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<document ID-CLB-Dataset="6649" ID-DOI="http://dx.doi.org/10.3897/zse.94.26859" ID-GBIF-Dataset="30d1d279-f033-4868-b0c9-925a406be2c5" ID-Pensoft-Pub="1860-0743-2-369" ID-Pensoft-UUID="83D45F0665FE5C729C47210ACD710944" ID-Zenodo-Dep="1344764" ID-ZooBank="C354D7DF198F4D4FA2A8F763FD03018D" ModsDocID="1860-0743-94-2-369" checkinTime="1621423395062" checkinUser="pensoft" docAuthor="Song, Zhi-Shun, Malenovsky, Igor, Chen, Jian-Qin, Deckert, Juergen &amp; Liang, Ai-Ping" docDate="2018" docId="5F32E155187AE727B82545CC3A6F69C2" docLanguage="en" docName="ZoosystEvol 94(2): 369-391" docOrigin="Zoosystematics and Evolution 94 (2)" docPubDate="2018-08-06" docSource="http://dx.doi.org/10.3897/zse.94.26859" docTitle="Orthopagus exoletus , comb. n. 2018, stat. rev." docType="treatment" docVersion="5" id="83D45F0665FE5C729C47210ACD710944" lastPageNumber="373" masterDocId="83D45F0665FE5C729C47210ACD710944" masterDocTitle="Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species" masterLastPageNumber="391" masterPageNumber="369" pageNumber="371" updateTime="1732751584564" updateUser="ExternalLinkService">
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<mods:title id="3A74792CE229F242696D8C1F602AE2E3">Taxonomic review of the planthopper genus Orthopagus (Hemiptera, Fulgoromorpha, Dictyopharidae), with descriptions of two new species</mods:title>
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<mods:namePart id="FED9EB730A3E78770CC39BAFF3600A71">Song, Zhi-Shun</mods:namePart>
<mods:affiliation id="AFB474794918A77768233B07F837B2A2">Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry &amp; Chemical Engineering, Ji &amp; angsu Second Normal University, Nanjing 210013, China</mods:affiliation>
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<mods:namePart id="6FE011DB8ACF235B2180124524CCF54F">Malenovsky, Igor</mods:namePart>
<mods:affiliation id="5095F6D7097793578D2D29A5C7E3741C">Southeast Asia Biodiversity Research Institute, Chinese Academy of Sciences, Yezin, Nay Pyi Taw 05282, Myanmar</mods:affiliation>
<mods:nameIdentifier id="6E6241E9D2FA79AF74D922472B2323A6" type="email">imalenovsky@mzm.cz</mods:nameIdentifier>
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<mods:namePart id="DA6E2EA44C0B43732A9965A632CAAF20">Chen, Jian-Qin</mods:namePart>
<mods:affiliation id="18E663CB9F9D2B8B65AB1053C1C56B1B">Jiangsu Key Laboratory of Biofunctional Molecules, School of Life Sciences, Chemistry &amp; Chemical Engineering, Ji</mods:affiliation>
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<mods:namePart id="7DFD1D52E9B21DC600B1C47E13A2E833">Deckert, Juergen</mods:namePart>
<mods:affiliation id="CEBD4A98D9DC6EC53268A4040399090D">Department of Botany and Zoology, Faculty of Science, Masaryk University, Kotlarska 2, 611 37 Brno, Czech Republic</mods:affiliation>
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<mods:namePart id="E7438E5BBC37F625339DEC9172B3990C">Liang, Ai-Ping</mods:namePart>
<mods:affiliation id="34C9ECBABE6ED8A16072601F6464CA08">Museum fuer Naturkunde, Leibniz Institute for Research on Evolution and Biodiversity Science, Invalidenstrasse 43, Berlin 10115, Germany &amp; Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<treatment id="5F32E155187AE727B82545CC3A6F69C2" ID-GBIF-Taxon="182236654" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2" httpUri="http://treatment.plazi.org/id/5F32E155187AE727B82545CC3A6F69C2" lastPageId="4" lastPageNumber="373" pageId="2" pageNumber="371" scope_family="Dictyopharidae" scope_order="Hemiptera">
<subSubSection id="7C2F57FAAFF36EB01390C0BA42BB6D52" pageId="2" pageNumber="371" type="nomenclature">
<paragraph id="CDB7C53D2EF8138F87982E3883188DF1" pageId="2" pageNumber="371">
<taxonomicName id="1E44AB7BA88718B6AE12F7F375AC9680" ID-CoL="758Q3" authority="(Melichar, 1903), comb. n." authorityName=", comb. n." authorityYear="2018" baseAuthorityName="Melichar" baseAuthorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus exoletus" order="Hemiptera" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="exoletus" status="stat. rev.">Orthopagus exoletus (Melichar, 1903), comb. n.</taxonomicName>
<taxonomicNameLabel id="8360E36E5F45C962F06A73E9E6572BD2" pageId="2" pageNumber="371">stat. rev.</taxonomicNameLabel>
<figureCitation id="98A0DA9A95DC4EF5DDCD8284A44DC4C4" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="2" pageNumber="371">Figs 2C-D</figureCitation>
<figureCitation id="D7554A58639E02B33E6C3CB6CD2FEC0A" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="2" pageNumber="371">, 3B</figureCitation>
<figureCitation id="7A7755D550799A7A3D962DADC202D162" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="2" pageNumber="371">, 5A-I</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B91A800EE4E6005A77211E65EDCE0AB4" pageId="3" pageNumber="372" type="reference_group">
<paragraph id="B6BB9FD3E09A6C77A2321FA5447E57E4" pageId="3" pageNumber="372">
<taxonomicName id="8067F7023B4F2964C26EBFEB124C089C" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="exoleta" status="stat. rev.">
<pageBreakToken id="D284EDE59A12A3009E6B8DF26D3CE77D" pageId="3" pageNumber="372" start="start">Udugama</pageBreakToken>
exoleta
</taxonomicName>
Melichar, 1903: 28, Pl. I, figs 7, 7a-b. Syntypes: 5 females, Moruwale, Sri Lanka (not examined). Synonymized under
<taxonomicName id="29837AC12DC14D86AD50B1F74D4F807D" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama splendens" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="splendens" status="stat. rev.">Udugama splendens</taxonomicName>
(Germar, 1830) by
<bibRefCitation id="42DB47F4CBD181F9526B3001B823949C" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" refId="B6" refString="Distant, WL, 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera). Taylor &amp; Francis, London" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant 1906</bibRefCitation>
: 249.
