215 lines
27 KiB
XML
215 lines
27 KiB
XML
<document id="B1BCC07BA0B346E5AFEA182F34FEEB73" ID-DOI="10.5281/zenodo.183429" ID-GBIF-Dataset="1ca3efaf-ed1b-48b0-8b05-d6a5e6005e54" ID-ISSN="1175-5326" ID-Zenodo-Dep="183429" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459907369404" checkinUser="plazi" docAuthor="Rintelen, Thomas Von & Glaubrecht, Matthias" docDate="2008" docId="5F2DDF50FFB99334FF26FF4B97209B4D" docLanguage="en" docName="zt01852p049.pdf" docOrigin="Zootaxa 1852" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Tylomelania hannelorae Rintelen & Glaubrecht, 2008, sp. nov." docType="treatment" docVersion="9" lastPageNumber="46" masterDocId="A314A728FFB0933DFFB1FFDC921D9D4F" masterDocTitle="Three new species of the freshwater snail genus Tylomelania (Caenogastropoda: Pachychilidae) from the Malili lake system, Sulawesi, Indonesia" masterLastPageNumber="49" masterPageNumber="37" pageNumber="46" updateTime="1698229100631" updateUser="plazi">
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<mods:mods id="44A116C4B934B508AFB8850FE346E18E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="74C0D34D54E2EA08FA5A5B6FE96A7E60">
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<mods:title id="FA24F273C3AB2165371234F4D197B0B0">Three new species of the freshwater snail genus Tylomelania (Caenogastropoda: Pachychilidae) from the Malili lake system, Sulawesi, Indonesia</mods:title>
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<mods:name id="EB04F8E3DC0E54C3A370444C2E85CD97" type="personal">
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<mods:roleTerm id="BD50047879DC66D8F24476FB70051C3F">Author</mods:roleTerm>
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<mods:namePart id="D8A357050217F2D0FA62DE1086EE3C8F">Rintelen, Thomas Von</mods:namePart>
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<mods:name id="42F517D0F4368422812587217BE1C37E" type="personal">
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<mods:role id="DF07B6E61F96D27774D7BE6E0CA292F1">
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<mods:roleTerm id="CCB30CA726704E3021E03BDB9E9F8EE0">Author</mods:roleTerm>
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<mods:namePart id="BF6429313B61487654C43BCE4E97F3F5">Glaubrecht, Matthias</mods:namePart>
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<mods:typeOfResource id="ADEE5FE5ECE73F44918F2DD0AB542C53">text</mods:typeOfResource>
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<mods:title id="BE9D2CE4FA87AED54EAB3C4A473E396F">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="653E5CCC05EFCEDDF6B5F625F9A1F297">
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<mods:date id="0F5AD6FE9FA5CF809E96F8B677E4CAF3">2008</mods:date>
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<mods:detail id="2EDCC1D5592AB7F86F6F044412037A40" type="volume">
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<mods:number id="C174402B5DC229B4143C1B3DB02EDF4C">1852</mods:number>
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<mods:end id="E044AC891CEE2F84DDAE2519629CE4B2">49</mods:end>
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<mods:classification id="98569F86D96F6DC60B4C596DB0DC195B">journal article</mods:classification>
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<mods:identifier id="B38187613891B0FBF1260799861D9D33" type="DOI">10.5281/zenodo.183429</mods:identifier>
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<mods:identifier id="2CF182CF6FDB3391FAB2B13DF06A7756" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="3852BB39B7F7934C6E535524AD7477B0" type="Zenodo-Dep">183429</mods:identifier>
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<treatment id="5F2DDF50FFB99334FF26FF4B97209B4D" ID-DOI="http://doi.org/10.5281/zenodo.5621359" ID-GBIF-Taxon="119373538" ID-Zenodo-Dep="5621359" LSID="urn:lsid:plazi:treatment:5F2DDF50FFB99334FF26FF4B97209B4D" httpUri="http://treatment.plazi.org/id/5F2DDF50FFB99334FF26FF4B97209B4D" lastPageNumber="46" pageId="9" pageNumber="46">
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<subSubSection id="9F9E3DCDFFB99334FF26FF4B90399DFE" box="[151,548,151,177]" pageId="9" pageNumber="46" type="nomenclature">
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<paragraph id="D73B6E46FFB99334FF26FF4B90399DFE" blockId="9.[151,548,151,177]" box="[151,548,151,177]" pageId="9" pageNumber="46">
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<heading id="8C73D92AFFB99334FF26FF4B90399DFE" bold="true" box="[151,548,151,177]" fontSize="11" level="1" pageId="9" pageNumber="46" reason="1">
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<emphasis id="E5F0B254FFB99334FF26FF4B90399DFE" bold="true" box="[151,548,151,177]" pageId="9" pageNumber="46">
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<taxonomicName id="108415C5FFB99334FF26FF4B93A39DFE" box="[151,446,151,177]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="hannelorae" status="sp. nov.">
