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<document id="B7ED6D6161116B3A18EEE1F79F0F5CBF" ID-DOI="10.1111/zoj.12294" ID-ISSN="0024-4082" ID-Zenodo-Dep="5339511" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630184182576" checkinUser="felipe" docAuthor="Mallon, Jordan C., Ryan, Michael J. &amp; Campbell, James A." docDate="2015" docId="5E4887BCFFA9FFEFFF7DF8BCC3C8F947" docLanguage="en" docName="zoj12294.pdf" docOrigin="Zoological Journal of the Linnean Society 175 (4)" docSource="http://dx.doi.org/10.1111/zoj.12294" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Arrhinoceratops brachyops Parks 1925" docType="treatment" docVersion="3" lastPageNumber="911" masterDocId="A271FFC4FFA8FFEEFFB8FFA8C77AFF9A" masterDocTitle="Skull ontogeny in Arrhinoceratops brachyops (Ornithischia: Ceratopsidae) and other horned dinosaurs" masterLastPageNumber="929" masterPageNumber="910" pageNumber="911" updateTime="1699178607014" updateUser="plazi">
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<mods:title id="712F5AB043A2F0FEF9BFAB73E3E24F83">Skull ontogeny in Arrhinoceratops brachyops (Ornithischia: Ceratopsidae) and other horned dinosaurs</mods:title>
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<mods:namePart id="5A6B87AA7B2E8FBAAB85D803998BBD79">Ryan, Michael J.</mods:namePart>
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<treatment id="5E4887BCFFA9FFEFFF7DF8BCC3C8F947" LSID="urn:lsid:plazi:treatment:5E4887BCFFA9FFEFFF7DF8BCC3C8F947" httpUri="http://treatment.plazi.org/id/5E4887BCFFA9FFEFFF7DF8BCC3C8F947" lastPageNumber="911" pageId="1" pageNumber="911">
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<paragraph id="D65E36AAFFA9FFEFFF7DF8BCC5BBF8B3" blockId="1.[197,705,1811,1833]" box="[197,705,1811,1833]" pageId="1" pageNumber="911">
<heading id="8D1681C6FFA9FFEFFF7DF8BCC5BBF8B3" box="[197,705,1811,1833]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="911" reason="8">
<taxonomicName id="11E14D29FFA9FFEFFF7DF8BCC55DF8BD" authorityName="Parks" authorityYear="1925" box="[197,551,1812,1833]" class="Reptilia" family="Ceratopsidae" genus="Arrhinoceratops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="929" phylum="Chordata" rank="species" species="brachyops">
<emphasis id="E495EAB8FFA9FFEFFF7DF8BCC55DF8BD" box="[197,551,1812,1833]" italics="true" pageId="1" pageNumber="911">ARRHINOCERATOPS BRACHYOPS</emphasis>
</taxonomicName>
<bibRefCitation id="B2704B5BFFA9FFEFFD88F8BCC5BBF8B3" author="Parks WA" box="[560,705,1811,1833]" pageId="1" pageNumber="911" pagination="1 - 15" refId="ref11507" refString="Parks WA. 1925. Arrhinoceratops brachyops, a new genus and species of Ceratopsia from the Edmonton Formation of Alberta. University of Toronto Studies, Geology Series 19: 1 - 15." type="book chapter" year="1925">PARKS, 1925</bibRefCitation>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9EFB6521FFA9FFEFFF28F892C429F99F" pageId="1" pageNumber="911" type="materials_examined">
<paragraph id="D65E36AAFFA9FFEFFF28F892C669F8F5" blockId="1.[144,758,1850,1903]" pageId="1" pageNumber="911">
<materialsCitation id="66893CF7FFA9FFEFFF28F892C669F8F5" collectionCode="ROM" pageId="1" pageNumber="911" specimenCode="ROM 796" specimenCount="1" typeStatus="holotype">
<emphasis id="E495EAB8FFA9FFEFFF28F892C610F8CA" box="[144,362,1850,1872]" italics="true" pageId="1" pageNumber="911">
<typeStatus id="095A8808FFA9FFEFFF28F892C789F8D5" box="[144,243,1850,1871]" pageId="1" pageNumber="911" type="holotype">Holotype</typeStatus>
specimen:
</emphasis>
<specimenCode id="86479ED1FFA9FFEFFECCF892C69FF8CA" box="[372,485,1850,1872]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="911" type="Museum">ROM 796</specimenCode>
, complete skull lacking lower jaws.