</paragraph>
<paragraph id="533C37387769754C1BADD04AA265EB3F" pageId="3" pageNumber="372">
<taxonomicName id="0BAD6115F47FFBAB98E4C610FB1EC9F1" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="3" pageNumber="372" phylum="Arthropoda" rank="species" species="exoleta" status="stat. rev.">Udugama exoleta</taxonomicName>
:
<bibRefCitation id="546B27D57103384CF87CDDF30673EBFE" author="Kirkaldy, GW" journalOrPublisher="Annales de la Societe entomologique de Belgique Bruxelles" pageId="12" pageNumber="381" pagination="9 - 14" refId="B12" refString="Kirkaldy, GW, 1908. Notes on some Sinhalese Rhynchota. Annales de la Societe entomologique de Belgique Bruxelles 52: 9 - 14" title="Notes on some Sinhalese Rhynchota." volume="52" year="1908">Kirkaldy 1908</bibRefCitation>
: 14.
</paragraph>
</subSubSection>
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<paragraph id="C5D885B6EF669A56F378ABE26AC50278" pageId="3" pageNumber="372">Material examined.</paragraph>
<paragraph id="3E5C572C99FB9FFB0E14AFCF7E8B7955" pageId="3" pageNumber="372">
<materialsCitation id="937F893C1A372FA8A3C50527B2597692" collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="T. R. Bell" country="INDIA" location="INDIA" specimenCount="2" specimenCount-female="2">
<emphasis id="6463251FB2113CDB38639189C7B8EA58" bold="true" pageId="3" pageNumber="372">
<collectingCountry id="8C8F3CB6036E0F9140D8321E184BB83E" name="India">INDIA</collectingCountry>
</emphasis>
:
<specimenCount id="540219E00520EEC5D711907D1BD8AB15" type="female">2 females</specimenCount>
, [no state indicated], 1934-394,
<collectorName id="447344C88827B008FA139831DDB6558C">T. R. Bell</collectorName>
leg. (BMNH)
</materialsCitation>
;
<materialsCitation id="A89840472FFE6D311634219C0C0C99CB" collectingDate="1907-10-03" collectorName="Distant" country="India" location="Calcutta" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal">
<collectingRegion id="F407EE5BB3E2580E8624DDF061651312" country="India" name="West Bengal">West Bengal</collectingRegion>
:
<specimenCount id="C9774486B1CB6C5F096277630A82A48F" type="female">1 female</specimenCount>
,
<location id="3DB4C01D641E74FF1BA04872391CA973" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:3DB4C01D641E74FF1BA04872391CA973" country="India" name="Calcutta" stateProvince="West Bengal">Calcutta</location>
[= Kolkata],
<collectingDate id="91148BCDB9F9D2279E6158C9ED122377" value="1907-10-03">3.x.1907</collectingDate>
(coll.
<collectorName id="21000ADA6FDEF54DFC4527F3F17B488D">Distant</collectorName>
, BMNH)
</materialsCitation>
;
<materialsCitation id="C01972605DCD372B422D740D94413224" collectingDate="2005-09-28" collectingDateMax="2005-10-02" collectingDateMin="2005-09-28" collectorName="C. R. Bartlett" country="India" county="Sindhudurg district" elevation="730" latitude="18.443611" location="Tamini village" longLatPrecision="21" longitude="73.42944" municipality="Pune district" specimenCount="2" specimenCount-male="2" stateProvince="Maharashtra">
<collectingRegion id="4F832B0291A5B9E6385EB77430A95C1A" country="India" name="Maharashtra">Maharashtra</collectingRegion>
:
<specimenCount id="AF0EF7D2246330998728B9F40F287E23" type="male">1 male</specimenCount>
,
<collectingCounty id="992B70BA256C9C3A508485C8987D1B0B">Sindhudurg district</collectingCounty>
, roadside on ridge
<locationDeviation id="0466C19CA15FA660BE2D98A9771AAB62" location="Amboli">
<quantity id="F58A2AE6BA3BD4C8A659FBD53AD130A7" metricMagnitude="3" metricUnit="m" metricValue="2.0" unit="km" value="2.0">2 km</quantity>
W Amboli
</locationDeviation>
,
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,
<geoCoordinate id="C0F43455C9A7D4C22F5812480EEDD0FC" degrees="73" direction="east" minutes="59" orientation="longitude" precision="15" seconds="23" value="73.989716">73°59'23&quot;E</geoCoordinate>
, alt.
<quantity id="37AD7C750BD0D7F02B421EF95D6A9D9E" metricMagnitude="2" metricUnit="m" metricValue="7.296912" unit="ft" value="2394.0">
<elevation id="7A81FEA370980CA97EED3E1C87E42C70" metricMagnitude="2" metricUnit="m" metricValue="7.296912" unit="ft" value="2394.0">2394 ft</elevation>
</quantity>
, pasture and successional, moist deciduous forest,
<collectingDate id="3B7987C76D012081C3F96FC9CA117BF6" value="2005-09-28">28.ix.2005</collectingDate>
,
<collectorName id="B2065F969D17EA3CCF6178E0C15F8A60">C. R. Bartlett</collectorName>
leg. (UDCC);
<specimenCount id="D299660AAF4A926256EC4E3158A4668E" type="male">1 male</specimenCount>
,
<collectingMunicipality id="E5F4F472BA1D9DAC66901FAE5E607719">Pune district</collectingMunicipality>
,
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<quantity id="E52349E5C995208B7143E44F7ED5E452" metricMagnitude="3" metricUnit="m" metricValue="5.0" unit="km" value="5.0">5 km</quantity>
E Mulshi Lake
</locationDeviation>
near
<location id="0E0AF540AA13EB925D876EA27FD74657" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:0E0AF540AA13EB925D876EA27FD74657" country="India" county="Sindhudurg district" latitude="18.443611" longLatPrecision="21" longitude="73.42944" municipality="Pune district" name="Tamini village" stateProvince="Maharashtra">Tamini village</location>
,
<geoCoordinate id="D99CA765FFCA00C101CA3327A31A30AD" degrees="18" direction="north" minutes="26" orientation="latitude" precision="15" seconds="37" value="18.443611">18°26'37&quot;N</geoCoordinate>
,
<geoCoordinate id="163614B6BD453B7FBC41C1657D16EE78" degrees="73" direction="east" minutes="25" orientation="longitude" precision="15" seconds="46" value="73.42944">73°25'46&quot;E</geoCoordinate>
, alt.