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<emphasis id="E5F0B254FFB99334FF26FF4B93A39DFE" bold="true" box="[151,446,151,177]" italics="true" pageId="9" pageNumber="46">Tylomelania hannelorae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FEC30F2FFFB99334FE77FF4B90399DFE" box="[454,548,151,177]" pageId="9" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="9F9E3DCDFFB99334FF26FF3E96B49C6B" pageId="9" pageNumber="46" type="materials_examined">
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<paragraph id="D73B6E46FFB99334FF26FF3E96B49C6B" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
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<emphasis id="E5F0B254FFB99334FF26FF3E93529DB3" bold="true" box="[151,335,226,252]" pageId="9" pageNumber="46">
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<typeStatus id="083FD0E4FFB99334FF26FF3E92CF9DB3" box="[151,210,226,252]" pageId="9" pageNumber="46">Type</typeStatus>
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material.
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</emphasis>
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<collectingCountry id="AF932ED6FFB99334FEEBFF3E93D29DB3" box="[346,463,226,252]" name="Indonesia" pageId="9" pageNumber="46">Indonesia</collectingCountry>
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, Sulawesi, Lake Mahalona, loc. 56-03:
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<typeStatus id="083FD0E4FFB99334FC00FF3E96039DB3" box="[945,1054,226,252]" pageId="9" pageNumber="46" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="4FBF72C3FFB99334FB80FF3E96649DB3" box="[1073,1145,226,252]" captionStart="FIGURE 6. T" captionStartId="8.[151,254,1816,1840]" captionTargetBox="[151,1436,397,1758]" captionTargetId="figure@8.[151,1436,394,1792]" captionTargetPageId="8" captionText="FIGURE 6. T. hannelorae sp. nov. (loc. 57 - 03). A – B. shells, A. holotype, MZB Gst. 12.114, B. paratypes, ZMB Moll. 190713. Scale bar = 1 cm. C. opercula, ZMB Moll. 190713. Scale bar = 1 mm. D – E. radula, ZMB Moll. 190713. D. segment, frontal, E. segment, apical (45 °). Scale bar = 0.1 mm, F – H. embryonic shells, ZMB Moll. 190713. F. lateral view, G. apical whorls, lateral, H. apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/183435/files/figure.png" pageId="9" pageNumber="46">Fig. 6</figureCitation>
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A;
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<quantity id="107CC3A3FFB99334FB10FF3E97169DB3" box="[1185,1291,226,252]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" pageId="9" pageNumber="46" unit="mm" value="11.8">11.8 mm</quantity>
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x
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<quantity id="107CC3A3FFB99334FA9FFF3E97929DB3" box="[1326,1423,226,252]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="9" pageNumber="46" unit="mm" value="5.5">5.5 mm</quantity>
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),
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<collectionCode id="B195F683FFB99334FF26FED692CA9C6B" box="[151,215,266,292]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="46">MZB</collectionCode>
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Gst. 12.114;
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<typeStatus id="083FD0E4FFB99334FEC5FED693F99C6B" box="[372,484,266,292]" pageId="9" pageNumber="46" type="paratype">paratypes</typeStatus>
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(
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<figureCitation id="4FBF72C3FFB99334FE45FED690269C6B" box="[500,571,266,292]" captionStart="FIGURE 6. T" captionStartId="8.[151,254,1816,1840]" captionTargetBox="[151,1436,397,1758]" captionTargetId="figure@8.[151,1436,394,1792]" captionTargetPageId="8" captionText="FIGURE 6. T. hannelorae sp. nov. (loc. 57 - 03). A – B. shells, A. holotype, MZB Gst. 12.114, B. paratypes, ZMB Moll. 190713. Scale bar = 1 cm. C. opercula, ZMB Moll. 190713. Scale bar = 1 mm. D – E. radula, ZMB Moll. 190713. D. segment, frontal, E. segment, apical (45 °). Scale bar = 0.1 mm, F – H. embryonic shells, ZMB Moll. 190713. F. lateral view, G. apical whorls, lateral, H. apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/183435/files/figure.png" pageId="9" pageNumber="46">Fig. 6</figureCitation>
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B),
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<collectionCode id="B195F683FFB99334FDD4FED690B89C6B" box="[613,677,266,292]" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="9" pageNumber="46">MZB</collectionCode>
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Gst. 12.115, n=3;
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<collectionCode id="B195F683FFB99334FCCEFED691A29C6B" box="[895,959,266,292]" httpUri="http://grbio.org/cool/8syf-d21i" name="Museum für Naturkunde Berlin (Zoological Collections)" pageId="9" pageNumber="46">ZMB</collectionCode>
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Moll.
