref8341" refString="Hering, M. (1931) Limacodidae (Cochliopodidae). In: Seitz, A. (Ed.), Die Gross-Schmetterlinge der Erde. Vol. 10. Alfred Kerner Verlag, Stuttgart, pp. 665 - 728." type="book" year="1931">Hering, 1931</bibRefCitation>
</taxonomicName>
(
<figureCitation id="4FF8E8A8A120AD50F8E1DADEAF76CA9F" box="[497,629,978,1005]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Figs 2, 3, 5</figureCitation>
) by dark bronzy brown forewings bearing distinct pale oblique medial fasciae running from the apex to ca. 1/3 of the inner margin. A pattern somewhat similar to the one of the dark form of
<taxonomicName id="10C38FAEA120AD50F9A5DD16AC11CD46" baseAuthorityName="Swinhoe" baseAuthorityYear="1890" box="[181,274,1051,1076]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="crispa">
<emphasis id="E5B7283FA120AD50F9A5DD16AC11CD46" box="[181,274,1051,1076]" italics="true" pageId="2" pageNumber="103">I. crispa</emphasis>
</taxonomicName>
(
<figureCitation id="4FF8E8A8A120AD50F831DD16AC65CD47" box="[289,358,1051,1077]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Fig. 2</figureCitation>
), but the fascia in this species is not as conspicuous. The males
<taxonomicName id="10C38FAEA120AD50FD3EDD16A9E3CD47" authority="Solovyev &amp; Saldaitis, 2021" authorityName="Solovyev &amp; Saldaitis" authorityYear="2021" box="[1070,1248,1051,1077]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="dudai" status="sp. nov.">
<emphasis id="E5B7283FA120AD50FD3EDD16A9E3CD47" box="[1070,1248,1051,1077]" italics="true" pageId="2" pageNumber="103">Iragoides dudai</emphasis>
</taxonomicName>
<emphasis id="E5B7283FA120AD50FDF6DD16A83CCD47" bold="true" box="[1254,1343,1051,1077]" pageId="2" pageNumber="103">
<taxonomicNameLabel id="FE849544A120AD50FDF6DD16A83CCD47" box="[1254,1343,1051,1077]" pageId="2" pageNumber="103" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are also similar in wing pattern to the females of
<taxonomicName id="10C38FAEA120AD50FB49DD32AFB5CD2A" baseAuthorityName="Swinhoe" baseAuthorityYear="1890" box="[601,694,1087,1112]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="crispa">
<emphasis id="E5B7283FA120AD50FB49DD32AFB5CD2A" box="[601,694,1087,1112]" italics="true" pageId="2" pageNumber="103">I. crispa</emphasis>
</taxonomicName>
(
<figureCitation id="4FF8E8A8A120AD50FBD5DD32AE0BCD2B" box="[709,776,1087,1113]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Fig. 4</figureCitation>
), however the female of the new species is unknown so far. In male genitalia the new species differs from
<taxonomicName id="10C38FAEA120AD50FBB3DD6EAE02CD0E" baseAuthorityName="Swinhoe" baseAuthorityYear="1890" box="[675,769,1123,1148]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="crispa">
<emphasis id="E5B7283FA120AD50FBB3DD6EAE02CD0E" box="[675,769,1123,1148]" italics="true" pageId="2" pageNumber="103">I. crispa</emphasis>
</taxonomicName>
(