<quantity id="E0DE31ACA888E5AE7953A13D084DA8C0" metricMagnitude="2" metricUnit="m" metricValue="6.239256" unit="ft" value="2047.0">
<elevation id="E35091252012BD695D7C475502AEBA07" metricMagnitude="2" metricUnit="m" metricValue="6.239256" unit="ft" value="2047.0">2047 ft</elevation>
</quantity>
, deciduous forest and open areas,
<collectingDate id="5C68D16265EA5513B778D171230C0ADD" value="2005-10-02">2.x.2005</collectingDate>
,
<collectorName id="950FCD142ACD17A5610DE30ADDA454DB">C. R. Bartlett</collectorName>
(UDCC)
</materialsCitation>
;
<materialsCitation id="41B3BA535E4A48BF72DBCAC278BF6075" collectingDate="2005-09-24" collectingDateMax="2005-09-25" collectingDateMin="2005-09-24" collectorName="C. R. Bartlett" country="India" elevation="104" latitude="15.378612" location="Bhagwan Mahaveer Sanctuary" longLatPrecision="21" longitude="74.23111" municipality="Sanguem district" specimenCount="2" specimenCount-female="2" stateProvince="Goa">
<collectingRegion id="E63D2DE563609B965C43CC4460DE529F" country="India" name="Goa">Goa</collectingRegion>
:
<specimenCount id="FB7FB9543D020E14E709972F5D55EB0E" type="female">2 females</specimenCount>
,
<collectingMunicipality id="8BB895047549C2714C1197A81D8F10E9">Sanguem district</collectingMunicipality>
, near
<location id="B9F6385924AEDFABD5552A72E9594C4D" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:B9F6385924AEDFABD5552A72E9594C4D" country="India" latitude="15.378612" longLatPrecision="21" longitude="74.23111" municipality="Sanguem district" name="Bhagwan Mahaveer Sanctuary" stateProvince="Goa">Bhagwan Mahaveer Sanctuary</location>
<locationDeviation id="ACA13392A3FF50A447C98C06BAC8C1B8" location="Molem">
<quantity id="E5B89C6716BBEA8C3DA9C22F47CF8EA7" metricMagnitude="2" metricUnit="m" metricValue="1.0" unit="m" value="100.0">100 m</quantity>
E Molem
</locationDeviation>
,
<geoCoordinate id="C98AD5BB67E264B362FBC9941C08679D" degrees="15" direction="north" minutes="22" orientation="latitude" precision="15" seconds="43" value="15.378612">15°22'43&quot;N</geoCoordinate>
,
<geoCoordinate id="1B5469CFCBB58B36481F42ECADD3894B" degrees="74" direction="east" minutes="13" orientation="longitude" precision="15" seconds="52" value="74.23111">74°13'52&quot;E</geoCoordinate>
, alt.
<quantity id="1B98E3E24FC60FE8CB133AF0B929F402" metricMagnitude="2" metricUnit="m" metricValue="1.0424159999999998" unit="ft" value="342.0">
<elevation id="3F5949FF6C43B7720D450FF74198A30A" metricMagnitude="2" metricUnit="m" metricValue="1.0424159999999998" unit="ft" value="342.0">342 ft</elevation>
</quantity>
, moist deciduous forest,
<collectingDate id="474F7E15439A951C65A118C301C190D4" value="2005-09-24" valueMax="2005-09-25" valueMin="2005-09-24">24-25.ix.2005</collectingDate>
,
<collectorName id="3B061BD1A386CC2E35576DA2F5100589">C. R. Bartlett</collectorName>
leg. (UDCC)
</materialsCitation>
;
<materialsCitation id="9F2A167EE752B1825AF33F0C3B556507" collectingDate="1914-09-16" collectingDateMax="1938-09-22" collectingDateMin="1914-09-16" collectorName="F. H. Gravely, Distant, Tenmalai, Thenmala" country="India" county="Malabar" elevation="61" location="Parambikulam" municipality="Nadungayam" specimenCount="5" specimenCount-female="3" specimenCount-male="2" stateProvince="Kerala">
<collectingRegion id="F02F7F769F299626C8DA930B202D9E14" country="India" name="Kerala">Kerala</collectingRegion>
:
<specimenCount id="2A3EC5D954119012E05D9019E765E05E" type="male">2 males</specimenCount>
,
<collectingCounty id="EE2E82E125954044996F955CFB1D614F">Malabar</collectingCounty>
,
<collectingMunicipality id="B3E2B21A2BFBB4A94A4AE0A3ED70A82C">Nadungayam</collectingMunicipality>
[forest near Nilambur],
<quantity id="3058E19A63BB679FD6A07A32DF15F01E" metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">
<elevation id="B58BC8C681B76DD3A73AC5A3EED99108" metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">200 ft</elevation>
</quantity>
,
<collectingDate id="DF227050560C60D8AADA2A3973B11C32" value="1938-09-16" valueMax="1938-09-22" valueMin="1938-09-16">16.-22.ix.1938</collectingDate>
(BMNH);
<specimenCount id="7132E3F9385929AF71C196159AF0CB84" type="female">1 female</specimenCount>
,
<location id="EA752546781A4B03AAAE4974749660C5" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:EA752546781A4B03AAAE4974749660C5" country="India" county="Malabar" municipality="Nadungayam" name="Parambikulam" stateProvince="Kerala">Parambikulam</location>
, alt.
<quantity id="1DA1228ACCDB34D82DE0B8F3BF5358F8" metricMagnitude="2" metricUnit="m" metricValue="7.4676" metricValueMax="9.7536" metricValueMin="5.1816" unit="ft" value="2450.0" valueMax="3200.0" valueMin="1700.0">
<elevation id="87D07905A883D5F27CA6F9E83A4B38F7" metricMagnitude="2" metricUnit="m" metricValue="7.4676" metricValueMax="9.7536" metricValueMin="5.1816" unit="ft" value="2450.0" valueMax="3200.0" valueMin="1700.0">1700-3200 ft</elevation>
</quantity>
,
<collectingDate id="614BE97A9A325B615E62F4D876755C54" value="1914-09-16" valueMax="1914-09-24" valueMin="1914-09-16">16.-24.ix.1914</collectingDate>
,
<collectorName id="D2918D6C2C7CF61ACA8E01BCD5141F1E">F. H. Gravely</collectorName>
leg. (coll.