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<date id="A33A4886FFB99334FBBFFED696749C6B" box="[1038,1129,266,292]" pageId="9" pageNumber="46" value="13-07-19">190713</date>
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, n=5.
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</paragraph>
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</subSubSection>
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<subSubSection id="9F9E3DCDFFB99334FF77FEEE90CB9C3B" pageId="9" pageNumber="46" type="etymology">
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<paragraph id="D73B6E46FFB99334FF77FEEE90CB9C3B" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
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<emphasis id="E5F0B254FFB99334FF77FEEE934E9C03" bold="true" box="[198,339,306,332]" pageId="9" pageNumber="46">Etymology.</emphasis>
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The new species has been named
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<taxonomicName id="108415C5FFB99334FD4FFEEE919E9C04" box="[766,899,306,331]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="hannelorae">
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<emphasis id="E5F0B254FFB99334FD4FFEEE919E9C04" box="[766,899,306,331]" italics="true" pageId="9" pageNumber="46">hannelorae</emphasis>
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</taxonomicName>
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in honour of Hannelore Glaubrecht, for her emotional participation in our Sulawesi research.
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</paragraph>
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</subSubSection>
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<subSubSection id="9F9E3DCDFFB99334FF77FE5E96C79EE5" pageId="9" pageNumber="46" type="description">
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<paragraph id="D73B6E46FFB99334FF77FE5E902B9F5B" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
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<emphasis id="E5F0B254FFB99334FF77FE5E93439CD3" bold="true" box="[198,350,386,412]" pageId="9" pageNumber="46">Description.</emphasis>
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Shell (
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<figureCitation id="4FBF72C3FFB99334FE08FE5E901E9CD3" box="[441,515,386,412]" captionStart="FIGURE 6. T" captionStartId="8.[151,254,1816,1840]" captionTargetBox="[151,1436,397,1758]" captionTargetId="figure@8.[151,1436,394,1792]" captionTargetPageId="8" captionText="FIGURE 6. T. hannelorae sp. nov. (loc. 57 - 03). A – B. shells, A. holotype, MZB Gst. 12.114, B. paratypes, ZMB Moll. 190713. Scale bar = 1 cm. C. opercula, ZMB Moll. 190713. Scale bar = 1 mm. D – E. radula, ZMB Moll. 190713. D. segment, frontal, E. segment, apical (45 °). Scale bar = 0.1 mm, F – H. embryonic shells, ZMB Moll. 190713. F. lateral view, G. apical whorls, lateral, H. apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/183435/files/figure.png" pageId="9" pageNumber="46">Fig. 6</figureCitation>
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A,B): Small, dark brown, elongate conic, spire angle 11–17°. Top whorls in adult specimens always corroded to a varying degree, 3–5 remaining whorls, can reach up to
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<quantity id="107CC3A3FFB99334FB5CFE76974A9C8B" box="[1261,1367,426,452]" metricMagnitude="-2" metricUnit="m" metricValue="1.39" pageId="9" pageNumber="46" unit="mm" value="13.9">13.9 mm</quantity>
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(
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<tableCitation id="9A065BFDFFB99334FAD9FE7692B59CA3" captionStart="TABLE 2" captionStartId="5.[151,239,626,650]" captionTargetBox="[159,1431,751,1103]" captionTargetPageId="5" captionText="TABLE 2. Shell parameters of the three new species. Values represent, from top, range, mean and standard deviation, and sample size. h—shell height; w—shell width; aph—aperture length; apw—aperture width; bwl—body whorl; angle—spire angle; n. a. — not applicable." httpUri="http://table.plazi.org/id/83FB3ECEFFB59338FF26FDAE90519F80" pageId="9" pageNumber="46" tableUuid="83FB3ECEFFB59338FF26FDAE90519F80">Tab. 2</tableCitation>
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). Axial and spiral ribs form reticulated pattern. Aperture oval, pointed at top and slightly siphonated at base. Columella and interior dark brown.