</materialsCitation>
</paragraph>
<paragraph id="D65E36AAFFA9FFEFFC9EFF6EC200FE0E" blockId="1.[806,1422,198,404]" pageId="1" pageNumber="911">
<materialsCitation id="66893CF7FFA9FFEFFC9EFF6EC200FE0E" collectionCode="ROM" county="Five" location="Red Deer River" municipality="Bleriot Ferry" pageId="1" pageNumber="911" specimenCode="ROM 796" specimenCount="1" stateProvince="Alberta" typeStatus="holotype">
<emphasis id="E495EAB8FFA9FFEFFC9EFF6EC3E8FF41" box="[806,1170,198,219]" italics="true" pageId="1" pageNumber="911">
<typeStatus id="095A8808FFA9FFEFFC9EFF6EC4F6FF41" box="[806,908,198,219]" pageId="1" pageNumber="911" type="holotype">Holotype</typeStatus>
locality and horizon:
</emphasis>
<collectingCounty id="3F3F4E26FFA9FFEFFB27FF6EC3A8FF46" box="[1183,1234,198,220]" pageId="1" pageNumber="911">Five</collectingCounty>
kilometres upstream of
<collectingMunicipality id="363AACD0FFA9FFEFFC37FF4CC361FF60" box="[911,1051,228,250]" pageId="1" pageNumber="911">Bleriot Ferry</collectingMunicipality>
, along the
<location id="D33E6071FFA9FFEFFB32FF4CC255FF60" LSID="urn:lsid:plazi:treatment:5E4887BCFFA9FFEFFF7DF8BCC3C8F947:D33E6071FFA9FFEFFB32FF4CC255FF60" box="[1162,1327,228,250]" county="Five" municipality="Bleriot Ferry" name="Red Deer River" pageId="1" pageNumber="911" stateProvince="Alberta">Red Deer River</location>
,
<collectingRegion id="1425F848FFA9FFEFFA8DFF4CC2F0FF60" box="[1333,1418,228,250]" country="Canada" name="Alberta" pageId="1" pageNumber="911">Alberta</collectingRegion>
.
<specimenCode id="86479ED1FFA9FFEFFC9EFEABC4ECFE83" box="[806,918,259,281]" collectionCode="ROM" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="911" type="Museum">ROM 796</specimenCode>
is from between the no. 8 and no. 10 coal seams, below the DMT, and within a narrow stratigraphic range spanning the upper half of the Horsethief Member to the lower half of the Morrin Member of the Horseshoe Canyon Formation (
<bibRefCitation id="B2704B5BFFA9FFEFFB28FED5C214FE0E" author="Mallon JC &amp; Holmes R &amp; Anderson JS &amp; Farke AA &amp; Evans DC" box="[1168,1390,381,404]" pageId="1" pageNumber="911" pagination="618 - 634" refId="ref11267" refString="Mallon JC, Holmes R, Anderson JS, Farke AA, Evans DC. 2014. New information on the rare horned dinosaur Arrhinoceratops brachyops (Ornithischia: Ceratopsidae) from the Upper Cretaceous of Alberta, Canada. Canadian Journal of Earth Sciences 51: 618 - 634." type="journal article" year="2014">
Mallon
<emphasis id="E495EAB8FFA9FFEFFB53FED6C261FE08" box="[1259,1307,381,403]" italics="true" pageId="1" pageNumber="911">et al</emphasis>
., 2014
</bibRefCitation>
).
</materialsCitation>
</paragraph>
<paragraph id="D65E36AAFFA9FFEFFC9EFE14C37CFD10" blockId="1.[806,1422,444,650]" pageId="1" pageNumber="911">
<emphasis id="E495EAB8FFA9FFEFFC9EFE14C373FE48" box="[806,1033,444,466]" italics="true" pageId="1" pageNumber="911">Referred specimens:</emphasis>
<collectionCode id="B0F0AE6FFFA9FFEFFBAAFE14C335FE4B" box="[1042,1103,444,465]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="1" pageNumber="911" type="Museum">CMN</collectionCode>
8882, partial skull of a juvenile, including dentaries with prominent lateral ridge;
<collectionCode id="B0F0AE6FFFA9FFEFFC9EFE51C418FD94" box="[806,866,505,526]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34778" lsid="urn:lsid:biocol.org:col:34778" name="Royal Ontario Museum" pageId="1" pageNumber="911" type="Museum">ROM</collectionCode>
1439, an almost complete skull with lower jaws, syncervical and left forelimb elements (scapula, humerus, radius, metacarpals and several phalanges);
<collectionCode id="B0F0AE6FFFA9FFEFFAB2FD9FC23CFDD6" box="[1290,1350,567,588]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34935" lsid="urn:lsid:biocol.org:col:34935" name="Canadian Museum of Nature" pageId="1" pageNumber="911" type="Museum">CMN</collectionCode>
56500 (field #WL 123), nasal horncore with triangular transverse cross section.
</paragraph>
<paragraph id="D65E36AAFFA9FFEFFC9EFD1AC252FD7D" blockId="1.[806,1421,690,743]" pageId="1" pageNumber="911">
<emphasis id="E495EAB8FFA9FFEFFC9EFD1AC3EAFD52" box="[806,1168,690,712]" italics="true" pageId="1" pageNumber="911">Tentatively referred specimens:</emphasis>
<collectionCode id="B0F0AE6FFFA9FFEFFB24FD1BC3AEFD52" box="[1180,1236,691,712]" country="South Africa" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="1" pageNumber="911">TMP</collectionCode>
1981.048.0006, partial frill with low, blunt epi-ossifications.
</paragraph>
<paragraph id="D65E36AAFFA9FFEFFC9EFCB8C3BAFC25" blockId="1.[806,1421,784,959]" pageId="1" pageNumber="911">
<emphasis id="E495EAB8FFA9FFEFFC9EFCB8C4BBFCBF" box="[806,961,784,805]" italics="true" pageId="1" pageNumber="911">Distribution:</emphasis>
From the upper Horsethief Member through the Morrin Member up to the DMT of the Horseshoe Canyon Formation,
<collectingRegion id="1425F848FFA9FFEFFB3DFCE5C3A6FCF9" box="[1157,1244,845,867]" country="Canada" name="Alberta" pageId="1" pageNumber="911">Alberta</collectingRegion>
,
<collectingCountry id="AEF6763AFFA9FFEFFB50FCE5C245FCF9" box="[1256,1343,845,867]" name="Canada" pageId="1" pageNumber="911">Canada</collectingCountry>
(
<emphasis id="E495EAB8FFA9FFEFFAF7FCE6C2F7FCF9" box="[1359,1421,846,867]" italics="true" pageId="1" pageNumber="911">sensu</emphasis>
<bibRefCitation id="B2704B5BFFA9FFEFFC9EFCC3C345FC18" author="Eberth DA &amp; Braman DR" box="[806,1087,875,898]" pageId="1" pageNumber="911" pagination="1053 - 1086" refId="ref9917" refString="Eberth DA, Braman DR. 2012. A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains. Canadian Journal of Earth Sciences 49: 1053 - 1086." type="journal article" year="2012">Eberth &amp; Braman, 2012</bibRefCitation>
).
<collectionCode id="B0F0AE6FFFA9FFEFFBEAFCC4C3F0FC1B" box="[1106,1162,876,897]" country="South Africa" httpUri="http://grbio.org/cool/4q52-xvg6" name="Transvaal Museum" pageId="1" pageNumber="911">TMP</collectionCode>
1981.048.0006 is from the Horseshoe Canyon Formation (unknown horizon) near Devon,
<collectingRegion id="1425F848FFA9FFEFFC03FC01C36BFC25" box="[955,1041,937,959]" country="Canada" name="Alberta" pageId="1" pageNumber="911">Alberta</collectingRegion>
(
<bibRefCitation id="B2704B5BFFA9FFEFFB9BFC01C3CEFC25" author="Tanke DH" box="[1059,1204,937,959]" pageId="1" pageNumber="911" pagination="19 - 30" refId="ref12027" refString="Tanke DH. 1984. Dinosaurs of the Devon area with reference to a large hadrosaur femur. Fossils Quarterly 3: 19 - 30." type="journal article" year="1984">Tanke, 1984</bibRefCitation>
).
</paragraph>
<paragraph id="D65E36AAFFA9FFEFFC9EFC40C429F99F" blockId="1.[806,1422,999,1541]" pageId="1" pageNumber="911">
<emphasis id="E495EAB8FFA9FFEFFC9EFC40C42CFC67" box="[806,854,1000,1021]" italics="true" pageId="1" pageNumber="911">Age:</emphasis>
The base of the Horseshoe Canyon Formation near Drumheller has been assigned an age of approximately 72.5 Ma (
<bibRefCitation id="B2704B5BFFA9FFEFFC1CFB8DC3D4FBA0" author="Eberth DA &amp; Braman DR" box="[932,1198,1061,1083]" pageId="1" pageNumber="911" pagination="1053 - 1086" refId="ref9917" refString="Eberth DA, Braman DR. 2012. A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains. Canadian Journal of Earth Sciences 49: 1053 - 1086." type="journal article" year="2012">Eberth &amp; Braman, 2012</bibRefCitation>
;
<bibRefCitation id="B2704B5BFFA9FFEFFB0FFB8DC2F8FBA1" author="Eberth DA &amp; Evans DC &amp; Brinkman DB &amp; Therrien F &amp; Tanke DH &amp; Russell LS" box="[1207,1410,1061,1083]" pageId="1" pageNumber="911" pagination="701 - 726" refId="ref9956" refString="Eberth DA, Evans DC, Brinkman DB, Therrien F, Tanke DH, Russell LS. 2013. Dinosaur biostratigraphy of the Edmonton Group (Upper Cretaceous), Alberta, Canada: evidence for climate influence. Canadian Journal of Earth Sciences 50: 701 - 726." type="journal article" year="2013">
Eberth
<emphasis id="E495EAB8FFA9FFEFFAB1FB8DC24DFBA0" box="[1289,1335,1061,1082]" italics="true" pageId="1" pageNumber="911">et al</emphasis>
., 2013
</bibRefCitation>
). The younger DMT in the middle of the formation lies just above the inferred position of the Campanian Maastrichtian boundary (
<bibRefCitation id="B2704B5BFFA9FFEFFB83FB28C23BFB0C" author="Eberth DA &amp; Braman DR" box="[1083,1345,1152,1174]" pageId="1" pageNumber="911" pagination="1053 - 1086" refId="ref9917" refString="Eberth DA, Braman DR. 2012. A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains. Canadian Journal of Earth Sciences 49: 1053 - 1086." type="journal article" year="2012">Eberth &amp; Braman, 2012</bibRefCitation>
, fig. 4), with an accepted age of 70.6 Ma (see discussion in
<bibRefCitation id="B2704B5BFFA9FFEFFAF8FB37C48DFB49" author="Eberth DA &amp; Braman DR" pageId="1" pageNumber="911" pagination="1053 - 1086" refId="ref9917" refString="Eberth DA, Braman DR. 2012. A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains. Canadian Journal of Earth Sciences 49: 1053 - 1086." type="journal article" year="2012">Eberth &amp; Braman, 2012</bibRefCitation>
;
<bibRefCitation id="B2704B5BFFA9FFEFFBBFFB16C38BFB4E" author="Eberth DA &amp; Evans DC &amp; Brinkman DB &amp; Therrien F &amp; Tanke DH &amp; Russell LS" box="[1031,1265,1214,1236]" pageId="1" pageNumber="911" pagination="701 - 726" refId="ref9956" refString="Eberth DA, Evans DC, Brinkman DB, Therrien F, Tanke DH, Russell LS. 2013. Dinosaur biostratigraphy of the Edmonton Group (Upper Cretaceous), Alberta, Canada: evidence for climate influence. Canadian Journal of Earth Sciences 50: 701 - 726." type="journal article" year="2013">
Eberth
<emphasis id="E495EAB8FFA9FFEFFBDEFB16C3E3FB49" box="[1126,1177,1214,1235]" italics="true" pageId="1" pageNumber="911">et al</emphasis>
., 2013
</bibRefCitation>
). The stratigraphic interval that extends upwards from the base of the formation to the DMT is approximately
<quantity id="11199B4FFFA9FFEFFAFFFB53C2F7FA8A" box="[1351,1421,1275,1297]" metricMagnitude="2" metricUnit="m" metricValue="1.65" pageId="1" pageNumber="911" unit="m" value="165.0">165 m</quantity>
thick (
<bibRefCitation id="B2704B5BFFA9FFEFFCC8FAB1C3F3FAB5" author="Eberth DA &amp; Braman DR" box="[880,1161,1305,1327]" pageId="1" pageNumber="911" pagination="1053 - 1086" refId="ref9917" refString="Eberth DA, Braman DR. 2012. A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains. Canadian Journal of Earth Sciences 49: 1053 - 1086." type="journal article" year="2012">Eberth &amp; Braman, 2012</bibRefCitation>