<collectorName id="A18D2041AD703F35CFD506A4831CA5A5">Distant</collectorName>
, BMNH);
<specimenCount id="EDBF6EF091670EC29CA987DC13C4F46F" type="female">2 females</specimenCount>
,
<collectorName id="38EE881638E8E47D6639967F563667F9">Tenmalai</collectorName>
[=
<collectorName id="38A55E5974FB92A4D38073C116CC063B">Thenmala</collectorName>
],
<collectingDate id="BC657A2499DC32B851D283EE49BBD65A" value="1937-05-12" valueMax="1937-05-15" valueMin="1937-05-12">12.-15.v.1937</collectingDate>
(BMNH)
</materialsCitation>
;
<materialsCitation id="D09E2DAC57C65C603085F7819F2E7E1F" collectingDate="2002-05-03" collectingDateMax="2002-05-15" collectingDateMin="2002-05-03" collectorName="L. Dembicky" country="India" elevation="1100" location="Kunchappanai" municipality="Nilgiri Hills" specimenCount="1" specimenCount-female="1" stateProvince="Tamil Nadu">
<collectingRegion id="D4A26EC8341F105257948D8BBC61AEB3" country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
:
<specimenCount id="5D5AE358AB6C6B7A036B25470A81D193" type="female">1 female</specimenCount>
,
<collectingMunicipality id="55AE04131B570592AECFC86D4816187F">Nilgiri Hills</collectingMunicipality>
,
<locationDeviation id="AE6CC3BA5EF845C445E2D01FD1692F7C" location="Kotagiri">
<quantity id="3DADFADE2A21D2AC39FDD1207BDF8B55" metricMagnitude="4" metricUnit="m" metricValue="1.1" unit="km" value="11.0">11 km</quantity>
SE Kotagiri
</locationDeviation>
,
<location id="83B62F08653406187A3E8D1770BB5334" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:83B62F08653406187A3E8D1770BB5334" country="India" municipality="Nilgiri Hills" name="Kunchappanai" stateProvince="Tamil Nadu">Kunchappanai</location>
, 11°24ʹN, 76°56ʹE, alt.
<quantity id="EB433354AA0B65B92C79F36B86F864CD" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
<elevation id="F1F85D9118DE50D8CC0CE3F71BD12B2E" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" unit="m" value="1100.0" valueMax="1200.0" valueMin="1000.0">
1100
<normalizedToken id="1E80F79FECAED9365A6D5CDED736F235" originalValue="±">+/-</normalizedToken>
100 m
</elevation>
</quantity>
,
<collectingDate id="9206243081827BB1700113813121FB85" value="2002-05-03" valueMax="2002-05-15" valueMin="2002-05-03">3.-15.v.2002</collectingDate>
,
<collectorName id="B2A14C8F6CEAFCAF03A39780413168B3">
L.
<normalizedToken id="FB4CC1A47ACEE14C2BA871E4C4C05946" originalValue="Dembický">Dembicky</normalizedToken>
</collectorName>
leg. (MMBC)
</materialsCitation>
.
<materialsCitation id="56C96FCA8BBBA2A6075F6F796738EBD9" collectingDate="2018-01-01" collectingDateMax="2018-12-31" collectingDateMin="2018-01-01" collectorName="Melichar" country="SRI LANKA" location="Puttalam" municipality="Puttalam district" specimenCount="1" specimenCount-female="1" stateProvince="North Western Province">
<emphasis id="A7FDF46FB0AA38CEDB4D51909730661A" bold="true" pageId="3" pageNumber="372">
<collectingCountry id="962FD5AC0548A0E54D1F7254F7ABF38C" name="Sri Lanka">SRI LANKA</collectingCountry>
</emphasis>
:
<collectingRegion id="B9CBFB4BF1B99C9FDE83D83E31A68517" country="Sri Lanka" name="North Western">North Western Province</collectingRegion>
:
<specimenCount id="062BC06E9EF63B477A08F4274B4F690E" type="female">1 female</specimenCount>
,
<collectingMunicipality id="D11C9901F625F25665277203DC0AA36F">Puttalam district</collectingMunicipality>
,
<location id="98C618000FDF276A3CE5108D2A7408FB" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:98C618000FDF276A3CE5108D2A7408FB" country="SRI LANKA" municipality="Puttalam district" name="Puttalam" stateProvince="North Western Province">Puttalam</location>
,
<normalizedToken id="96BF503BA0678C873C8B1E2D2BA8DD7F" originalValue="“12.”">&quot;12.&quot;</normalizedToken>
[=?1912] (coll.