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</paragraph>
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<paragraph id="D73B6E46FFB99334FF77FDFE96689F73" blockId="9.[151,1436,226,1538]" box="[198,1141,546,572]" pageId="9" pageNumber="46">External morphology: Headfoot black, mantle edge serrated to a varying degree.</paragraph>
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<paragraph id="D73B6E46FFB99334FF74FD9696BE9F2B" blockId="9.[151,1436,226,1538]" box="[197,1187,586,612]" pageId="9" pageNumber="46">
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Operculum (
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<figureCitation id="4FBF72C3FFB99334FEE8FD9693BD9F2B" box="[345,416,586,612]" captionStart="FIGURE 6. T" captionStartId="8.[151,254,1816,1840]" captionTargetBox="[151,1436,397,1758]" captionTargetId="figure@8.[151,1436,394,1792]" captionTargetPageId="8" captionText="FIGURE 6. T. hannelorae sp. nov. (loc. 57 - 03). A – B. shells, A. holotype, MZB Gst. 12.114, B. paratypes, ZMB Moll. 190713. Scale bar = 1 cm. C. opercula, ZMB Moll. 190713. Scale bar = 1 mm. D – E. radula, ZMB Moll. 190713. D. segment, frontal, E. segment, apical (45 °). Scale bar = 0.1 mm, F – H. embryonic shells, ZMB Moll. 190713. F. lateral view, G. apical whorls, lateral, H. apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/183435/files/figure.png" pageId="9" pageNumber="46">Fig. 6</figureCitation>
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C): ovate, last whorl strongly inflated, multispiral with 10 whorls.
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</paragraph>
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<paragraph id="D73B6E46FFB99334FF74FDAE90799E63" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
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Radula (
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<figureCitation id="4FBF72C3FFB99334FE9AFDAE93699FC3" box="[299,372,626,652]" captionStart="FIGURE 6. T" captionStartId="8.[151,254,1816,1840]" captionTargetBox="[151,1436,397,1758]" captionTargetId="figure@8.[151,1436,394,1792]" captionTargetPageId="8" captionText="FIGURE 6. T. hannelorae sp. nov. (loc. 57 - 03). A – B. shells, A. holotype, MZB Gst. 12.114, B. paratypes, ZMB Moll. 190713. Scale bar = 1 cm. C. opercula, ZMB Moll. 190713. Scale bar = 1 mm. D – E. radula, ZMB Moll. 190713. D. segment, frontal, E. segment, apical (45 °). Scale bar = 0.1 mm, F – H. embryonic shells, ZMB Moll. 190713. F. lateral view, G. apical whorls, lateral, H. apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/183435/files/figure.png" pageId="9" pageNumber="46">Fig. 6</figureCitation>
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D,E): 28–44 rows,
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<quantity id="107CC3A3FFB99334FDEAFDAE90F69FC3" box="[603,747,626,652]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="3.2" metricValueMin="2.1" pageId="9" pageNumber="46" unit="mm" value="2.65" valueMax="3.2" valueMin="2.1">2.1–3.2 mm</quantity>
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long, on average 13.7 teeth per mm (n=5). Central tooth with very large and elongate squarish major denticle and one minor denticle on each side. Glabella narrow, with convex base. Lateral teeth with very much enlarged squarish major denticles and one minor denticle on each side. Marginal teeth shovel-like, inner and outer marginals with three denticles each, the outermost ones considerably wider than the inner ones.
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</paragraph>
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<paragraph id="D73B6E46FFB99334FF74FCE6972D9E1B" blockId="9.[151,1436,226,1538]" box="[197,1328,826,852]" pageId="9" pageNumber="46">
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Reproductive biology: Brood pouch contains 1–2 embryos, their size can reach
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<quantity id="107CC3A3FFB99334FBDAFCE696D89E1B" box="[1131,1221,826,852]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="9" pageNumber="46" unit="mm" value="3.1">3.1 mm</quantity>
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(Tab. 3).