<figureCitation id="4FF8E8A8A120AD50FA01DD6EAE67CD0F" box="[785,868,1123,1149]" captionStart="FIGURES 16–21" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[170,1419,190,1728]" captionTargetId="figure-17@9.[156,1432,181,1741]" captionTargetPageId="9" captionText="FIGURES 16–21. Male terminalia of Iragoides spp., Euphlyctina spp. and Sansarea spp. (genitalia in ventral view, aedeagus detached in lateral view, sternite VIII (only for Sansarea spp.): 16—Iragoides dudai sp. nov., holotype, the apical spine on the aedeagus is shown by arrow (genital slide CAS-20-06, WIGJ); 17—I. crispa, India (KalimpongAlgarah), the apical spine on the aedeagus is shown by arrow (genital slide 14178, MWM/ZSM); 18—Euphlyctina butvilai sp. nov., holotype (genital slide 10- 41, ZFMK); 19—E. phaeopasta, India (Darjeeling) (genital slide SAV-08-43, MfN); 20—Sansarea alenae sp. nov., holotype, papilla-shaped medial process of gnathos is shown by arrow, enlarged medial part of gnathos is shown in upper left side (genital slide CAS-20-03, WIGJ); 21—S. circulifera, China (Sichuan) (genital slide 17718, MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089399" httpUri="https://zenodo.org/record/5089399/files/figure.png" pageId="2" pageNumber="103">Fig. 17</figureCitation>
, the apical spine is shown by the arrow) by a very small and strongly sclerotized apical spine on the aedeagus (
<figureCitation id="4FF8E8A8A120AD50FA2BDD8AAE8DCDD3" box="[827,910,1159,1185]" captionStart="FIGURES 16–21" captionStartId="9.[151,264,1764,1788]" captionTargetBox="[170,1419,190,1728]" captionTargetId="figure-17@9.[156,1432,181,1741]" captionTargetPageId="9" captionText="FIGURES 16–21. Male terminalia of Iragoides spp., Euphlyctina spp. and Sansarea spp. (genitalia in ventral view, aedeagus detached in lateral view, sternite VIII (only for Sansarea spp.): 16—Iragoides dudai sp. nov., holotype, the apical spine on the aedeagus is shown by arrow (genital slide CAS-20-06, WIGJ); 17—I. crispa, India (KalimpongAlgarah), the apical spine on the aedeagus is shown by arrow (genital slide 14178, MWM/ZSM); 18—Euphlyctina butvilai sp. nov., holotype (genital slide 10- 41, ZFMK); 19—E. phaeopasta, India (Darjeeling) (genital slide SAV-08-43, MfN); 20—Sansarea alenae sp. nov., holotype, papilla-shaped medial process of gnathos is shown by arrow, enlarged medial part of gnathos is shown in upper left side (genital slide CAS-20-03, WIGJ); 21—S. circulifera, China (Sichuan) (genital slide 17718, MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089399" httpUri="https://zenodo.org/record/5089399/files/figure.png" pageId="2" pageNumber="103">Fig. 16</figureCitation>
, the apical spine is shown by the arrow).
</paragraph>
</subSubSection>
<subSubSection id="9FD9A7A6A120AD50F9D7DDA7AE95CC0B" pageId="2" pageNumber="103" type="distribution">
<paragraph id="D77CF42DA120AD50F9D7DDA7AF30CD9B" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DDA7AC62CDB6" bold="true" box="[199,353,1194,1220]" pageId="2" pageNumber="103">Distribution.</emphasis>
The new species is known from the Erlang
<collectingRegion id="15073ACFA120AD50FA51DDA6AE7ACDB7" box="[833,889,1195,1221]" country="Myanmar" name="Shan" pageId="2" pageNumber="103">Shan</collectingRegion>
mountain range at the eastern edge of the Tibetan plateau in China’s
<collectingRegion id="15073ACFA120AD50F878DDC2AF33CD9B" box="[360,560,1231,1257]" country="China" name="Sichuan" pageId="2" pageNumber="103">Sichuan province</collectingRegion>
.
</paragraph>
<paragraph id="D77CF42DA120AD50F9D7DDFEAE95CC0B" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DDFEACD5CC7E" bold="true" box="[199,470,1266,1293]" pageId="2" pageNumber="103">Phenology and habitat</emphasis>
. Four males were collected at the end of June at
<quantity id="103B59C8A120AD50FAECDDFEA95ACC7F" box="[1020,1113,1267,1293]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="2" pageNumber="103" unit="m" value="2000.0">2,000 m</quantity>
asl, in well preserved mixed forest habitat dominated by various broad-leaved trees such as oaks (
<emphasis id="E5B7283FA120AD50FA85DC1AA9CBCC43" box="[917,1224,1303,1329]" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="10C38FAEA120AD50FA85DC1AA953CC43" box="[917,1104,1303,1329]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="dentata">Quercus dentata</taxonomicName>
,
<taxonomicName id="10C38FAEA120AD50FD4ADC1AA9CBCC43" box="[1114,1224,1303,1329]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="glauca">Q. glauca</taxonomicName>
</emphasis>
), poplars (
<emphasis id="E5B7283FA120AD50FC50DC1AAC89CC26" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="10C38FAEA120AD50FC50DC1AAC0CCC27" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="cathayana">Populus cathayana</taxonomicName>
,
<taxonomicName id="10C38FAEA120AD50F80BDC36AC89CC26" box="[283,394,1339,1364]" class="Magnoliopsida" family="Salicaceae" genus="Populus" kingdom="Plantae" order="Malpighiales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="simonii">P. simonii</taxonomicName>
</emphasis>
), elm (
<taxonomicName id="10C38FAEA120AD50F8CEDC36AF9ECC27" authorityName="Jacq" authorityYear="1798" box="[478,669,1339,1365]" class="Insecta" family="Tingidae" genus="Ulmus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="parvifolia">
<emphasis id="E5B7283FA120AD50F8CEDC36AF9ECC27" box="[478,669,1339,1365]" italics="true" pageId="2" pageNumber="103">Ulmus parvifolia</emphasis>
</taxonomicName>
), rhododendrons (
<emphasis id="E5B7283FA120AD50FA60DC36A858CC27" box="[880,1371,1339,1365]" italics="true" pageId="2" pageNumber="103">
<taxonomicName id="10C38FAEA120AD50FA60DC36A9C0CC27" box="[880,1219,1339,1365]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="brachycarpum">Rhododendron brachycarpum</taxonomicName>
,
<taxonomicName id="10C38FAEA120AD50FDC0DC36A858CC27" box="[1232,1371,1339,1365]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="dauricum">R. dauricum</taxonomicName>
</emphasis>
), and bamboos (
<taxonomicName id="10C38FAEA120AD50F81DDC52ACAACC0B" box="[269,425,1375,1401]" class="Liliopsida" family="Poaceae" genus="Phyllostachys" kingdom="Animalia" order="Poales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="genus">
<emphasis id="E5B7283FA120AD50F81DDC52ACAACC0B" box="[269,425,1375,1401]" italics="true" pageId="2" pageNumber="103">Phyllostachys</emphasis>
</taxonomicName>
ssp.,
<taxonomicName id="10C38FAEA120AD50F8F8DC52AF47CC0B" authorityName="C.M.A.Stapleton" authorityYear="1994" box="[488,580,1375,1401]" class="Liliopsida" family="Poaceae" genus="Borinda" higherTaxonomySource="CoL" kingdom="Plantae" order="Poales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="genus">
<emphasis id="E5B7283FA120AD50F8F8DC52AF47CC0B" box="[488,580,1375,1401]" italics="true" pageId="2" pageNumber="103">Borinda</emphasis>
</taxonomicName>
ssp.,
<taxonomicName id="10C38FAEA120AD50FB93DC52AE1DCC0B" box="[643,798,1375,1401]" class="Liliopsida" family="Poaceae" genus="Fargesia" kingdom="Plantae" order="Poales" pageId="2" pageNumber="103" phylum="Tracheophyta" rank="species" species="undetermined">
<emphasis id="E5B7283FA120AD50FB93DC52AFE4CC0A" box="[643,743,1375,1400]" italics="true" pageId="2" pageNumber="103">Fargesia</emphasis>
spp.
</taxonomicName>
) (
<figureCitation id="4FF8E8A8A120AD50FA25DC52AE8BCC0B" box="[821,904,1375,1401]" captionStart="FIGURES 33–35" captionStartId="12.[151,264,1259,1283]" captionTargetBox="[159,1429,181,1235]" captionTargetId="figure-28@12.[159,1429,181,1235]" captionTargetPageId="12" captionText="FIGURES 33–35. Biotopes: 33—China, W. Sichuan, road Yaan/Kangding, Erlang Shan Mt., biotope of Iragoides dudai sp. nov., photo by R. Butvila; 34, 35—Nu Jiang valley, biotope of Sansarea alenae sp. nov. and Euphlyctina butvilai sp. nov., photo by R. Butvila." figureDoi="http://doi.org/10.5281/zenodo.5089409" httpUri="https://zenodo.org/record/5089409/files/figure.png" pageId="2" pageNumber="103">Fig. 33</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9FD9A7A6A120AD50F9D7DC8EAE17CCB0" pageId="2" pageNumber="103" type="etymology">
<paragraph id="D77CF42DA120AD50F9D7DC8EAE17CCB0" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DC8EAC4BCCEF" bold="true" box="[199,328,1411,1437]" pageId="2" pageNumber="103">Etymology</emphasis>
. The new species is named after our colleague, prominent Lithuanian collector and director of the World Insect Gallery, Juozas Dűda (Joniđkis,
<collectingCountry id="AFD4B4BDA120AD50FB87DCAAAE0BCCB0" box="[663,776,1447,1474]" name="Lithuania" pageId="2" pageNumber="103">Lithuania</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="9FD9A7A6A120AD50F9D7DCC6AC53CE5B" pageId="2" pageNumber="103" type="discussion">
<paragraph id="D77CF42DA120AD50F9D7DCC6AC53CE5B" blockId="2.[151,1437,474,1833]" pageId="2" pageNumber="103">
<emphasis id="E5B7283FA120AD50F9D7DCC6AC37CC97" bold="true" box="[199,308,1483,1509]" pageId="2" pageNumber="103">Remarks</emphasis>
. The species
<taxonomicName id="10C38FAEA120AD50F8DDDCC6AF29CC96" baseAuthorityName="Swinhoe" baseAuthorityYear="1890" box="[461,554,1483,1508]" class="Insecta" family="Limacodidae" genus="Iragoides" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="103" phylum="Arthropoda" rank="species" species="crispa">
<emphasis id="E5B7283FA120AD50F8DDDCC6AF29CC96" box="[461,554,1483,1508]" italics="true" pageId="2" pageNumber="103">I. crispa</emphasis>
</taxonomicName>
has several distinct forms: a pale one (
<figureCitation id="4FF8E8A8A120AD50FAF1DCC6A927CC97" box="[993,1060,1483,1509]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Fig. 3</figureCitation>
), a dark one (
<figureCitation id="4FF8E8A8A120AD50FDD0DCC6A807CC97" box="[1216,1284,1483,1509]" captionStart="FIGURES 1–15" captionStartId="8.[151,264,1753,1777]" captionTargetBox="[243,1340,188,1712]" captionTargetId="figure-28@8.[234,1354,179,1727]" captionTargetPageId="8" captionText="FIGURES 1–15. Dorsal view of Limacodidae adults: 1—Iragoides dudai sp. nov., holotype ♂ (WIGJ); 2—I. crispa, dark form, ♂, Myanmar (Chin State), Witt-limac 200 (MWM/ZSM); 3—ditto, pale form, ♂, China (Yunnan), Witt-limac 197 (MWM/ ZSM); 4—ditto, dark form, ♀, N. Vietnam (MWM/ZSM); 5—I. elongata, ♂, N. Vietnam (MWM/ZSM); 6—Sansarea alenae sp. nov., holotype ♂ (WIGJ); 7—Euphlyctina butvilai sp. nov., holotype ♂ (ZFMK); 8—E. butvilai sp. nov., ♂, paratype, China (Yunnan) (CASV); 9—E. phaeopasta, ♂, India (Darjeeling) (MfN); 10—Narosoideus witti sp. nov., holotype ♂ (MWM/ZSM); 11—N. flavidorsalis, ♂, Russia (Primorye) (MWM/ZSM); 12—N. morion, ♂, N. Vietnam (MWM/ZSM); 13—Griseothosea mousta sp. nov., holotype ♂ (MWM/ZSM); 14—G. fasciata, ♂, Nepal (Dhaulagiri Himal) (MWM/ZSM); 15—G. cruda, ♂, Malaysia (Sabah) (MWM/ZSM)." figureDoi="http://doi.org/10.5281/zenodo.5089397" httpUri="https://zenodo.org/record/5089397/files/figure.png" pageId="2" pageNumber="103">Fig. 2</figureCitation>
), and various intermediate forms. The typical darker individuals seem to occur mainly in
<collectingCountry id="AFD4B4BDA120AD50FACADCE2A915CF7B" box="[986,1046,1519,1545]" name="India" pageId="2" pageNumber="103">India</collectingCountry>
,
<collectingCountry id="AFD4B4BDA120AD50FD30DCE2A965CF7B" box="[1056,1126,1519,1545]" name="Nepal" pageId="2" pageNumber="103">Nepal</collectingCountry>
,
<collectingCountry id="AFD4B4BDA120AD50FD60DCE2A9DCCF7B" box="[1136,1247,1519,1545]" name="Myanmar" pageId="2" pageNumber="103">Myanmar</collectingCountry>
,
<collectingCountry id="AFD4B4BDA120AD50FDF9DCE2A82FCF7B" box="[1257,1324,1519,1545]" name="China" pageId="2" pageNumber="103">China</collectingCountry>
(
<collectingRegion id="15073ACFA120AD50FC29DCE2A89BCF7B" box="[1337,1432,1519,1545]" country="China" name="Sichuan" pageId="2" pageNumber="103">Sichuan</collectingRegion>
,
<collectingRegion id="15073ACFA120AD50F987DF1EADE0CF5F" box="[151,227,1555,1581]" country="China" name="Henan" pageId="2" pageNumber="103">Henan</collectingRegion>
,
<collectingRegion id="15073ACFA120AD50F9FEDF1EAC4BCF5F" box="[238,328,1555,1581]" country="China" name="Yunnan" pageId="2" pageNumber="103">Yunnan</collectingRegion>
),
<collectingCountry id="AFD4B4BDA120AD50F84CDF1EACC3CF5F" box="[348,448,1555,1581]" name="Thailand" pageId="2" pageNumber="103">Thailand</collectingCountry>
and
<collectingCountry id="AFD4B4BDA120AD50F8E7DF1EAF5ACF5F" box="[503,601,1555,1581]" name="Vietnam" pageId="2" pageNumber="103">Vietnam</collectingCountry>
, while the paler ones have so far been recorded only in
<collectingRegion id="15073ACFA120AD50FDC4DF1EA82DCF5F" box="[1236,1326,1555,1581]" country="China" name="Shaanxi" pageId="2" pageNumber="103">Shaanxi</collectingRegion>
and
<collectingRegion id="15073ACFA120AD50FC76DF1EADBCCF23" country="China" name="Yunnan" pageId="2" pageNumber="103">Yunnan</collectingRegion>
in
<collectingCountry id="AFD4B4BDA120AD50F9F3DF3AAC29CF23" box="[227,298,1591,1617]" name="China" pageId="2" pageNumber="103">China</collectingCountry>
. Such variations of forms could be due to environmental factors influencing the expression of specific genes; however, it is impossible at this stage to give a conclusive verdict. The specimens of different forms show no differences in male genitalia. Moreover, the molecular distances of the fragment of the gene
<emphasis id="E5B7283FA120AD50FDB3DF72A9D0CFEB" box="[1187,1235,1663,1689]" italics="true" pageId="2" pageNumber="103">COI</emphasis>
between pale and dark forms range from 0 to 4.9%, whereas the distances between specimens of dark forms from 1.3 to 4.9%. Despite of the maximum intraspecific divergence of 4.9%, we did not find any morphological difference and therefore, we accept these forms as belonging to the same taxon. Further investigation is needed when more material or information is available.
</paragraph>
</subSubSection>
</treatment>
</document>
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