<collectorName id="D210981C19D2C2A8241B8AE1F41DEE73">Melichar</collectorName>
, MMBC)
</materialsCitation>
;
<materialsCitation id="6E917E109CE76AD41AA19DEE0576B87D" collectingDate="1970-03-10" collectingDateMax="1970-03-19" collectingDateMin="1970-03-10" collectorName="D. Davis, B. Rowe" country="Sri Lanka" county="Anuradnapura district" elevation="61" location="Hunuwilagama" municipality="Wilpattu National Wildlife Park" specimenCount="1" specimenCount-male="1" stateProvince="North Central Province">
<collectingRegion id="EED90691F348F862FB5036CB3B7C042D" country="Sri Lanka" name="North Central">North Central Province</collectingRegion>
:
<specimenCount id="2F231E12699ECB052289731EAA8A9904" type="male">1 male</specimenCount>
,
<collectingCounty id="6FFC37F3EA783F92D45A35D145E707C7">Anuradnapura district</collectingCounty>
,
<collectingMunicipality id="3F7CFEEFE09B06CA96E92281DA66D8B9">Wilpattu National Wildlife Park</collectingMunicipality>
,
<location id="A31F36E8D5B7B903881939635780450A" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:A31F36E8D5B7B903881939635780450A" country="Sri Lanka" county="Anuradnapura district" municipality="Wilpattu National Wildlife Park" name="Hunuwilagama" stateProvince="North Central Province">Hunuwilagama</location>
,
<location id="29112F908E0B495AC2435F947F7EB513" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:29112F908E0B495AC2435F947F7EB513" country="Sri Lanka" county="Anuradnapura district" municipality="Wilpattu National Wildlife Park" name="Wildlife Soc. Bungalow" stateProvince="North Central Province">Wildlife Soc. Bungalow</location>
,
<quantity id="F978804108A92342F908B3318B91E894" metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">
<elevation id="1593D38BC0735760534853152E06CF0C" metricMagnitude="1" metricUnit="m" metricValue="6.096" unit="ft" value="200.0">200 ft</elevation>
</quantity>
,
<collectingDate id="3D10B0323405CAD8DE005306866404D7" value="1970-03-10" valueMax="1970-03-19" valueMin="1970-03-10">10-19.iii.1970</collectingDate>
,
<collectorName id="556D3F6AF4E0021141704AA888E8EE0A">D. Davis</collectorName>
&amp;
<collectorName id="BBA56C6DF0DBC4EBBB0FC19993EFC1AD">B. Rowe</collectorName>
leg. (USNM)
</materialsCitation>
;
<materialsCitation id="5DB56D5B8B44EC037654B134B55FD1E4" collectingDate="1970-01-31" collectingDateMax="1995-11-29" collectingDateMin="1970-01-31" collectorName="M. Schaffer, Samaragamuwa, D. Davis, B. Rowe" country="Sri Lanka" county="Kandy district" elevation="107" location="Ratnapura district" municipality="Talwatte" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Central Province">
<collectingRegion id="E0137D3D455200A9CB7DE66D50B0E123" country="Sri Lanka" name="Central">Central Province</collectingRegion>
:
<specimenCount id="2FD060D1482FF9A62E410E638DF7F62D" type="female">1 female</specimenCount>
,
<collectingCounty id="551BA195189EC15389AD154849D68BFE">Kandy district</collectingCounty>
,
<collectingMunicipality id="483391CE17EE869A821829DBAEF9A935">Talwatte</collectingMunicipality>
,
<collectingDate id="20A9E34CE394116AE76539B193E019BD" value="1995-11-29">29.xi.1995</collectingDate>
,
<collectorName id="A5B59F8DA9E6A2E8A9544B402D0A95D1">M. Schaffer</collectorName>
leg. (BMNH);
<collectorName id="6CE35FC12255C5890887709F76C05E97">Samaragamuwa</collectorName>
:
<specimenCount id="34569488710E9C1575881D91137814D5" type="male">1 male</specimenCount>
,
<location id="2E9E28A70AF361FD3F83CE34AA26F573" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:2E9E28A70AF361FD3F83CE34AA26F573" country="Sri Lanka" county="Kandy district" municipality="Talwatte" name="Ratnapura district" stateProvince="Central Province">Ratnapura district</location>
,
<location id="B6A2690BE3D4973D3EFB97AAF6D8FD37" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:B6A2690BE3D4973D3EFB97AAF6D8FD37" country="Sri Lanka" county="Kandy district" municipality="Talwatte" name="Uggalkaltota" stateProvince="Central Province">Uggalkaltota</location>
,
<location id="30E27F8C6BDAC6C4F6620CC968DAF417" LSID="urn:lsid:plazi:treatment:5F32E155187AE727B82545CC3A6F69C2:30E27F8C6BDAC6C4F6620CC968DAF417" country="Sri Lanka" county="Kandy district" municipality="Talwatte" name="Irrigation Bungalow" stateProvince="Central Province">Irrigation Bungalow</location>
, alt.
<quantity id="2A3B9D00EBB395C67384CD94FBBF4738" metricMagnitude="2" metricUnit="m" metricValue="1.0668" unit="ft" value="350.0">
<elevation id="003830DF46886F41C6B277358BCB6C92" metricMagnitude="2" metricUnit="m" metricValue="1.0668" unit="ft" value="350.0">350 ft</elevation>
</quantity>
,
<collectingDate id="F71A644E52D3D35A3E8D225D63CBE7DC" value="1970-01-31" valueMax="1970-02-08" valueMin="1970-01-31">31.i-8.ii.1970</collectingDate>
,
<collectorName id="A465458C7D3FC813852539224671E01B">D. Davis</collectorName>
&amp;
<collectorName id="8B37360F8F7033D4BF661B647264207F">B. Rowe</collectorName>
leg. (USNM)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="E40C26AA74F66C2DECD30889625A84A0" pageId="3" pageNumber="372" type="redescription">
<paragraph id="6E3BA440D91583479B159F782DBEC522" pageId="3" pageNumber="372">Redescription.</paragraph>
<paragraph id="B47007FBD08A50C33F78450EDC6E4DC5" pageId="3" pageNumber="372">Measurements (2 males, 8 females). Body length (from apex of head to tip of forewings): male 11.1 mm, female 12.3-14.7 mm; head length (from apex of cephalic process to base of eyes): male 1.28-1.30 mm, female 1.28-1.40 mm; head width (including eyes): male 1.40-1.45 mm, female 1.48-1.60 mm; forewing length: male 9.0-9.4 mm, female 10.2-12.2 mm.</paragraph>
<paragraph id="DD7A069924F0C26A0C165A3F513CC885" pageId="3" pageNumber="372">
Coloration. General coloration as in generic description (Fig.
<figureCitation id="4D275D7C54F6D63B1B7A28410E80D197" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Habitus of Orthopagus species. (A, B) O. bartletti sp. n., male, holotype, Karnataka, India; (C, D) O. exoletus (Melichar), male and female, Sri Lanka; (E, F) O. hainanensis sp. n., male and female, paratypes, Hainan, China; (G, H) O. lunulifer Uhler, male and female, China; (I, J) O. philippinus Melichar, male and female, Philippines; (K, L) O. splendens (Germar), male and female, China." figureDoi="10.3897/zse.94.26859.figure2" httpUri="https://binary.pensoft.net/fig/221398" pageId="3" pageNumber="372">2A</figureCitation>
). Vertex predominantly light ochraceous, with 3-4 pairs of small dark brown markings: an elongate patch on each side of midline apically, a small spot at each lateral keel medially, a roundish spot on each side of midline at basal third, and a small spot in each postero-lateral corner (Fig.
<figureCitation id="DF3BAD2C35CD082A8B34C5E97031BBEB" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5A</figureCitation>
). Frons light ochraceous with small dark brown spots along intermediate and lateral carinae, frons base slightly infuscated (Fig.
<figureCitation id="2F4243A30B7044A224462717F6CF1F6D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5C</figureCitation>
). Forewing membrane pattern as in Fig.