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</paragraph>
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<paragraph id="D73B6E46FFB99334FF74FCB996C79EE5" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
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Embryonic shells (
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<figureCitation id="4FBF72C3FFB99334FE17FCB993F29E30" box="[422,495,869,895]" captionStart="FIGURE 6. T" captionStartId="8.[151,254,1816,1840]" captionTargetBox="[151,1436,397,1758]" captionTargetId="figure@8.[151,1436,394,1792]" captionTargetPageId="8" captionText="FIGURE 6. T. hannelorae sp. nov. (loc. 57 - 03). A – B. shells, A. holotype, MZB Gst. 12.114, B. paratypes, ZMB Moll. 190713. Scale bar = 1 cm. C. opercula, ZMB Moll. 190713. Scale bar = 1 mm. D – E. radula, ZMB Moll. 190713. D. segment, frontal, E. segment, apical (45 °). Scale bar = 0.1 mm, F – H. embryonic shells, ZMB Moll. 190713. F. lateral view, G. apical whorls, lateral, H. apical view. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/183435/files/figure.png" pageId="9" pageNumber="46">Fig. 6</figureCitation>
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F–H): Ovate-conic, with strong axial ribs emerging on the 2nd to 3rd whorl and fading on the 5th whorl. Shallow, widely spaced spiral ribs emerge on 3rd to 4th whorl (Tab. 3).
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</paragraph>
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</subSubSection>
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<subSubSection id="9F9E3DCDFFB99334FF74FC6491EC998D" pageId="9" pageNumber="46" type="distribution">
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<paragraph id="D73B6E46FFB99334FF74FC6496FF9E9D" blockId="9.[151,1436,226,1538]" box="[197,1250,952,978]" pageId="9" pageNumber="46">
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<emphasis id="E5F0B254FFB99334FF74FC6493E09E9D" bold="true" box="[197,509,952,978]" pageId="9" pageNumber="46">Distribution and habitat.</emphasis>
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South Sulawesi, Lake Mahalona, cape on NW shore (
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<figureCitation id="4FBF72C3FFB99334FBC9FC6496A69E9D" box="[1144,1211,952,978]" captionStart="FIGURE 1" captionStartId="1.[151,254,1586,1610]" captionTargetBox="[154,1436,281,1555]" captionTargetId="figure@1.[151,1436,277,1560]" captionTargetPageId="1" captionText="FIGURE 1. Sulawesi and the Malili lake system with sample sites. Locality numbers are the original field numbers and correspond to those in Tab. 1." httpUri="https://zenodo.org/record/183430/files/figure.png" pageId="9" pageNumber="46">Fig. 1</figureCitation>
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B).
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</paragraph>
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<paragraph id="D73B6E46FFB99334FF74FC3C96569EB5" blockId="9.[151,1436,226,1538]" box="[197,1099,992,1018]" pageId="9" pageNumber="46">
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This species was collected on rocks in shallow water (less than
|
||
<quantity id="107CC3A3FFB99334FC1CFC3C91ED9EB5" box="[941,1008,992,1018]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="46" unit="m" value="0.5">0.5 m</quantity>
|
||
depth).