: fig. 4), and suggests a rate of sedimentation of
<quantity id="11199B4FFFA9FFEFFBE7FA90C3D6FAD7" box="[1119,1196,1336,1357]" metricMagnitude="-2" metricUnit="m" metricValue="8.7" pageId="1" pageNumber="911" unit="cm" value="8.7">8.7 cm</quantity>
kyr
<superScript id="21949BE2FFA9FFEFFB66FA9EC394FAD8" attach="left" box="[1246,1262,1334,1346]" fontSize="5" pageId="1" pageNumber="911">1</superScript>
. Identifiable
<taxonomicName id="11E14D29FFA9FFEFFC9EFAFFC4A7FAF6" authorityName="PARKS" authorityYear="1925" box="[806,989,1367,1388]" class="Reptilia" family="Ceratopsidae" genus="Arrhinoceratops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ornithischia" pageId="1" pageNumber="911" phylum="Chordata" rank="genus">
<emphasis id="E495EAB8FFA9FFEFFC9EFAFFC4A7FAF6" box="[806,989,1367,1388]" italics="true" pageId="1" pageNumber="911">Arrhinoceratops</emphasis>
</taxonomicName>
remains are known to occur through a stratigraphic interval of
<quantity id="11199B4FFFA9FFEFFB85FADDC309FA11" box="[1085,1139,1397,1419]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="1" pageNumber="911" unit="m" value="75.0">75 m</quantity>
that extends down section from the base of the DMT (
<bibRefCitation id="B2704B5BFFA9FFEFFBD1FA3CC23CFA30" author="Eberth DA &amp; Evans DC &amp; Brinkman DB &amp; Therrien F &amp; Tanke DH &amp; Russell LS" box="[1129,1350,1428,1450]" pageId="1" pageNumber="911" pagination="701 - 726" refId="ref9956" refString="Eberth DA, Evans DC, Brinkman DB, Therrien F, Tanke DH, Russell LS. 2013. Dinosaur biostratigraphy of the Edmonton Group (Upper Cretaceous), Alberta, Canada: evidence for climate influence. Canadian Journal of Earth Sciences 50: 701 - 726." type="journal article" year="2013">
Eberth
<emphasis id="E495EAB8FFA9FFEFFB7BFA3DC389FA33" box="[1219,1267,1428,1450]" italics="true" pageId="1" pageNumber="911">et al</emphasis>
., 2013
</bibRefCitation>
). This suggests a temporal range of approximately 860 kyr for that taxon in the region, between 71.5 and 70.6 Ma.
</paragraph>
</subSubSection>
<subSubSection id="9EFB6521FFA9FFEFFC21F994C3C8F947" pageId="1" pageNumber="911" type="description">
<paragraph id="D65E36AAFFA9FFEFFC21F994C260F9CE" blockId="1.[921,1306,1596,1620]" box="[921,1306,1596,1620]" pageId="1" pageNumber="911">DESCRIPTION OF CMN 8882</paragraph>
<paragraph id="D65E36AAFFA9FFEFFC9EF9C4C3C8F947" blockId="1.[806,1422,1644,1757]" pageId="1" pageNumber="911">
As reconstructed (
<figureCitation id="4EDA2A2FFFA9FFEFFC4EF9C4C34FF918" box="[1014,1077,1644,1666]" captionStart="Figure 1" captionStartId="2.[164,239,1083,1102]" captionTargetBox="[167,778,648,1052]" captionTargetId="figure-362@2.[164,780,646,1054]" captionTargetPageId="2" captionText="Figure 1. Skull reconstruction of CMN 8882 (Arrhinoceratops brachyops) in left lateral view. Missing portions are reconstructed with dashed lines." figureDoi="http://doi.org/10.5281/zenodo.5339513" httpUri="https://zenodo.org/record/5339513/files/figure.png" pageId="1" pageNumber="911">Fig. 1</figureCitation>
), the skull has a basal length (rostral to occipital condyle) of approximately
<quantity id="11199B4FFFA9FFEFFA94F922C2F3F93A" box="[1324,1417,1674,1696]" metricMagnitude="-1" metricUnit="m" metricValue="4.65" pageId="1" pageNumber="911" unit="mm" value="465.0">465 mm</quantity>
, and is
<quantity id="11199B4FFFA9FFEFFCCFF901C4A9F924" box="[887,979,1705,1726]" metricMagnitude="-1" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="911" unit="mm" value="900.0">900 mm</quantity>
in total length. Other relevant measurements are provided in
<tableCitation id="9B630311FFA9FFEFFBE1F96FC3C8F947" box="[1113,1202,1735,1757]" captionStart="Table 1" captionStartId="2.[164,228,1211,1230]" captionText="Table 1. Skull measurements for CMN 8882 (Arrhinoceratops brachyops)" pageId="1" pageNumber="911">Table 1.</tableCitation>
</paragraph>
</subSubSection>
</treatment>
</document>