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<mods:title id="F249F5209F7736756B0D52578F67AD28">Cutting the ribbon: bathyal Nemertea from seeps along the Costa Rica margin, with descriptions of 2 new genera and 9 new species</mods:title>
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<mods:namePart id="EB51DFB453D1E2CD1913336EF269F8E3">Sagorny, Christina</mods:namePart>
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<mods:affiliation id="E4119739BAD932CBD04846423DF1D2AE">Institute of Evolutionary Biology and Animal Ecology, Rheinische Friedrich-Wilhelms-Universität Bonn, An der Immenburg 1, 53121 Bonn, Germany. &amp; Scripps Institution of Oceanography, 9500 Gilman Drive, La Jolla, California 92093 - 0202, USA.</mods:affiliation>
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<heading id="8C513E28FFADFFF0FD9EF9CCFC2C2E2F" box="[613,975,1593,1620]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="147" reason="2">
Genus
<taxonomicName id="10A6F2C7FFADFFF0FD4CF9CCFCBE2E2F" authority="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic, 2022" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[695,861,1593,1620]" class="Polychaeta" family="Hesionidae" genus="Chernyshevia" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="147" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="E5D25556FFADFFF0FD4CF9CCFCBE2E2F" bold="true" box="[695,861,1593,1620]" italics="true" pageId="15" pageNumber="147">Chernyshevia</emphasis>
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<taxonomicNameLabel id="FEE1E82DFFADFFF0FC9FF9CFFC2C2E2F" box="[868,975,1594,1620]" pageId="15" pageNumber="147" rank="genus">gen. nov.</taxonomicNameLabel>
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<typeStatus id="081D37E6FFADFFF0FF46F96BFF192EC3" box="[189,250,1694,1720]" pageId="15" pageNumber="147">Type</typeStatus>
species
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<taxonomicName id="10A6F2C7FFADFFF0FF46F939FDE42E9D" authority="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic, 2022" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[189,519,1740,1766]" class="Polychaeta" family="Hesionidae" genus="Chernyshevia" kingdom="Animalia" order="Phyllodocida" pageId="15" pageNumber="147" phylum="Annelida" rank="species" species="escarpiaphila" status="gen. et sp. nov.">
<emphasis id="E5D25556FFADFFF0FF46F939FDE42E9D" box="[189,519,1740,1766]" italics="true" pageId="15" pageNumber="147">Chernyshevia escarpiaphila</emphasis>
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<paragraph id="D7198944FFADFFF0FF46F8F8FED52F5C" blockId="15.[189,310,1805,1831]" box="[189,310,1805,1831]" pageId="15" pageNumber="147">
<heading id="8C513E28FFADFFF0FF46F8F8FED52F5C" bold="true" box="[189,310,1805,1831]" fontSize="11" level="3" pageId="15" pageNumber="147" reason="3">
<emphasis id="E5D25556FFADFFF0FF46F8F8FED52F5C" bold="true" box="[189,310,1805,1831]" pageId="15" pageNumber="147">Diagnosis</emphasis>
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<paragraph id="D7198944FFADFFF0FF46F8C9FEC62F9F" blockId="15.[187,1399,1852,2020]" pageId="15" pageNumber="147">Monostiliferan nemertean lacking eyes, with one pair of ventro-lateral cephalic furrows continuing as unforked ciliated canal; cerebral organs close in front of brain; rhynchocoel extending to posterior end of body; apical organ present; cephalic glands extending to brain; lateral nerve cords with small accessory nerves; mid-dorsal blood vessel without vascular plug; intestinal cecum with anterior diverticula; sexes separate.</paragraph>
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<caption id="83D9D9CCFFB2FFEFFF40F8EFFAE72F9F" ID-DOI="http://doi.org/10.5281/zenodo.7258870" ID-Zenodo-Dep="7258870" httpUri="https://zenodo.org/record/7258870/files/figure.png" pageId="16" pageNumber="148" startId="16.[187,230,1818,1844]" targetBox="[259,1337,273,1766]" targetPageId="16">
<paragraph id="D7198944FFB2FFEFFF40F8EFFAE72F9F" blockId="16.[187,1401,1818,2021]" pageId="16" pageNumber="148">
<emphasis id="E5D25556FFB2FFEFFF40F8EFFEEF2F4F" bold="true" box="[187,268,1818,1844]" pageId="16" pageNumber="148">Fig. 7.</emphasis>
Maximum likelihood (ML) tree for the monostiliferan supermatrix based on the concatenated four gene dataset (16S, COI, 18S, H3). Black circles indicate nodal support of 100/100/1.0; asterisks indicate nodal support of either 100/ or /1.0. Alternative branches found in the BI analysis are given as blue dotted lines.