|
||
</paragraph>
|
||
<paragraph id="D73B6E46FFB99334FF74FBD491EC998D" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
|
||
<emphasis id="E5F0B254FFB99334FF74FBD493D5996D" bold="true" box="[197,456,1032,1058]" pageId="9" pageNumber="46">Taxonomic remarks.</emphasis>
|
||
<taxonomicName id="108415C5FFB99334FE61FBD5906E996D" box="[464,627,1033,1058]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="hannelorae">
|
||
<emphasis id="E5F0B254FFB99334FE61FBD5906E996D" box="[464,627,1033,1058]" italics="true" pageId="9" pageNumber="46">T. hannelorae</emphasis>
|
||
</taxonomicName>
|
||
is the smallest species within the Malili lakes species flock. Its size in combination with the reticulate shell sculpture generally can serve to distinguish it unambiguously from all other species in the system. While
|
||
<taxonomicName id="108415C5FFB99334FD9CFB8590D3993D" box="[557,718,1113,1138]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="hannelorae">
|
||
<emphasis id="E5F0B254FFB99334FD9CFB8590D3993D" box="[557,718,1113,1138]" italics="true" pageId="9" pageNumber="46">T. hannelorae</emphasis>
|
||
</taxonomicName>
|
||
might be mistaken for subadult specimens of
|
||
<taxonomicName id="108415C5FFB99334FB58FB85902199D5" authority="Rintelen, Bouchet & Glaubrecht, 2007" authorityName="Rintelen, Bouchet & Glaubrecht" authorityYear="2007" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="confusa">
|
||
<emphasis id="E5F0B254FFB99334FB58FB85977D993D" box="[1257,1376,1113,1138]" italics="true" pageId="9" pageNumber="46">T. confusa</emphasis>
|
||
<bibRefCitation id="B31513B7FFB99334FAD9FB84902199D5" author="Rintelen" pageId="9" pageNumber="49" refString="Rintelen, T. von, Bouchet, P. & Glaubrecht, M. (2007) Ancient lakes as hotspots of diversity: a morphological review of an endemic species flock of Tylomelania (Caenogastropoda: Cerithioidea: Pachychilidae) in the Malili lake system on Sulawesi, Indonesia. Hydrobiologia, 592, 11 - 94." type="journal article" year="2007">Rintelen, Bouchet & Glaubrecht, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, at first glance, the shell corrosion of the upper whorls characteristic for older animals will allow identifying any specimen as a fully grown adult.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9F9E3DCDFFB99334FF74FB0C97209B4D" pageId="9" pageNumber="46" type="description">
|
||
<paragraph id="D73B6E46FFB99334FF74FB0C97209B4D" blockId="9.[151,1436,226,1538]" pageId="9" pageNumber="46">
|
||
<emphasis id="E5F0B254FFB99334FF74FB0C93C899A5" bold="true" box="[197,469,1232,1258]" pageId="9" pageNumber="46">Molecular phylogeny.</emphasis>
|
||
The topology of the 16S ML phylogram based on 851 bp of mitochondrial 16S rDNA (
|
||
<figureCitation id="4FBF72C3FFB99334FF43FB249324985D" box="[242,313,1272,1298]" captionStart="FIGURE 7" captionStartId="10.[151,254,1915,1939]" captionTargetBox="[185,1408,191,1837]" captionTargetId="figure@10.[151,1436,160,1891]" captionTargetPageId="10" captionText="FIGURE 7. Molecular phylogeny of Tylomelania. Maximum Likelihood phylogram based on 851 bp of mitochondrial 16 S rDNA (outgroup not shown). Malili lake species are marked by grey boxes, the position of the new species is indicated by a black bar. Numbers on branches are ML bootstrap values." httpUri="https://zenodo.org/record/183436/files/figure.png" pageId="9" pageNumber="46">Fig. 7</figureCitation>
|
||
) is basically identical to the tree presented in
|
||
<bibRefCitation id="B31513B7FFB99334FCECFB249646985D" box="[861,1115,1272,1298]" pageId="9" pageNumber="49" refString="Rintelen, T. von, Wilson, A. B., Meyer, A. & Glaubrecht, M. (2004) Escalation and trophic specialization drive adaptive radiation of viviparous freshwater gastropods in the ancient lakes on Sulawesi, Indonesia. Proceedings of the Royal Society of London B Biological Sciences, 271, 2541 - 2549." type="journal article">
|
||
Rintelen
|
||
<emphasis id="E5F0B254FFB99334FC79FB249619985E" box="[968,1028,1272,1297]" italics="true" pageId="9" pageNumber="46">et al.</emphasis>
|
||
(2004)
|
||
</bibRefCitation>
|
||
except for the inclusion of two of the new species described here,
|
||
<taxonomicName id="108415C5FFB99334FDD0FAFC90C39876" box="[609,734,1312,1337]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB99334FDD0FAFC90C39876" box="[609,734,1312,1337]" italics="true" pageId="9" pageNumber="46">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="108415C5FFB99334FCA7FAFC91D49876" box="[790,969,1312,1337]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="sinabartfeldi">