<figureCitation id="4F5A91B3522A152BA4D63C22C3A1A20C" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Forewing of Orthopagus species. (A) O. bartletti sp. n.; (B) O. exoletus (Melichar); (C) O. hainanensis sp. n.; (D) O. lunulifer Uhler; (E) O. philippinus Melichar; (F) O. splendens (Germar)." figureDoi="10.3897/zse.94.26859.figure3" httpUri="https://binary.pensoft.net/fig/221399" pageId="3" pageNumber="372">3B</figureCitation>
. Hind wing membrane with a relatively narrow dark brown streak along the apical portion of CuA1 vein, extending along hind wing apical margin in some specimens.
</paragraph>
<paragraph id="A47C661982324DC516951A591943D1C3" pageId="3" pageNumber="372">
Structure. Head with cephalic process very short, not inflated (Figs
<figureCitation id="3AEA0FF2AC4DFAD7BDCD621A6D99B319" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5A-C</figureCitation>
). Vertex (Fig.
<figureCitation id="C9B539B6ECC9B0CF15147A195C8ADCD0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5A</figureCitation>
) with ratio of length at midline to width between eyes 1.2-1.5. Transition of vertex to frons narrowly rounded in lateral view (Fig.
<figureCitation id="C894A4EC5843A7C3F9F9C2CB952596C5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5B</figureCitation>
). Frons relatively broad, with ratio of length at midline to maximum width 2.3-2.7.
</paragraph>
<paragraph id="6F73EE3034073EBBA7499815AD1A7735" pageId="3" pageNumber="372">
Male genitalia. Pygofer, in lateral view, with dorso-posterior margin produced into a relatively large but narrow, tooth-like, apically obtuse process (Fig.
<figureCitation id="3377284F02049B6D59AFD2B9A09D66DD" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5E</figureCitation>
); in ventral view (Fig.
<figureCitation id="59626C81269A1C9783C67E12A71CC9C0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5F</figureCitation>
) much longer than in dorsal view (Fig.
<figureCitation id="818B7512B4BE3E4546F1A229EF66C772" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5D</figureCitation>
) with ratio of ventral to dorsal width about 2.6. Gonostyles (Fig.
<figureCitation id="A5DEF6FD3B44609D7E905529C8315340" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5E</figureCitation>
) large and broad medially, with dorsal margin weakly sinuate medially. Aedeagus (Fig.
<figureCitation id="DE90B647443D9B160B56C2C4B409A196" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5G-I</figureCitation>
) relatively small and slender, endosomal processes curved laterad and slightly ventro- or dorsoanteriad, membranous, without distinct minute superficial spines; phallobase with three pairs of relatively small (not conspicuously inflated) membranous lobes: a pair of elongate lateral lobes with their apices gradually convergent and tapering posteriad (Fig.
<figureCitation id="6680402295BC672234DE7B08EA272D6F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5H-I</figureCitation>
), a pair of elongate thumb-like ventral lobes, directed ventroposteriad (Fig.
<figureCitation id="C476CD22297D76BAE7CA25B5CFB84DB8" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5H</figureCitation>
), and a pair of small thumb-like dorsal lobes, directed dorsolaterad (Fig.
<figureCitation id="B2AB0D133F2D7C524F08AA2507899FF2" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5G</figureCitation>
). Segment X, in lateral view, elongate, basal half narrow, gradually widening to apex beyond middle (Fig.
<figureCitation id="5A548532E35E0323C701F738D0313A82" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5E</figureCitation>
); in dorsal view, widest medially, with ratio of length to maximum width 1.1 (Fig.
<figureCitation id="BD20B8D3C35EB7CB71CCD9ED8B17AE68" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Orthopagus exoletus (Melichar). (A) Head, pronotum and mesonotum, dorsal view. (B) Head and pronotum, lateral view. (C) Head and pronotum, ventral view. (D) Male segment X and pygofer, dorsal view. (E) Male pygofer, gonostyles, and segment X, lateral view. (F) Male pygofer and gonostyles, ventral view. (G) Aedeagus, dorsal view. (H) Aedeagus, lateral view. (I) Aedeagus, ventral view." figureDoi="10.3897/zse.94.26859.figure5" httpUri="https://binary.pensoft.net/fig/221401" pageId="3" pageNumber="372">5D</figureCitation>
).
</paragraph>
<paragraph id="B2D98BCF2C7DAABDC3BABC9B71065D5D" pageId="3" pageNumber="372">Female genitalia as in generic description.</paragraph>
</subSubSection>
<subSubSection id="B5982D27A37C270037C7BBDDAB1E8E91" pageId="3" pageNumber="372" type="distribution">
<paragraph id="D3475FC1972EB121FBC139B8414FF97C" pageId="3" pageNumber="372">Distribution.</paragraph>
<paragraph id="7F5A26374F87A37ACC40FA9291BE3388" pageId="3" pageNumber="372">
India (southwestern part and West Bengal), Sri Lanka (Fig.
<figureCitation id="ABF2AE0E609DA61868B4E2E14EB68A81" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Geographical distribution of the Orthopagus species." figureDoi="10.3897/zse.94.26859.figure11" httpUri="https://binary.pensoft.net/fig/221408" pageId="3" pageNumber="372">11</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="85E1D5DA9764F1E8002B9FD253055B11" pageId="4" pageNumber="373" type="remarks">
<paragraph id="6B40D10E8C14E7FAB78EE03AF6ECDFB6" pageId="4" pageNumber="373">
<pageBreakToken id="387D3B9EC4950700F27A4686F1FCC9F9" pageId="4" pageNumber="373" start="start">Remarks</pageBreakToken>
.
</paragraph>
<paragraph id="D42F669F975DB25957F7E7944088B4D6" pageId="4" pageNumber="373">
<taxonomicName id="815E32A1595876073C5771AE5C3D5571" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoleta">
<emphasis id="F5F9C680CD918F3FA4ACFDA1F590A48F" italics="true" pageId="4" pageNumber="373">Udugama exoleta</emphasis>
</taxonomicName>
was described from Sri Lanka as the type species of
<taxonomicName id="DAFA877C406BF0318613E7A80985BCB9" authorityName="Melichar" authorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="D835EAEF5D2C6C0AB4B3AD688E6F8AD0" italics="true" pageId="4" pageNumber="373">Udugama</emphasis>
</taxonomicName>
(
<bibRefCitation id="315C5AE40573313BFD569793E3B70018" author="Melichar, L" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" refId="B20" refString="Melichar, L, 1903. Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." title="Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." year="1903">Melichar 1903</bibRefCitation>
).
<bibRefCitation id="4480F4464BDE3EF60163D33106D410B1" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" refId="B6" refString="Distant, WL, 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera). Taylor &amp; Francis, London" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant (1906)</bibRefCitation>
synonymized this species name with
<taxonomicName id="780D2E8491C08CAFEDAC199ECDF620DB" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama splendens" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="splendens">
<emphasis id="2D98A4283521820641D80D0ECE78E920" italics="true" pageId="4" pageNumber="373">Udugama splendens</emphasis>
</taxonomicName>
described from Java, Indonesia. However,
<bibRefCitation id="89D923B72EF9D2D7B5E779BC959FFC73" author="Kirkaldy, GW" journalOrPublisher="Annales de la Societe entomologique de Belgique Bruxelles" pageId="12" pageNumber="381" pagination="9 - 14" refId="B12" refString="Kirkaldy, GW, 1908. Notes on some Sinhalese Rhynchota. Annales de la Societe entomologique de Belgique Bruxelles 52: 9 - 14" title="Notes on some Sinhalese Rhynchota." volume="52" year="1908">Kirkaldy (1908)</bibRefCitation>
, probably based on comparisons of figures in
<bibRefCitation id="8E5ED91243E080160C31E3CD155B0765" author="Melichar, L" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" refId="B20" refString="Melichar, L, 1903. Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." title="Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." year="1903">Melichar (1903)</bibRefCitation>
and
<bibRefCitation id="8F56E3F1A131EB2C2CAB8EBA9A5EA9D6" author="Distant, WL" journalOrPublisher="Taylor &amp; Francis, London" pageId="12" pageNumber="381" refId="B6" refString="Distant, WL, 1906. The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera). Taylor &amp; Francis, London" title="The Fauna of British India, including Ceylon and Burma. Rhynchota Vol. III (Heteroptera-Homoptera)." year="1906">Distant (1906)</bibRefCitation>
, commented that
<taxonomicName id="276CA9D6435C7FCF1AC0AB35A0B6287B" lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis id="E015A450E0A61E4A2DFE9C11DE45A848" italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
was &quot;very different&quot; from
<taxonomicName id="1205F1D408E1AB0EE536BA459E13D5B0" lsidName="U. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis id="9C6B5CCA529674008C50CF96FA686B9E" italics="true" pageId="4" pageNumber="373">U. splendens</emphasis>
</taxonomicName>
in having a much longer face. Nevertheless, the synonymy of
<taxonomicName id="37472AABC38C1FC3817F2DB31702AEF8" lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis id="4F4E0A4C6E532328B8C03BD0CF86D9C6" italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
with
<taxonomicName id="55BC200BDA148B48FF4769E4B72B1260" lsidName="U. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis id="650AD33B131A307F1E7C329E4F4F995D" italics="true" pageId="4" pageNumber="373">U. splendens</emphasis>
</taxonomicName>
was accepted by
<bibRefCitation id="86AD690027B35C89285E374A76C32BF8" author="Melichar, L" journalOrPublisher="Zoologisch-Botanischen Gesellschaft in Wien" pageId="12" pageNumber="381" pagination="1 - 221" refId="B21" refString="Melichar, L, 1912. Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K. Zoologisch-Botanischen Gesellschaft in Wien 7 (1): 1 - 221" title="Monographie der Dictyophorinen (Homoptera). Abhandlungen der K. K." volume="7" year="1912">Melichar (1912)</bibRefCitation>
, and later included in
<bibRefCitation id="4CC6B06514224B362EC03D8BECF666D4" DOI="https://doi.org/10.1093/aesa/40.3.544" author="Metcalf, ZP" journalOrPublisher="Smith College, Northampton, USA" pageId="12" pageNumber="381" refId="B23" refString="Metcalf, ZP, 1946. General catalogue of the Hemiptera, Fasci. IV. Fulgoroidea, Part 8 Dictyopharidae. Smith College, Northampton, USA, DOI: https://doi.org/10.1093/aesa/40.3.544" title="General catalogue of the Hemiptera, Fasci. IV. Fulgoroidea, Part 8 Dictyopharidae." url="https://doi.org/10.1093/aesa/40.3.544" year="1946">
<normalizedToken id="DD0854AAA3F6D53D32979E9BB2A8E4F6" originalValue="Metcalfs">Metcalf's</normalizedToken>
(1946)
</bibRefCitation>
catalogue of world
<taxonomicName id="1F4115CAAA4D63CFFE5FA0F13DBBFCC4" authorityName="Spinola" authorityYear="1839" class="Insecta" family="Dictyopharidae" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="family">Dictyopharidae</taxonomicName>
.
</paragraph>
<paragraph id="A6BB5BD4C373656426BC84F0C551339F" pageId="4" pageNumber="373">
Based on our critical review of the published information and examination of
<taxonomicName id="C843AE37B5B984CE8AF90E13C65BE9B3" authorityName="Uhler" authorityYear="1897" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="4D900FF221925088AEB80B4E62E5D6DA" italics="true" pageId="4" pageNumber="373">Orthopagus</emphasis>
</taxonomicName>
material from Sri Lanka which agrees with the original description of
<taxonomicName id="6D0058C2EABB97AC6BA61074D200F04E" lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis id="4EDF1B88003C3DAC863C11E7ECF6FD48" italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
, we propose here to resurrect
<taxonomicName id="16A8E7C029A9B5A40FC0488CAD2FA91A" authorityName=", comb. n." authorityYear="2018" baseAuthorityName="Melichar" baseAuthorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus exoletus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis id="AE9B2553F582B84BD7A3902E3B64D053" italics="true" pageId="4" pageNumber="373">Orthopagus exoletus</emphasis>
</taxonomicName>
comb. n. from the synonymy with
<taxonomicName id="E0BB06BFCC65FA1BE40B6E1825A048EE" lsidName="O. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis id="EAFFDE985C4A295C28ADEAD63C2182FF" italics="true" pageId="4" pageNumber="373">O. splendens</emphasis>
</taxonomicName>
and to restore it as a valid species.
<taxonomicName id="40FE6387A034A682A26D98370829B7FF" authorityName=", comb. n." authorityYear="2018" baseAuthorityName="Melichar" baseAuthorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus exoletus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoletus">
<emphasis id="2F549CB15C2D7BFC2DD6C06913A8D3E9" italics="true" pageId="4" pageNumber="373">Orthopagus exoletus</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="A2B5D3D9C5DBBA28B8C1F552F3A8B2BE" lsidName="O. splendens" pageId="4" pageNumber="373" rank="species" species="splendens">
<emphasis id="FED359BA015E3D2BBD355A56138F0883" italics="true" pageId="4" pageNumber="373">O. splendens</emphasis>
</taxonomicName>
by the coloration of the vertex and the structure of the male genitalia, particularly the structure of the endosomal processes of aedeagus, lobes of the phallobase and the shape of the segment X. The relative length of frons mentioned by
<bibRefCitation id="4758965BDBA31BA41A49A15B07A6508D" author="Kirkaldy, GW" journalOrPublisher="Annales de la Societe entomologique de Belgique Bruxelles" pageId="12" pageNumber="381" pagination="9 - 14" refId="B12" refString="Kirkaldy, GW, 1908. Notes on some Sinhalese Rhynchota. Annales de la Societe entomologique de Belgique Bruxelles 52: 9 - 14" title="Notes on some Sinhalese Rhynchota." volume="52" year="1908">Kirkaldy (1908)</bibRefCitation>
is probably not a relevant diagnostic character because the length of the head in
<taxonomicName id="B46C5DCB3A09B19BB9D7F39B61E70A57" authorityName="Uhler" authorityYear="1897" class="Insecta" family="Dictyopharidae" genus="Orthopagus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthopagus" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="5D9FE67D6EDFD3587CE722C068686292" italics="true" pageId="4" pageNumber="373">Orthopagus</emphasis>
</taxonomicName>
species varies within a certain range.
</paragraph>
<paragraph id="B50FE89146605CB792CB8EEFBC186E94" pageId="4" pageNumber="373">
According to
<bibRefCitation id="0DDFC14B63C7A56DD1D5A175307EDB90" author="Melichar, L" journalOrPublisher="Transactions of the Sapporo Natural History Society" pageId="12" pageNumber="381" refId="B20" refString="Melichar, L, 1903. Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." title="Homopteren Fauna von Ceylon. Berlin: Verlag von Felix L. Dames, 248 pp." year="1903">Melichar (1903)</bibRefCitation>
,
<taxonomicName id="40D42CB72E18ED5BFBC765E248F38E9D" lsidName="U. exoleta" pageId="4" pageNumber="373" rank="species" species="exoleta">
<emphasis id="A88887B39E1088A5D3186D6291C90B9C" italics="true" pageId="4" pageNumber="373">U. exoleta</emphasis>
</taxonomicName>
was described based on five female specimens from
<normalizedToken id="8216F4784C523D5390CE19CD26A54678" originalValue="“Moruwale”">&quot;Moruwale&quot;</normalizedToken>
, deposited in the collection of the museum in Colombo, Sri Lanka. This material was not available to our study. Nevertheless, we have studied one female from Sri Lanka,
<normalizedToken id="D5714D026F5EC3FAEE028EE45861A4E8" originalValue="“Puttalam”">&quot;Puttalam&quot;</normalizedToken>
preserved in
<normalizedToken id="B4DF342BF9C0D33EB15AF6693BFC52E4" originalValue="Melichars">Melichar's</normalizedToken>
personal collection in MMBC. Even though this specimen bears original identification labels handwritten by Melichar as &quot;
<taxonomicName id="CEDBFFE99410D30B139495A71A946B9A" authorityName="Melichar" authorityYear="1903" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="genus">
<emphasis id="848102C4D74F539EFC6E5B7A2B27A2C3" italics="true" pageId="4" pageNumber="373">Udugama</emphasis>
</taxonomicName>
&quot; and &quot;
<taxonomicName id="940CB953D1442E5569CC54A66EFBDE35" authorityName="Song &amp; Malenovský &amp; Chen &amp; Deckert &amp; Liang" authorityYear="2018" class="Insecta" family="Dictyopharidae" genus="Udugama" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Udugama exoleta" order="Hemiptera" pageId="4" pageNumber="373" phylum="Arthropoda" rank="species" species="exoleta">
<emphasis id="CB9CA951EB0082B270CA8F8235574439" italics="true" pageId="4" pageNumber="373">Udugama exoleta</emphasis>
</taxonomicName>
det. Melichar&quot; and a dark red label
<normalizedToken id="DD7D28C88AF8C57AAD92885FE260CE87" originalValue="“Typus”">&quot;Typus&quot;</normalizedToken>
originally also attached to the specimen by Melichar, it probably cannot be considered as a syntype because it differs in its locality and deposition from the information published in the original description and probably it was also collected later than the original species description had been published. Melichar did not use type labels in the modern sense. He had rather adopted the practice of placing a
<normalizedToken id="5AE64A98BE9DB50D7FBE30B583C66C02" originalValue="ʻtypeʼ">ʻtypeʼ</normalizedToken>
label on one or more specimens of the most taxa (even on species previously described by other authors and identified by Melichar), specimens presumably which he himself used for comparison (
<bibRefCitation id="CCE8F9E1B7C1AAE43DD622C300BBE30D" author="Young, DA" journalOrPublisher="Budapest" pageId="13" pageNumber="382" pagination="465 - 467" refId="B52" refString="Young, DA, Soos, A, 1964. Types of Cicadellinae (Homoptera, Cicadellidae) in the Hungarian Natural History Museum. Annales historico-naturales Musei nationalis hungarici. Budapest 56: 465 - 467" title="Types of Cicadellinae (Homoptera, Cicadellidae) in the Hungarian Natural History Museum. Annales historico-naturales Musei nationalis hungarici." volume="56" year="1964">
Young and
<normalizedToken id="6B1D497A2607B1661DE278A0A3F2A2AF" originalValue="Soós">Soos</normalizedToken>
1964
</bibRefCitation>
,
<bibRefCitation id="DE03492F09F00584F4207DB88CF00431" author="Wilson, MR" journalOrPublisher="Brno" pageId="13" pageNumber="382" pagination="137 - 153" refId="B48" refString="Wilson, MR, Malenovsky, I, 2007. Tropiduchidae described by Leopold Melichar (Hemiptera, Fulgoromorpha). Acta Musei Moraviae, Scientiae biologicae. Brno 92: 137 - 153" title="Tropiduchidae described by Leopold Melichar (Hemiptera, Fulgoromorpha). Acta Musei Moraviae, Scientiae biologicae." volume="92" year="2007">
Wilson and
<normalizedToken id="5509BE6D35C063FF8FA6262FDCA42DD3" originalValue="Malenovský">Malenovsky</normalizedToken>
2007
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>