<taxonomicName id="10A6F2C7FFB2FFEFFE70F871FB9E2FE4" authority="Chernyshev, Abukawa &amp; Kajihara, 2015" authorityName="Chernyshev, Abukawa &amp; Kajihara" authorityYear="2015" box="[395,1149,1924,1951]" class="Pilidiophora" family="Valenciniidae" genus="Sonnenemertes" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="148" phylum="Nemertea" rank="species" species="cantelli">
<emphasis id="E5D25556FFB2FFEFFE70F871FD782FE5" box="[395,667,1924,1950]" italics="true" pageId="16" pageNumber="148">Sonnenemertes cantelli</emphasis>
<bibRefCitation id="B337F4B5FFB2FFEFFD58F871FB9E2FE4" author="Chernyshev A. V. &amp; Abukawa S. &amp; Kajihara H." box="[675,1149,1924,1951]" pageId="16" pageNumber="148" pagination="119 - 127" refId="ref19531" refString="Chernyshev A. V., Abukawa S. &amp; Kajihara H. 2015. Sonnenemertes cantelli gen. et sp. nov. (Heteronemertea) - A new Oxypolella - like nemertean from the abyssal plain adjacent to the KurilKamchatka Trench. Deep Sea Research Part II: Topical Studies in Oceanography 111: 119 - 127. https: // doi. org / 10.1016 / j. dsr 2.2014.07.014" type="journal article" year="2015">Chernyshev, Abukawa &amp; Kajihara, 2015</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="10A6F2C7FFB2FFEFFB4EF870FE1B2FB9" authority="Sundberg &amp; Gibson, 1995" authorityName="Sundberg &amp; Gibson" authorityYear="1995" class="Pilidiophora" family="Lineidae" genus="Micrura" kingdom="Animalia" order="Heteronemertea" pageId="16" pageNumber="148" phylum="Nemertea" rank="species" species="callima">
<emphasis id="E5D25556FFB2FFEFFB4EF870FA952FE5" box="[1205,1398,1924,1950]" italics="true" pageId="16" pageNumber="148">Micrura callima</emphasis>
<bibRefCitation id="B337F4B5FFB2FFEFFF40F852FE1B2FB9" author="Sundberg P. &amp; Gibson R." box="[187,504,1959,1986]" pageId="16" pageNumber="148" pagination="101 - 141" refId="ref23975" refString="Sundberg P. &amp; Gibson R. 1995. The nemerteans (Nemertea) of Rottnest Island, Western Australia. Zoologica Scripta 24: 101 - 141. https: // doi. org / 10.1111 / j. 1463 - 6409.1995. tb 00395. x" type="journal article" year="1995">Sundberg &amp; Gibson, 1995</bibRefCitation>
</taxonomicName>
were used for outgroup rooting. Specimens belonging to new species are given in bold. Specimen-ID, sampling locality, and depth (for deep-sea species) are provided.
</paragraph>
</caption>
<subSubSection id="9FBCDACFFFB3FFEEFF46FEFCFAF42AF6" pageId="17" pageNumber="149" type="etymology">
<paragraph id="D7198944FFB3FFEEFF46FEFCFEA72958" blockId="17.[189,324,265,291]" box="[189,324,265,291]" pageId="17" pageNumber="149">
<heading id="8C513E28FFB3FFEEFF46FEFCFEA72958" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="17" pageNumber="149" reason="3">
<emphasis id="E5D25556FFB3FFEEFF46FEFCFEA72958" bold="true" box="[189,324,265,291]" pageId="17" pageNumber="149">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="D7198944FFB3FFEEFF46FEC3FDFC2908" blockId="17.[189,1399,310,371]" pageId="17" pageNumber="149">The genus is named after Dr Alexei Chernyshev to acknowledge his significant contributions to the study of deep-sea nemerteans.</paragraph>
<paragraph id="D7198944FFB3FFEEFF46FE6CFE2E29C8" blockId="17.[189,461,409,435]" box="[189,461,409,435]" pageId="17" pageNumber="149">
<heading id="8C513E28FFB3FFEEFF46FE6CFE2E29C8" bold="true" box="[189,461,409,435]" fontSize="11" level="3" pageId="17" pageNumber="149" reason="3">
<emphasis id="E5D25556FFB3FFEEFF46FE6CFE2E29C8" bold="true" box="[189,461,409,435]" pageId="17" pageNumber="149">Phylogenetic remarks</emphasis>
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</paragraph>
<paragraph id="D7198944FFB3FFEEFF46FE32FAF42AF6" blockId="17.[189,1399,455,653]" pageId="17" pageNumber="149">
The genus is sister to the represented specimen of
<taxonomicName id="10A6F2C7FFB3FFEEFCD3FE3DFB392999" authority="(Girard, 1852)" baseAuthorityName="Girard" baseAuthorityYear="1852" box="[808,1242,455,482]" class="Hoplonemertea" family="Tetrastemmatidae" genus="Tetrastemma" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="species" species="elegans">
<emphasis id="E5D25556FFB3FFEEFCD3FE3DFBC0299A" box="[808,1059,455,481]" italics="true" pageId="17" pageNumber="149">Tetrastemma elegans</emphasis>
(Girard, 1852)
</taxonomicName>
(
<figureCitation id="4F9D95C1FFB3FFEEFB16FE32FAD5299A" box="[1261,1334,455,481]" captionStart="Fig" captionStartId="16.[187,230,1818,1844]" captionTargetBox="[259,1337,273,1766]" captionTargetId="figure-12@16.[236,1349,266,1783]" captionTargetPageId="16" captionText="Fig. 7. Maximum likelihood (ML) tree for the monostiliferan supermatrix based on the concatenated four gene dataset (16S, COI, 18S, H3). Black circles indicate nodal support of 100/100/1.0; asterisks indicate nodal support of either 100/ or /1.0. Alternative branches found in the BI analysis are given as blue dotted lines. Sonnenemertes cantelli Chernyshev, Abukawa &amp; Kajihara, 2015 and Micrura callima Sundberg &amp; Gibson, 1995 were used for outgroup rooting. Specimens belonging to new species are given in bold. Specimen-ID, sampling locality, and depth (for deep-sea species) are provided." figureDoi="http://doi.org/10.5281/zenodo.7258870" httpUri="https://zenodo.org/record/7258870/files/figure.png" pageId="17" pageNumber="149">Fig. 7</figureCitation>
) that is in need of revision. This clade is sister to the monotypic genus
<taxonomicName id="10A6F2C7FFB3FFEEFC2BFE1FFF1F2A5D" authority="Junoy, Andrade &amp; Giribet, 2010" authorityName="Junoy, Andrade &amp; Giribet" authorityYear="2010" class="Hoplonemertea" genus="Vieitezia" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="genus">
<emphasis id="E5D25556FFB3FFEEFC2BFE1FFBD72A78" box="[976,1076,490,515]" italics="true" pageId="17" pageNumber="149">Vieitezia</emphasis>
<bibRefCitation id="B337F4B5FFB3FFEEFBC7FE1FFF1F2A5D" author="Junoy J. &amp; Andrade S. C. S. &amp; Giribet G." pageId="17" pageNumber="149" pagination="616 - 629" refId="ref20760" refString="Junoy J., Andrade S. C. S. &amp; Giribet G. 2010. Phylogenetic placement of a new hoplonemertean species commensal on ascidians. Invertebrate Systematics 24: 616 - 629. https: // doi. org / 10.1071 / IS 10036" type="journal article" year="2010">Junoy, Andrade &amp; Giribet, 2010</bibRefCitation>
</taxonomicName>
, including specimens of the
<typeStatus id="081D37E6FFB3FFEEFDABFDF9FD602A5D" box="[592,643,524,550]" pageId="17" pageNumber="149">type</typeStatus>
species
<taxonomicName id="10A6F2C7FFB3FFEEFD1CFDF8FA902A5D" authority="Junoy, Andrade &amp; Giribet, 2010" authorityName="Junoy, Andrade &amp; Giribet" authorityYear="2010" box="[743,1395,524,550]" class="Hoplonemertea" genus="Vieitezia" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="species" species="luzmurubeae">
<emphasis id="E5D25556FFB3FFEEFD1CFDF8FC0F2A5D" box="[743,1004,524,550]" italics="true" pageId="17" pageNumber="149">Vieitezia luzmurubeae</emphasis>
<bibRefCitation id="B337F4B5FFB3FFEEFC0FFDF9FA902A5D" author="Junoy J. &amp; Andrade S. C. S. &amp; Giribet G." box="[1012,1395,524,550]" pageId="17" pageNumber="149" pagination="616 - 629" refId="ref20760" refString="Junoy J., Andrade S. C. S. &amp; Giribet G. 2010. Phylogenetic placement of a new hoplonemertean species commensal on ascidians. Invertebrate Systematics 24: 616 - 629. https: // doi. org / 10.1071 / IS 10036" type="journal article" year="2010">Junoy, Andrade &amp; Giribet, 2010</bibRefCitation>
</taxonomicName>
, and specimens of
<taxonomicName id="10A6F2C7FFB3FFEEFE6EFDDAFC852A33" authority="(Burger, 1904)" baseAuthorityName="Burger" baseAuthorityYear="1904" box="[405,870,558,584]" class="Hoplonemertea" family="Tetrastemmatidae" genus="Tetrastemma" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="species" species="vittigerum">
<emphasis id="E5D25556FFB3FFEEFE6EFDDAFD4E2A33" box="[405,685,559,584]" italics="true" pageId="17" pageNumber="149">Tetrastemma vittigerum</emphasis>
(
<bibRefCitation id="B337F4B5FFB3FFEEFD46FDDBFCBE2A33" author="Burger O." box="[701,861,558,584]" pageId="17" pageNumber="149" pagination="20" refId="ref18900" refString="Burger O. 1904. Nemertini. In: Das Tierreich 20. Lieferung Platyhelminthes. R. Friedlander &amp; Sohn, Berlin." type="book chapter" year="1904">Bürger, 1904</bibRefCitation>
)
</taxonomicName>
. The genus is nested within a clade that also includes the new genus
<taxonomicName id="10A6F2C7FFB3FFEEFE24FDA4FD762A10" authority="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic, 2022" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[479,661,593,619]" class="Pilidiophora" family="Lineidae" genus="Alvinonemertes" kingdom="Animalia" order="Heteronemertea" pageId="17" pageNumber="149" phylum="Nemertea" rank="genus" status="gen. nov.">
<emphasis id="E5D25556FFB3FFEEFE24FDA4FD762A10" box="[479,661,593,619]" italics="true" pageId="17" pageNumber="149">Alvinonemertes</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE1E82DFFB3FFEEFD66FDA4FCEF2A10" box="[669,780,593,619]" pageId="17" pageNumber="149" rank="genus">gen. nov.</taxonomicNameLabel>
, as well as the represented specimens of the genera
<taxonomicName id="10A6F2C7FFB3FFEEFF44FD86FDD22AF6" authority="Bergendal, 1900" authorityName="Bergendal" authorityYear="1900" box="[191,561,627,653]" class="Hoplonemertea" family="Prosorhochmidae" genus="Gononemertes" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="genus">
<emphasis id="E5D25556FFB3FFEEFF44FD86FE892AF6" box="[191,362,627,653]" italics="true" pageId="17" pageNumber="149">Gononemertes</emphasis>
Bergendal, 1900
</taxonomicName>
and
<taxonomicName id="10A6F2C7FFB3FFEEFD91FD86FB522AF6" authority="Chernyshev &amp; Polyakova, 2019" authorityName="Chernyshev &amp; Polyakova" authorityYear="2019" box="[618,1201,627,653]" class="Enopla" family="Oerstediidae" genus="Galathenemertes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="genus">
<emphasis id="E5D25556FFB3FFEEFD91FD86FCD12AF6" box="[618,818,627,653]" italics="true" pageId="17" pageNumber="149">Galathenemertes</emphasis>
<bibRefCitation id="B337F4B5FFB3FFEEFCC2FD86FB522AF6" author="Chernyshev A. V. &amp; Polyakova N. E." box="[825,1201,627,653]" pageId="17" pageNumber="149" pagination="102148" refId="ref19479" refString="Chernyshev A. V. &amp; Polyakova N. E. 2019. Nemerteans from the deep-sea expedition KuramBio II with descriptions of three new hoplonemerteans from the Kuril-Kamchatka Trench. Progress in Oceanography 178: 102148. https: // doi. org / 10.1016 / j. pocean. 2019.102148" type="journal article" year="2019">Chernyshev &amp; Polyakova, 2019</bibRefCitation>
</taxonomicName>
(
<figureCitation id="4F9D95C1FFB3FFEEFB3BFD86FAEB2AF6" box="[1216,1288,627,653]" captionStart="Fig" captionStartId="16.[187,230,1818,1844]" captionTargetBox="[259,1337,273,1766]" captionTargetId="figure-12@16.[236,1349,266,1783]" captionTargetPageId="16" captionText="Fig. 7. Maximum likelihood (ML) tree for the monostiliferan supermatrix based on the concatenated four gene dataset (16S, COI, 18S, H3). Black circles indicate nodal support of 100/100/1.0; asterisks indicate nodal support of either 100/ or /1.0. Alternative branches found in the BI analysis are given as blue dotted lines. Sonnenemertes cantelli Chernyshev, Abukawa &amp; Kajihara, 2015 and Micrura callima Sundberg &amp; Gibson, 1995 were used for outgroup rooting. Specimens belonging to new species are given in bold. Specimen-ID, sampling locality, and depth (for deep-sea species) are provided." figureDoi="http://doi.org/10.5281/zenodo.7258870" httpUri="https://zenodo.org/record/7258870/files/figure.png" pageId="17" pageNumber="149">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9FBCDACFFFB3FFEEFF46FD41FC2A2C6D" pageId="17" pageNumber="149" type="discussion">
<paragraph id="D7198944FFB3FFEEFF46FD41FECC2AB5" blockId="17.[189,303,692,718]" box="[189,303,692,718]" pageId="17" pageNumber="149">
<heading id="8C513E28FFB3FFEEFF46FD41FECC2AB5" bold="true" box="[189,303,692,718]" fontSize="11" level="3" pageId="17" pageNumber="149" reason="3">
<emphasis id="E5D25556FFB3FFEEFF46FD41FECC2AB5" bold="true" box="[189,303,692,718]" pageId="17" pageNumber="149">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="D7198944FFB3FFEEFF46FD14FE942BD3" blockId="17.[188,1401,737,936]" pageId="17" pageNumber="149">
The genus
<taxonomicName id="10A6F2C7FFB3FFEEFEB9FD14FE002A80" authority="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic, 2022" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[322,483,737,763]" class="Polychaeta" family="Hesionidae" genus="Chernyshevia" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="149" phylum="Annelida" rank="genus" status="gen. nov.">
<emphasis id="E5D25556FFB3FFEEFEB9FD14FE002A80" box="[322,483,737,763]" italics="true" pageId="17" pageNumber="149">Chernyshevia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FEE1E82DFFB3FFEEFE10FD17FDB42A87" box="[491,599,738,764]" pageId="17" pageNumber="149" rank="genus">gen. nov.</taxonomicNameLabel>
is erected as new genus based on its phylogenetic position and the questionable position of different species of
<taxonomicName id="10A6F2C7FFB3FFEEFD37FCF0FC812B65" authorityName="Ehrenberg" authorityYear="1831" box="[716,866,773,798]" class="Hoplonemertea" family="Tetrastemmatidae" genus="Tetrastemma" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="genus">
<emphasis id="E5D25556FFB3FFEEFD37FCF0FC812B65" box="[716,866,773,798]" italics="true" pageId="17" pageNumber="149">Tetrastemma</emphasis>
</taxonomicName>
(
<taxonomicName id="10A6F2C7FFB3FFEEFC80FCF0FC122B65" baseAuthorityName="Girard" baseAuthorityYear="1852" box="[891,1009,772,798]" class="Hoplonemertea" family="Tetrastemmatidae" genus="Tetrastemma" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="species" species="elegans">
<emphasis id="E5D25556FFB3FFEEFC80FCF0FC122B65" box="[891,1009,772,798]" italics="true" pageId="17" pageNumber="149">T. elegans</emphasis>
</taxonomicName>
and
<taxonomicName id="10A6F2C7FFB3FFEEFBD2FCF0FB5D2B65" baseAuthorityName="Burger" baseAuthorityYear="1904" box="[1065,1214,773,798]" class="Hoplonemertea" family="Tetrastemmatidae" genus="Tetrastemma" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="species" species="vittigerum">
<emphasis id="E5D25556FFB3FFEEFBD2FCF0FB5D2B65" box="[1065,1214,773,798]" italics="true" pageId="17" pageNumber="149">T. vittigerum</emphasis>
</taxonomicName>
) that are nested within the same clade as
<taxonomicName id="10A6F2C7FFB3FFEEFE19FCD3FD602B3B" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[482,643,806,832]" class="Polychaeta" family="Hesionidae" genus="Chernyshevia" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="149" phylum="Annelida" rank="genus">
<emphasis id="E5D25556FFB3FFEEFE19FCD3FD602B3B" box="[482,643,806,832]" italics="true" pageId="17" pageNumber="149">Chernyshevia</emphasis>
</taxonomicName>
. Moreover, some morphological differences between specimens of the genera
<taxonomicName id="10A6F2C7FFB3FFEEFE9DFCBCFE292B19" authorityName="Junoy, Andrade &amp; Giribet" authorityYear="2010" box="[358,458,841,866]" class="Hoplonemertea" genus="Vieitezia" kingdom="Animalia" order="Monostilifera" pageId="17" pageNumber="149" phylum="Nemertea" rank="genus">
<emphasis id="E5D25556FFB3FFEEFE9DFCBCFE292B19" box="[358,458,841,866]" italics="true" pageId="17" pageNumber="149">Vieitezia</emphasis>
</taxonomicName>
and
<taxonomicName id="10A6F2C7FFB3FFEEFDFEFCBDFD452B19" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[517,678,840,866]" class="Polychaeta" family="Hesionidae" genus="Chernyshevia" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="149" phylum="Annelida" rank="genus">
<emphasis id="E5D25556FFB3FFEEFDFEFCBDFD452B19" box="[517,678,840,866]" italics="true" pageId="17" pageNumber="149">Chernyshevia</emphasis>
</taxonomicName>
, such as the absence or presence of an accessory nerve, the dimension of the cephalic glands, and the presence of separate sexes, might also support the presence of a new genus.
</paragraph>
<paragraph id="D7198944FFB3FFEEFF46FC3BFE6F2B93" blockId="17.[189,396,974,1000]" box="[189,396,974,1000]" pageId="17" pageNumber="149">
<heading id="8C513E28FFB3FFEEFF46FC3BFE6F2B93" bold="true" box="[189,396,974,1000]" fontSize="11" level="3" pageId="17" pageNumber="149" reason="3">
<emphasis id="E5D25556FFB3FFEEFF46FC3BFE6F2B93" bold="true" box="[189,396,974,1000]" pageId="17" pageNumber="149">Species included</emphasis>
</heading>
</paragraph>
<paragraph id="D7198944FFB3FFEEFF46FC09FC2A2C6D" blockId="17.[189,969,1020,1046]" box="[189,969,1020,1046]" pageId="17" pageNumber="149">
Only the
<typeStatus id="081D37E6FFB3FFEEFED1FC09FEBE2C6D" box="[298,349,1020,1046]" pageId="17" pageNumber="149">type</typeStatus>
species
<taxonomicName id="10A6F2C7FFB3FFEEFE3BFC09FCE92C6D" authorityName="Sagorny &amp; Döhren &amp; Rouse &amp; Tilic" authorityYear="2022" box="[448,778,1020,1046]" class="Polychaeta" family="Hesionidae" genus="Chernyshevia" kingdom="Animalia" order="Phyllodocida" pageId="17" pageNumber="149" phylum="Annelida" rank="species" species="escarpiaphila">
<emphasis id="E5D25556FFB3FFEEFE3BFC09FCE92C6D" box="[448,778,1020,1046]" italics="true" pageId="17" pageNumber="149">Chernyshevia escarpiaphila</emphasis>
</taxonomicName>
gen. et. sp. nov.
</paragraph>
</subSubSection>
</treatment>
</document>