|
||
<emphasis id="E5F0B254FFB99334FCA7FAFC91D49876" box="[790,969,1312,1337]" italics="true" pageId="9" pageNumber="46">T. sinabartfeldi</emphasis>
|
||
</taxonomicName>
|
||
(all attempts to sequence
|
||
<taxonomicName id="108415C5FFB99334FB4BFAFC97869876" box="[1274,1435,1312,1337]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="hannelorae">
|
||
<emphasis id="E5F0B254FFB99334FB4BFAFC97869876" box="[1274,1435,1312,1337]" italics="true" pageId="9" pageNumber="46">T. hannelorae</emphasis>
|
||
</taxonomicName>
|
||
failed). Both species belong to a clade of smooth-shelled or spirally-ribbed taxa (clade Malili 2, compare
|
||
<bibRefCitation id="B31513B7FFB99334FAD6FA94934098C5" author="Rintelen" pageId="9" pageNumber="49" refString="Rintelen, T. von, Wilson, A. B., Meyer, A. & Glaubrecht, M. (2004) Escalation and trophic specialization drive adaptive radiation of viviparous freshwater gastropods in the ancient lakes on Sulawesi, Indonesia. Proceedings of the Royal Society of London B Biological Sciences, 271, 2541 - 2549." type="journal article" year="2004">
|
||
Rintelen
|
||
<emphasis id="E5F0B254FFB99334FF68FAAC930B98C6" box="[217,278,1392,1417]" italics="true" pageId="9" pageNumber="46">et al.</emphasis>
|
||
2004
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="B31513B7FFB99334FEDBFAAC93B498C5" author="Rintelen" box="[362,425,1392,1418]" pageId="9" pageNumber="49" refString="Rintelen, T. von, Bouchet, P. & Glaubrecht, M. (2007) Ancient lakes as hotspots of diversity: a morphological review of an endemic species flock of Tylomelania (Caenogastropoda: Cerithioidea: Pachychilidae) in the Malili lake system on Sulawesi, Indonesia. Hydrobiologia, 592, 11 - 94." type="journal article" year="2007">2007</bibRefCitation>
|
||
), where they form a poorly supported subclade which is sistergroup to all remaining species of clade Malili 2 but for one individual of
|
||
<taxonomicName id="108415C5FFB99334FD6EFA44964898FD" authority="Kruimel, 1913" authorityName="Kruimel" authorityYear="1913" box="[735,1109,1432,1458]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="sarasinorum">
|
||
<emphasis id="E5F0B254FFB99334FD6EFA44918D98FE" box="[735,912,1432,1457]" italics="true" pageId="9" pageNumber="46">T. sarasinorum</emphasis>
|
||
(
|
||
<bibRefCitation id="B31513B7FFB99334FC2EFA44965098FD" author="Kruimel" box="[927,1101,1432,1458]" pageId="9" pageNumber="49" refString="Kruimel, J. H. (1913) Verzeichnis der von Herrn E. C. Abendanon in Celebes gesammelten Susswasser-Mollusken. Bijdragen tot de Dierkunde, 19, 217 - 235." type="journal article" year="1913">Kruimel, 1913</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from the outlet bay of Lake Towuti clustering with
|
||
<taxonomicName id="108415C5FFB99334FE17FA1C903E9896" box="[422,547,1472,1497]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB99334FE17FA1C903E9896" box="[422,547,1472,1497]" italics="true" pageId="9" pageNumber="46">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="108415C5FFB99334FDEAFA1C91109896" box="[603,781,1472,1497]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="sinabartfeldi">
|
||
<emphasis id="E5F0B254FFB99334FDEAFA1C91109896" box="[603,781,1472,1497]" italics="true" pageId="9" pageNumber="46">T. sinabartfeldi</emphasis>
|
||
</taxonomicName>
|
||
. While the three haplotypes of
|
||
<taxonomicName id="108415C5FFB99334FBCDFA1C97339896" box="[1148,1326,1472,1497]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="sinabartfeldi">
|
||
<emphasis id="E5F0B254FFB99334FBCDFA1C97339896" box="[1148,1326,1472,1497]" italics="true" pageId="9" pageNumber="46">T. sinabartfeldi</emphasis>
|
||
</taxonomicName>
|
||
are identical,
|
||
<taxonomicName id="108415C5FFB99334FF69FA34934F9B4E" box="[216,338,1512,1537]" class="Gastropoda" family="Pachychilidae" genus="Tylomelania" kingdom="Animalia" pageId="9" pageNumber="46" phylum="Mollusca" rank="species" species="baskasti">
|
||
<emphasis id="E5F0B254FFB99334FF69FA34934F9B4E" box="[216,338,1512,1537]" italics="true" pageId="9" pageNumber="46">T. baskasti</emphasis>
|
||
</taxonomicName>
|
||
appears paraphyletic with an intraspecific sequence divergence of 0.6% (p-distance).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |