treatments-xml/data/5D/A7/2A/5DA72A4401B052989412F1C115CC0853.xml

<document id="AC28AF0BEDDFC8695766EE99498617D3" ID-CLB-Dataset="26391" ID-DOI="http://dx.doi.org/10.3897/evolsyst.3.33417" ID-GBIF-Dataset="5c22a401-90a8-4f21-b461-4acddee5378c" ID-Pensoft-Pub="2535-0730-1-75" ID-ZooBank="87CCD4C89F21471C85DD2AECD896FC1D" ModsDocAuthor="" ModsDocDate="2019" ModsDocID="2535-0730-1-75" ModsDocOrigin="Evolutionary Systematics 3 (1)" ModsDocTitle="A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar" checkinTime="1561619016944" checkinUser="pensoft" docAuthor="Scherz, Mark D., Koehler, Joern, Vences, Miguel &amp; Glaw, Frank" docDate="2019" docId="5DA72A4401B052989412F1C115CC0853" docLanguage="en" docName="EvolutSyst 3(1): 75-83" docOrigin="Evolutionary Systematics 3 (1)" docSource="http://dx.doi.org/10.3897/evolsyst.3.33417" docTitle="Platypelis ando Scherz, Koehler, Vences &amp; Glaw, 2019, sp. nov." docType="treatment" docUuid="386749F6-1762-4DCF-9560-72C5347331FC" docUuidSource="ZooBank" docVersion="6" lastPageNumber="75" masterDocId="FFA41A6FFFB1994EFFE9FFBAED391818" masterDocTitle="A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar" masterLastPageNumber="83" masterPageNumber="75" pageNumber="75" updateTime="1701376499090" updateUser="ExternalLinkService">
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<mods:title id="ECB4580BC46F29B67ABC3DD81991F518">A new yellow-toed Platypelis species (Anura, Microhylidae, Cophylinae) from the Maroantsetra region, northeastern Madagascar</mods:title>
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<mods:namePart id="98FB6B1B559300A15B3BCEA500C3D2BA">Scherz, Mark D.</mods:namePart>
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<mods:namePart id="A7F2F815041ABDBAA0A4FB7B7DAC5105">Koehler, Joern</mods:namePart>
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<mods:namePart id="65C0B3715BFDBA611FB2543B8169C9B3">Vences, Miguel</mods:namePart>
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<mods:namePart id="1E35C954BDDAA237DADD4A44AC2C90E9">Glaw, Frank</mods:namePart>
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<subSubSection id="109B38E13EAAEF4FF3CB740DC397A069" pageId="0" pageNumber="75" type="nomenclature">
<paragraph id="B3CCC25B266FBDA2105B5BE28EFA67CC" pageId="0" pageNumber="75">
<taxonomicName id="4F45E6CEC901F31F27DB2C05F0ABDD89" LSID="http://zoobank.org/386749F6-1762-4DCF-9560-72C5347331FC" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
<taxonomicNameLabel id="F43DC0310796908BE110298E472009EF" pageId="0" pageNumber="75">sp. nov.</taxonomicNameLabel>
Figs 1, 2, 3, 4, Table 1
</paragraph>
</subSubSection>
<subSubSection id="35033BE27062DFB7A7CA66D940D323C5" pageId="0" pageNumber="75" type="holotype">
<paragraph id="521A7D1C58D607E6F0322D1A123E8B71" pageId="0" pageNumber="75">Holotype.</paragraph>
<paragraph id="62C5A63F64D86ED9B7AFFE4A079C3B27" pageId="0" pageNumber="75">
ZSM 293/2010 (FGZC 4285), adult male, collected on 3 April 2010 in Ambodivoangy (
<geoCoordinate id="9235915BF20409CC9E1A1CE78CAAAA12" direction="south" orientation="latitude" precision="5" value="-15.2899">15.2899S</geoCoordinate>
, 
<geoCoordinate id="E49018E8CF69EEF2323C3BA1191C4152" direction="east" orientation="longitude" precision="5" value="49.6203">49.6203E</geoCoordinate>
, ca. 100 m a.s.l.), Analanjirofo Region, northeastern Madagascar, by P.-S. Gehring, F. Glaw, J. 
<normalizedToken id="C7A3470930A281763E04D15D8EDB424C" originalValue="Köhler">Koehler</normalizedToken>
, M. Pabijan, and F. M. Ratsoavina.
</paragraph>
</subSubSection>
<subSubSection id="41624C9F6B9FDA77170CAC87BD86F20E" pageId="0" pageNumber="75" type="paratypes">
<paragraph id="DFC5B5A222A871D5A992EC2C033647AB" pageId="0" pageNumber="75">Paratypes.</paragraph>
<paragraph id="AD14E0386C90811B36B35972D1EFC38C" pageId="0" pageNumber="75">ZSM 291/2010 (FGZC 4200), adult male, and ZSM 292/2010 (FGZC 4226), probably a male, collected on 31 March 2010 from the same locality as the holotype by the same collectors.</paragraph>
</subSubSection>
<subSubSection id="C3E37E1CF8117276205F0F796500C010" pageId="0" pageNumber="75" type="diagnosis">
<paragraph id="1BA31C42FD334F0C830857CA8FAE7E50" pageId="0" pageNumber="75">Diagnosis.</paragraph>
<paragraph id="69309AE7F2E5F1BA67CEF1DAFC826D6D" pageId="0" pageNumber="75">
The new species is assigned to the genus 
<taxonomicName id="74D800BB0E50A32F18B176282AE823C7" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
based on molecular phylogenetic relationships (Fig. 1). 
<taxonomicName id="0BEE9625CA824CE8BA50B9051FE3003F" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
sp. nov. is characterised by the following combination of characters: (1) Small size, with adult male SVL 16.9-18.7 mm; (2) manus with second finger shorter than fourth, pes with fifth toe shorter than third; (3) discs of fingers and toes yellowish to orangish in life; (4) presence of a dark dorsal chevron; (5) presence of dorsal tubercles; (6) short supratympanic dark brown marking; (7) males with prepollical tubercle but lacking a finger-like prepollex as typical for 
<taxonomicName id="E286D7807F7507458DBBD96B6A1C4772" class="Amphibia" family="Microhylidae" genus="Anodonthyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anodonthyla" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Anodonthyla</taxonomicName>
<normalizedToken id="47473B6DEB6CEC75A52D22383676DE30" originalValue="Müller">Mueller</normalizedToken>
, 1892.
</paragraph>
<paragraph id="4A609CCA4E392BA08737A857C512B1EB" pageId="0" pageNumber="75">
The new species is distinguished from 
<taxonomicName id="E2C65BE5ACED8C947E2C6F7AD10574B0" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis cowanii" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="cowanii">Platypelis cowanii</taxonomicName>
, 
<taxonomicName id="87B4254A1EA772AD6C4812F60D5313D9" lsidName="P. mavomavo" pageId="0" pageNumber="75" rank="species" species="mavomavo">P. mavomavo</taxonomicName>
, 
<taxonomicName id="DB1D99D689881224EA8BDE5A6D5ED949" lsidName="P. grandis" pageId="0" pageNumber="75" rank="species" species="grandis">P. grandis</taxonomicName>
, 
<taxonomicName id="6F042F92CE582FC133014F3DD27B5561" lsidName="P. tsaratananaensis" pageId="0" pageNumber="75" rank="species" species="tsaratananaensis">P. tsaratananaensis</taxonomicName>
, 
<taxonomicName id="49F69F9DF9A0A1B3085F9AB96C04D851" lsidName="P. pollicaris" pageId="0" pageNumber="75" rank="species" species="pollicaris">P. pollicaris</taxonomicName>
, 
<taxonomicName id="093F1BDF73AA1B89257D50DB02FE112C" lsidName="P. alticola" pageId="0" pageNumber="75" rank="species" species="alticola">P. alticola</taxonomicName>
, 
<taxonomicName id="BC0A424A13AC8BF51E3D6A14E9693336" lsidName="P. olgae" pageId="0" pageNumber="75" rank="species" species="olgae">P. olgae</taxonomicName>
, 
<taxonomicName id="3A5EC8960912B74AD4CE19ADE030AC82" lsidName="P. tuberifera" pageId="0" pageNumber="75" rank="species" species="tuberifera">P. tuberifera</taxonomicName>
, 
<taxonomicName id="FB021B83CDE7F9C3E655004086BAB8CB" lsidName="P. barbouri" pageId="0" pageNumber="75" rank="species" species="barbouri">P. barbouri</taxonomicName>
and 
<taxonomicName id="AB446CA495725C8FB1FE993727285B97" lsidName="P. milloti" pageId="0" pageNumber="75" rank="species" species="milloti">P. milloti</taxonomicName>
by considerably smaller size (16.9-18.7 vs&gt;25 mm). Among 
<taxonomicName id="0BA7E9A6E9E3689BDD68FE4AF22D8C98" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
species of similar size, it can be distinguished from 
<taxonomicName id="7D0452E3C5F45CBF4E6FC572A7ACDFB5" lsidName="P. tetra" pageId="0" pageNumber="75" rank="species" species="tetra">P. tetra</taxonomicName>
by its smaller dorsal tubercles, absence of large white spots on the dorsum (vs presence), and presence of a brown chevron-shaped marking on the dorsum (vs absence); and from 
<taxonomicName id="4A8C5289070B2BFA15BBEA163FDE6C0B" lsidName="P. karenae" pageId="0" pageNumber="75" rank="species" species="karenae">P. karenae</taxonomicName>
by its brown colouration and dorsal patterning (vs yellow colouration and lack of dorsal patterning), short supratympanic dark brown marking (vs extended along the flank), and less pointed snout. Morphologically and genetically, 
<taxonomicName id="DD0EEB0A6ACE7E091336E432F15112F9" lsidName="P. ando" pageId="0" pageNumber="75" rank="species" species="ando">P. ando</taxonomicName>
sp. nov. most closely resembles 
<taxonomicName id="3FFCCF5B12353F5B4630C5D3338DF1C4" lsidName="P. ravus" pageId="0" pageNumber="75" rank="species" species="ravus">P. ravus</taxonomicName>
. It differs from that species in the lack of yellowish colour on its venter (vs present), yellowish to orangish dorsal finger and toe tip colouration (vs brownish), and by a chevron-shaped brown marking on dorsum (vs W-shaped).
</paragraph>
<paragraph id="18E71269F6A836C5BC736FB09D100CBF" pageId="0" pageNumber="75">
From all members of the externally similar 
<taxonomicName id="8F052E9344745158A628125F666FFB86" class="Amphibia" family="Microhylidae" genus="Cophyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cophyla" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Cophyla</taxonomicName>
, except 
<taxonomicName id="C331B2502B85FF05AB41BA11F0754BE0" lsidName="C. occultans" pageId="0" pageNumber="75" rank="species" species="occultans">C. occultans</taxonomicName>
and 
<taxonomicName id="44AD011322C7B1F619B527C4ED441D95" genus="C." lsidName="C." pageId="0" pageNumber="75" rank="genus">C.</taxonomicName>
sp. 
<normalizedToken id="17B75213AA596E75F80FEE1B371945F4" originalValue="‘fortuna’">'fortuna'</normalizedToken>
(
<bibRefCitation id="43997196B0F7C1C57D8A225B6FC4DC2E" pageId="0" pageNumber="75">Rakotoarison et al. in press</bibRefCitation>
), the new species differs in having a smaller body size (16.9-18.7 mm vs 21.6-33.6 mm). This includes 
<taxonomicName id="DE19E5E0E7D0F09A3F8B7E133761CADB" lsidName="C. maharipeo" pageId="0" pageNumber="75" rank="species" species="maharipeo">C. maharipeo</taxonomicName>
, which is similar in having yellow or orange finger and toe tips, but is larger in size. From 
<taxonomicName id="3A8931CD835632DEFB47DCB9E820C798" lsidName="C. occultans" pageId="0" pageNumber="75" rank="species" species="occultans">C. occultans</taxonomicName>
, 
<taxonomicName id="80A88327FB788A2202C430E59BFE8A3E" lsidName="P. ando" pageId="0" pageNumber="75" rank="species" species="ando">P. ando</taxonomicName>
sp. nov. differs in several call parameters (see below), but is very difficult to distinguish in external morphology, despite clear genetic evidence that these two species are not closely related. From 
<taxonomicName id="C8E97236A2F4418AB1241E361000B5B0" genus="C." lsidName="C." pageId="0" pageNumber="75" rank="genus">C.</taxonomicName>
sp. 
<normalizedToken id="E78022480C88A2D395A3B6D89733E0DA" originalValue="‘fortuna’">'fortuna'</normalizedToken>
, it differs in having the fifth toe distinctly shorter than the third (vs slightly longer than the third), and presence of a brown chevron on the dorsum (vs absence).
</paragraph>
<paragraph id="84DD95B5FBC25EC438C16A41AD9BDE3B" pageId="0" pageNumber="75">
From all members of the genus 
<taxonomicName id="BB930290AA81070E9529B8C2A1B0A067" class="Amphibia" family="Microhylidae" genus="Anodonthyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Anodonthyla" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Anodonthyla</taxonomicName>
, the species can be distinguished by the absence of a distinct finger-like prepollex in males.
</paragraph>
<paragraph id="CF4FFA68AE8FB3D6A78F5876ECA31A66" pageId="0" pageNumber="75">
The new species differs bioacoustically from other 
<taxonomicName id="59B593EC274F885222ECAB1F44C583DF" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
species with known advertisement calls as follows: from 
<taxonomicName id="4591542A0A788AC3950C2753EB7B5C2E" lsidName="P. barbouri" pageId="0" pageNumber="75" rank="species" species="barbouri">P. barbouri</taxonomicName>
, 
<taxonomicName id="7548A99D4561BA5F268CAC1E649CA877" lsidName="P. karenae" pageId="0" pageNumber="75" rank="species" species="karenae">P. karenae</taxonomicName>
, 
<taxonomicName id="BBC70E3E2A544CEA25B031E71268A278" lsidName="P. milloti" pageId="0" pageNumber="75" rank="species" species="milloti">P. milloti</taxonomicName>
, 
<taxonomicName id="5B4683186C83A70AC2E227F37DD0E5CD" lsidName="P. pollicaris" pageId="0" pageNumber="75" rank="species" species="pollicaris">P. pollicaris</taxonomicName>
, 
<taxonomicName id="88FED5041D6D808909397A2EB0A426CC" lsidName="P. tsaratananaensis" pageId="0" pageNumber="75" rank="species" species="tsaratananaensis">P. tsaratananaensis</taxonomicName>
, and 
<taxonomicName id="37297362392959D726BBEEA11E80241B" lsidName="P. tuberifera" pageId="0" pageNumber="75" rank="species" species="tuberifera">P. tuberifera</taxonomicName>
by significantly longer call duration (= note duration; single note calls); and in addition from 
<taxonomicName id="9DE37A627BD85951EC0C22EB5766B48C" lsidName="P. barbouri" pageId="0" pageNumber="75" rank="species" species="barbouri">P. barbouri</taxonomicName>
, 
<taxonomicName id="C5F76BC8F333BB4A267604BE56F17C3F" lsidName="P. milloti" pageId="0" pageNumber="75" rank="species" species="milloti">P. milloti</taxonomicName>
, 
<taxonomicName id="98865138FF463FD9B75EE05E83341405" lsidName="P. pollicaris" pageId="0" pageNumber="75" rank="species" species="pollicaris">P. pollicaris</taxonomicName>
, 
<taxonomicName id="36984306B54641708C1B602BAE3DAE09" lsidName="P. ravus" pageId="0" pageNumber="75" rank="species" species="ravus">P. ravus</taxonomicName>
, 
<taxonomicName id="9B918E85D7935F29D92B1824AD3582BD" lsidName="P. tsaratananaensis" pageId="0" pageNumber="75" rank="species" species="tsaratananaensis">P. tsaratananaensis</taxonomicName>
, and 
<taxonomicName id="CD87F6E6D5DBE5E83BD5787B582C9105" lsidName="P. tuberifera" pageId="0" pageNumber="75" rank="species" species="tuberifera">P. tuberifera</taxonomicName>
by significantly higher dominant frequency (see bioacoustics section below).
</paragraph>
</subSubSection>
<subSubSection id="1E7ECB8D88606337A529422105867CB4" pageId="0" pageNumber="75" type="description">
<paragraph id="09B1D0C95B0E0FBC5066700B2F4363C5" pageId="0" pageNumber="75">Description of the holotype.</paragraph>
<paragraph id="878BF863F4628570931B7737B6D9E45D" pageId="0" pageNumber="75">
Adult male in a good state of preservation, tongue taken as tissue sample. Snout-vent length 18.7 mm; for other measurements, see Table 1. Body long and rather round in preservative (more slender in life; see Fig. 2 
<normalizedToken id="4478CC8FC4943E8C57689D8C86762C6F" originalValue="a–b">a-b</normalizedToken>
); head slightly wider than long (HW/HL 1.04), snout rounded in dorsal and lateral view; nostrils not protuberant, nearer to tip of snout than to eye; canthus rostralis distinct, concave; loreal region slightly concave; tympanum hidden; supratympanic fold indistinct, starting at the posterior corner of the eye and ending anterior to the insertion of the forelimb, dark in colour; tongue attached anteriorly and was posteriorly free; maxillary teeth present; vomerine teeth not visible or palpable (presence of rudimentary vomerine teeth cannot be excluded and would require osteological examination); choanae diminutive, round. Forelimbs slender; subarticular tubercles small, single; outer metacarpal tubercle not visible, inner metacarpal tubercle distinct, forming a large and distinct protuberance at the base of the first finger; hand without webbing; finger discs distinctly broadly rounded, somewhat truncate, with small lateral fringes; relative length of fingers 1&lt;2&lt;4&lt;3; nuptial pads absent. Hindlimbs slender, tibiotarsal articulation reaching tympanum when hindlimb adpressed along body; tibia length 37.9% of SVL; inner metatarsal tubercle small, oblong; outer metatarsal tubercle absent; webbing between third, fourth, and fifth toes rather well developed, webbing formula 1(1) 2i(2) 2e(2) 3i(3) 3e(2) 4i(2.5) 4e(2.5) 5(1); subarticular tubercles on toes indistinct; toes flattened and their discs relatively broad and truncate; relative length of toes 1&lt;2&lt;5&lt;3&lt;4; third toe distinctly shorter than fifth. Dorsal skin smooth, without dorsolateral folds. A very weak mid-dorsal ridge was present in life, but is not evident in preservative. Ventral skin smooth on throat, weakly granular on abdomen and ventral legs.
</paragraph>
<caption id="5C0F01F7DB9E475BADE8B7C8EEBF67F8" pageId="0" pageNumber="75">
<paragraph id="4189CEEF995F0CE73166D85CD40A963B" pageId="0" pageNumber="75">
Table 1. Measurement data on 
<taxonomicName id="F0D2B4E0A6C12C4971C6BDBA523B2D34" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
sp. nov. All measurements in mm. For abbreviations, see the material and methods. HT = holotype, PT = paratype.
</paragraph>
</caption>
<paragraph id="7AB3EE240C194FBFB08D59EB1F14687C" pageId="0" pageNumber="75">
<table id="5165E003AD41DD3FD69D2FF8747BDAF9" pageId="0" pageNumber="75">
<tr id="11A1F28625F95271148381514D7A6544" pageId="0" pageNumber="75">
<th id="DA36931715521C5BFC09B40C91762973" colspan="1" pageId="0" pageNumber="75" rowspan="1">Voucher</th>
<th id="76158AF70617E8DC6C0D48AE83692752" colspan="1" pageId="0" pageNumber="75" rowspan="1">Field no</th>
<th id="D384100B88AA9BEC11284E2FE0DA8809" colspan="1" pageId="0" pageNumber="75" rowspan="1">Sex</th>
<th id="12977108EC0B896232CA686050544C98" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0E51AE" pageId="0" pageNumber="75">SVL</abbrev>
</th>
<th id="140422B07E41B6354BF83D7940AD5D59" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EG2AE" pageId="0" pageNumber="75">HW</abbrev>
</th>
<th id="9B3B956F4333DE3B2FB6874B3BA2DC4D" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EO2AE" pageId="0" pageNumber="75">HL</abbrev>
</th>
<th id="7907C090EA78096E733937ED1C6D5D42" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EW2AE" pageId="0" pageNumber="75">TD</abbrev>
</th>
<th id="CB16706C86DC5E4151F9ADBB5CA594BC" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0E52AE" pageId="0" pageNumber="75">ED</abbrev>
</th>
<th id="F9E8E2A8396BFFFEB7E9D5D64F91E09A" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EG3AE" pageId="0" pageNumber="75">END</abbrev>
</th>
<th id="97C3AB7CCD8AC62C4CA3B7AC6E7BC9B7" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EO3AE" pageId="0" pageNumber="75">NSD</abbrev>
</th>
<th id="A0A463F4C6034CEFFF9DE9733601FE8A" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EW3AE" pageId="0" pageNumber="75">NND</abbrev>
</th>
<th id="9B31A12F5B1BDE68955D6DB2146546CC" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0E53AE" pageId="0" pageNumber="75">FORL</abbrev>
</th>
<th id="1AADB4A3E9A0B51CB5A679BDE8E4F015" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EG4AE" pageId="0" pageNumber="75">HAL</abbrev>
</th>
<th id="21E6B56053011751FAAFA53994033382" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EO4AE" pageId="0" pageNumber="75">HIL</abbrev>
</th>
<th id="E72A5250CE5ACB738151029F18E31456" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0EW4AE" pageId="0" pageNumber="75">FOTL</abbrev>
</th>
<th id="7938ED79AA8B65767A3DA0BFCF95379C" colspan="1" pageId="0" pageNumber="75" rowspan="1">
<abbrev id="ABBRID0E54AE" pageId="0" pageNumber="75">FOL</abbrev>
</th>
<th id="C53580438143ED353C29F53E286734C6" colspan="1" pageId="0" pageNumber="75" rowspan="1">TIBL</th>
</tr>
<tr id="F33EF6AB53F06BB8FAE2B4DAA5AD17A2" pageId="0" pageNumber="75">
<td id="7D96C8B35B1C5DD95706B01C803BDF55" colspan="1" pageId="0" pageNumber="75" rowspan="1">ZSMHT</td>
</tr>
<tr id="29CF4ADA1E45D820F07356AF77E26B3F" pageId="0" pageNumber="75">
<td id="621D3FB344A47E342832783385D70975" colspan="1" pageId="0" pageNumber="75" rowspan="1">ZSMPT</td>
</tr>
<tr id="CDE1F18FFF094D7F871F3DDC94C7EE6F" pageId="0" pageNumber="75">
<td id="68C05853D473922D6FA610BCB0373FA3" colspan="1" pageId="0" pageNumber="75" rowspan="1">ZSMPT</td>
</tr>
</table>
</paragraph>
<paragraph id="C1349C720A706797E372DDA904223201" pageId="0" pageNumber="75">In life, the holotype was olive brown in dorsal colouration with a slightly green-tinged cream saddle marking on its middle, demarcated posteriorly with a dark brown broken border, and anteriorly bordering a dark brown chevron over the suprascapular region that extended to the middle of the eyes, where it stopped abruptly behind an olive-green bar between the eyes (Fig. 2a). The lateral head surface was as the surface of the snout, mottled olive and brown. The same colour was present on the dorsum behind the saddle marking. The dorsal surface of the hindlimbs was a more muted version of this colour, with several dark grey crossbands on each limb segment. The forelimbs were yellowish over the brachium, becoming more orange-tinged distally, with a single, nearly black crossband on the antebrachium and a spot of the same colour on the outer manus. A whitish annulus was present at the base of each terminal phalange. Finger and toe tips were yellowish in colour. Ventrally, it was pale mauve in colour, and the skin was quite transparent, flecked with diminutive cream spots. The digit tips were ventrally also clearly yellow in colouration (Fig. 2b). The iris was gold with black reticulations.</paragraph>
<caption id="47A58807223E6515662E67CF45DEFD4F" pageId="0" pageNumber="75">
<paragraph id="D70BB71623B3582A46429ADD5F47A56B" pageId="0" pageNumber="75">
Figure 2. 
<taxonomicName id="E00B97EFAEE52D17D4960DE3752ADA79" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
specimens in life. (
<normalizedToken id="3668FDF256864AFFA90C60B9B9EAB673" originalValue="a–b">a-b</normalizedToken>
) Holotype (ZSM 293/2010) of 
<taxonomicName id="0A44BF13BD8BC78FABDD9656FDA43259" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
sp. nov. in (a) dorsolateral and (b) ventral view; (
<normalizedToken id="DD48AB74CC9FDB8BC43E35B8744FFC3A" originalValue="c–d">c-d</normalizedToken>
) paratype of 
<taxonomicName id="267095B9771018FF6EC8C467D497F88D" lsidName="P. ando" pageId="0" pageNumber="75" rank="species" species="ando">P. ando</taxonomicName>
sp. nov. (ZSM 292/2010) in (c) lateral and (d) ventral view; (
<normalizedToken id="8BDF673D7A4DC563603B05E434EB98F5" originalValue="e–f">e-f</normalizedToken>
) holotype (ZSM 349/2005) of 
<taxonomicName id="37BE7A75A0EAA7850411D1816AD801DC" lsidName="P. ravus" pageId="0" pageNumber="75" rank="species" species="ravus">P. ravus</taxonomicName>
in (e) dorsolateral and (f) ventral view.
</paragraph>
</caption>
<paragraph id="9B3216BAF888AF9F1EEF907F0ADCB76F" pageId="0" pageNumber="75">After almost nine years in preservative, the specimen has faded considerably, resulting in the loss of distinction in its pattern (Fig. 3). The body and legs are overall beige, with dark oval markings in the suprascapular region, curving over the tympanum, and scattered irregularly over the rest of the dorsum, including some markings in the inguinal region. Faint crossbands on limbs, one of which is distinct on the forearm. A broad, poorly-defined, pale chevron is indistinctly visible on the dorsum.</paragraph>
<caption id="7CF642507861D74EB11B15EC0BF7396F" pageId="0" pageNumber="75">
<paragraph id="DD09D8CF0E36AADEDA54352D69DFC3AC" pageId="0" pageNumber="75">
Figure 3. Preserved holotype (ZSM 293/2010) of 
<taxonomicName id="B9264C81A56F7109728D80E535D0C09F" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
sp. nov. in dorsal and ventral view.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="94BA3E5BF5B11E44813EBB20163A241D" pageId="0" pageNumber="75" type="variation">
<paragraph id="E5A14A105B6DE5387EE8E1DA80F43173" pageId="0" pageNumber="75">Variation.</paragraph>
<paragraph id="537649F4714C6560E65AE845A3090794" pageId="0" pageNumber="75">
For variation in measurements, see Table 1. In general, the paratypes agree well with the holotype, but with the following noteworthy differences: The holotype is the plumpest specimen in the type series, with ZSM 291/2010 and ZSM 292/2010 (Fig. 2) being rather slim. In colouration, the holotype is the lightest specimen of the type series in preservative. Its pattern resembles strongly those of ZSM 291/2010 and 292/2010 (Fig. 2). ZSM 292/2010 in life (Fig. 2 
<normalizedToken id="4BC56B3B986A7A1A01F0872B8012DB52" originalValue="c–d">c-d</normalizedToken>
) apparently possessed a distinctly greenish marking in its inguinal region, of which there are unfortunately no clear photographs. This specimen also had a whiter venter than the holotype.
</paragraph>
</subSubSection>
<subSubSection id="F3218BFBCC8944F19C139DCB4531B657" pageId="0" pageNumber="75" type="bioacoustics">
<paragraph id="811924167B77045709B997BFBF592FC7" pageId="0" pageNumber="75">Bioacoustics.</paragraph>
<paragraph id="48D79CB0410A806545344F498BE95CDD" pageId="0" pageNumber="75">
The advertisement call recorded on the night of the 3rd of April 2010 in Ambodivoangy (estimated air temperature ca. 25 °C) from the holotype, ZSM 293/2010, consists of a single moderately long, high-pitched tonal whistle, repeated at regular intervals (Fig. 4). Numerical call parameters of 14 calls are as follows: call duration (= note duration) 433 
<normalizedToken id="9BE0CBE7C051E89AD31BDD471AF04A0A" originalValue="±">+/-</normalizedToken>
5.8 ms (424-441 ms); inter-call intervals 2655 
<normalizedToken id="CB8B4B7176E1DBCD26E3078C2E73CAEB" originalValue="±">+/-</normalizedToken>
365 ms (2200-3567 ms); call repetition rate within call series approximately 20 calls/minute; dominant frequency 5402 
<normalizedToken id="CA1BE664E831B8149346A49DC8050802" originalValue="±">+/-</normalizedToken>
22 Hz (5380-5432 Hz); prevalent bandwidth 5100-5550 Hz; second frequency band at app. 7800-8200 Hz, and third at 10200-11000 Hz, the latter with the lowest energy of all three recognisable bands. Each note is characterized by a distinct upward modulation of the dominant frequency, starting at around 5250 Hz and increasing up to 5440 Hz before ending with a slight final drop in dominant frequency at around 5300 Hz.
</paragraph>
<caption id="1DD8FE4DF61A99E175D3D6F875492028" pageId="0" pageNumber="75">
<paragraph id="CF1E3F520970237E006958F976AF874F" pageId="0" pageNumber="75">
Figure 4. Audiospectrogram (Hanning window function at 256 FFT width; high-pass filtered at 3750 Hz) and oscillograms of the advertisement call of 
<taxonomicName id="BE346CE7579077324378CEF2A932AC81" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
sp. nov. (holotype, ZSM 293/2010) recorded on 3 April 2010 at Ambodivoangy, northern Madagascar. Lower oscillogram shows a 10 second section of a regular call series.
</paragraph>
</caption>
<paragraph id="ADC699453F6F12AB656F57F2B01370C0" pageId="0" pageNumber="75">
Call comparison: The advertisement call of the sister species 
<taxonomicName id="B4FC6FD510EB80B1EB5A664643692DEB" lsidName="P. ravus" pageId="0" pageNumber="75" rank="species" species="ravus">P. ravus</taxonomicName>
(see 
<bibRefCitation id="921250DB5C9FFDAE8B7FDDAC9300756D" author="Glaw, F" journalOrPublisher="European Journal of Taxonomy" pageId="0" pageNumber="75" pagination="1 - 9" title="A tiny new species of Platypelis from the Marojejy National Park in northeastern Madagascar (Amphibia: Microhylidae)." url="https://doi.org/10.5852/ejt.2012.9" volume="9" year="2012">Glaw et al. 2012</bibRefCitation>
) is rather similar and temporal call parameters overlap with those of 
<taxonomicName id="3988E044BF9F46B251AA1532F39489B1" lsidName="P. ando" pageId="0" pageNumber="75" rank="species" species="ando">P. ando</taxonomicName>
sp. nov.: call duration 384-443 ms (vs. 424-441 ms); inter-call interval 2504-3200 ms (vs. 2200-3567 ms). However, 
<taxonomicName id="AACD61AAFBEF26F2ED0F9D3298B84B54" lsidName="P. ravus" pageId="0" pageNumber="75" rank="species" species="ravus">P. ravus</taxonomicName>
has a distinctly lower dominant frequency, with a mean value of 4010 Hz versus 5402 Hz in the new species. Despite the great similarity in structure of the calls of both species, such differences in dominant frequency are barely explainable with the slight differences in body size of calling males (SVL 19.1 versus 18.7 mm) and thus argue for species-specific differences (see 
<bibRefCitation id="5459E2DCC42ADAE4185ED5362CA0CA93" author="Koehler, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="1 - 124" title="The use of bioacoustics in anuran taxonomy: theory, terminology, methods and recommendations for best practice." url="https://doi.org/10.11646/zootaxa.4251.1.1" volume="4251" year="2017">
<normalizedToken id="290B38D58FDC2AA5506B5639096F7F6F" originalValue="Köhler">Koehler</normalizedToken>
et al. 2017
</bibRefCitation>
).
</paragraph>
<paragraph id="E0D6E41597684454D4F259E44C5FBFEB" pageId="0" pageNumber="75">
Compared to the call of 
<taxonomicName id="C97AD68C80BDF53E2161F29E774BEAB0" lsidName="P. ando" pageId="0" pageNumber="75" rank="species" species="ando">P. ando</taxonomicName>
sp. nov., the advertisement calls of other 
<taxonomicName id="A562258DFA69DB61B498C7BDC5395C64" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
species differ significantly. The call of 
<taxonomicName id="A7F2CE8B5A17E3B31FFE283D08B51B26" lsidName="P. tuberifera" pageId="0" pageNumber="75" rank="species" species="tuberifera">P. tuberifera</taxonomicName>
is shorter (280 ms) and has a lower dominant frequency of 2300-3000 Hz (
<bibRefCitation id="AB115BC1484B337826EBF25A5D44A724" author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlags GbR, Cologne" pageId="0" pageNumber="75" title="A fieldguide to the amphibians and reptiles of Madagascar." year="1994">Glaw and Vences 1994</bibRefCitation>
). The call of 
<taxonomicName id="D78D6EC94A14FE49BBEEAAA43D9BBECB" lsidName="P. barbouri" pageId="0" pageNumber="75" rank="species" species="barbouri">P. barbouri</taxonomicName>
has significantly shorter duration (160 ms) and is repeated at much longer intervals (3200 ms), with a lower dominant frequency of 3850 Hz (
<bibRefCitation id="CD3314F9A1224DDA1719DB08DC471CA4" author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlags GbR, Cologne" pageId="0" pageNumber="75" title="A fieldguide to the amphibians and reptiles of Madagascar." year="1994">Glaw and Vences 1994</bibRefCitation>
). Calls of 
<taxonomicName id="7020247399317FFE7BA5B9D292AF0200" lsidName="P. milloti" pageId="0" pageNumber="75" rank="species" species="milloti">P. milloti</taxonomicName>
are very short (55-65 ms) and exhibit a dominant frequency of approximately 3000 Hz (
<bibRefCitation id="5D9751C7662E82249D5ACDECBD72C759" author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlags GbR, Cologne" pageId="0" pageNumber="75" title="A fieldguide to the amphibians and reptiles of Madagascar." year="1994">Glaw and Vences 1994</bibRefCitation>
). Calls of 
<taxonomicName id="943FB1A38F3A37CBFAECF5C3285DC87B" lsidName="P. pollicaris" pageId="0" pageNumber="75" rank="species" species="pollicaris">P. pollicaris</taxonomicName>
from Andasibe have shorter call duration (160-180 ms) and a dominant frequency of about 3000 Hz (
<bibRefCitation id="6D52B323E6A0914340F0A6C63EEB0237" author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlags GbR, Cologne" pageId="0" pageNumber="75" title="A fieldguide to the amphibians and reptiles of Madagascar." year="1994">Glaw and Vences 1994</bibRefCitation>
). The call of 
<taxonomicName id="26F152A55F763B76184BB82D6952ECCB" lsidName="P. tsaratananaensis" pageId="0" pageNumber="75" rank="species" species="tsaratananaensis">P. tsaratananaensis</taxonomicName>
is very short (79-145 ms duration) at a dominant frequency of 3057-3186 Hz (
<bibRefCitation id="6A9C786514D2BA294E87753EE6CA0C3B" author="Rakotoarison, A" journalOrPublisher="Zootaxa" pageId="0" pageNumber="75" pagination="1 - 25" title="Taxonomy and natural history of arboreal microhylid frogs (Platypelis) from the Tsaratanana Massif in northern Madagascar, with description of a new species." url="https://doi.org/10.11646/zootaxa3563.1.1" volume="3563" year="2012">Rakotoarison et al. 2012</bibRefCitation>
), that of 
<taxonomicName id="F9F03630FAE1A3DCAB4254B8E4BC4499" lsidName="P. karenae" pageId="0" pageNumber="75" rank="species" species="karenae">P. karenae</taxonomicName>
has a duration of 131-145 ms and a dominant frequency 4600-5200 Hz (
<bibRefCitation id="7F11A78088852608BE796D1F8D56BCAE" author="Rosa, GM" journalOrPublisher="Salamandra" pageId="0" pageNumber="75" pagination="201 - 214" title="A new phytotelmic species of Platypelis (Microhylidae: Cophylinae) from the Betampona Reserve, eastern Madagascar." volume="50" year="2014">Rosa et al. 2014</bibRefCitation>
). The call of the morphologically similar 
<taxonomicName id="3BEE35A92272892A7A9299476889E4EA" class="Amphibia" family="Microhylidae" genus="Cophyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cophyla occultans" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="occultans">Cophyla occultans</taxonomicName>
from Nosy Be differs by a slightly longer note duration of 500-550 ms and shorter inter-call intervals (1210-1360 ms) at a dominant frequency of approximately 4000 Hz (
<bibRefCitation id="2DF17548C12494955B0C169EA747775F" author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlags GbR, Cologne" pageId="0" pageNumber="75" title="A fieldguide to the amphibians and reptiles of Madagascar." year="1994">Glaw and Vences 1994</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="26CCCD66B9563F1E72748E2F5BA5A65C" pageId="0" pageNumber="75" type="natural history">
<paragraph id="A133CC97A333CF407388206B657B603B" pageId="0" pageNumber="75">Natural history.</paragraph>
<paragraph id="B709C6CDC8906C915078639C06974F2B" pageId="0" pageNumber="75">
As is typical for 
<taxonomicName id="3263EDADA801484726B2BE93D00560EC" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
species, calling activity was only heard after dusk. ZSM 291/2010 was found calling 1.8 m above the ground. Nothing further is known about the habits of this species, but based on the reproductive ecology of congeners, it is likely to reproduce in phytotelms and have endotrophic nidicolous tadpoles.
</paragraph>
</subSubSection>
<subSubSection id="3E9805C1832345A86AE60C53038BEB38" pageId="0" pageNumber="75" type="available names">
<paragraph id="D05832A406EDC25C3EE61AEDF7B2C70E" pageId="0" pageNumber="75">Available names.</paragraph>
<paragraph id="9DAFDC0052963297024926BE93E8D2FA" pageId="0" pageNumber="75">
Only two available synonyms of any 
<taxonomicName id="EC4177241A6DBF554DC9103506049A9F" class="Amphibia" family="Microhylidae" genus="Cophyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cophyla" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Cophyla</taxonomicName>
or 
<taxonomicName id="2D4B6109678C6FB1BDFDB2598094FEC4" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
refer to small-sized species that could possibly refer to our new species. 
<taxonomicName id="12E5EAB4BA55F9322D5E1491346476F0" class="Amphibia" family="Microhylidae" genus="Cophyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cophyla tuberculata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="tuberculata">Cophyla tuberculata</taxonomicName>
Ahl, 1929 
<normalizedToken id="B0192B16E23A21BF92A0D0A73E8D3B32" originalValue="‘1928’">'1928'</normalizedToken>
is currently a synonym of 
<taxonomicName id="122F3601731DA5846A6068B650AD39E7" lsidName="P. grandis" pageId="0" pageNumber="75" rank="species" species="grandis">P. grandis</taxonomicName>
. The two syntypes are juveniles according to 
<bibRefCitation id="4AB933BFB6935E867A59E0CEA56A672D" author="Blommers-Schloesser, RMA" journalOrPublisher="Faune de Madagascar" pageId="0" pageNumber="75" pagination="1 - 397" title="Amphibiens (premiere partie)." volume="75" year="1991">
<normalizedToken id="BB0A8A5AE6544253CAFDC4416BBF4011" originalValue="Blommers-Schlösser">Blommers-Schloesser</normalizedToken>
and Blanc (1991)
</bibRefCitation>
, but have an SVL of 26 mm, and are therefore larger than the new species. 
<taxonomicName id="7CEB9F65450E8A5F68FAC0C1B3328185" class="Amphibia" family="Microhylidae" genus="Paracophyla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Paracophyla tuberculata" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="tuberculata">Paracophyla tuberculata</taxonomicName>
Millot &amp; 
<normalizedToken id="4EAF0BF6B37503DF7F472946AA20ED97" originalValue="Guibé">Guibe</normalizedToken>
, 1951 is currently considered a synonym of 
<taxonomicName id="26176399599A99CD0FFF28B2934A2DB3" lsidName="P. barbouri" pageId="0" pageNumber="75" rank="species" species="barbouri">P. barbouri</taxonomicName>
. The holotype of that species, MNHN-RA-1957.715, differs from our new species in having a more rugose dorsum, broader finger discs, and a darker venter. Additionally, it is from 
<normalizedToken id="F7F0F4ED262B6ED056997CBC4BB1450C" originalValue="Périnet">Perinet</normalizedToken>
(=Analamazaotra) in the Central East of Madagascar, more than 400 km south of Ambodivoangy. 
<bibRefCitation id="4B2D2AEBD013AF7EA6EBAA13D2C9B3F0" author="Blommers-Schloesser, RMA" journalOrPublisher="Faune de Madagascar" pageId="0" pageNumber="75" pagination="1 - 397" title="Amphibiens (premiere partie)." volume="75" year="1991">
<normalizedToken id="2D96EAA0B904786A5F86124D7A493CA6" originalValue="Blommers-Schlösser">Blommers-Schloesser</normalizedToken>
and Blanc (1991)
</bibRefCitation>
concluded that it is conspecific with 
<taxonomicName id="D75C5D8F2CA098A8538E32EA714EE833" lsidName="P. barbouri" pageId="0" pageNumber="75" rank="species" species="barbouri">P. barbouri</taxonomicName>
, and we agree that it is a member of that species complex, which is in need of revision.
</paragraph>
</subSubSection>
<subSubSection id="0E6B1BB2A3BF008A3DBB72BA0AA85DF0" pageId="0" pageNumber="75" type="etymology">
<paragraph id="7AACE8E84D314A17679EEA3BDA9C1DC6" pageId="0" pageNumber="75">Etymology.</paragraph>
<paragraph id="0637E72385DA3023477D4679C5C65855" pageId="0" pageNumber="75">We dedicate this species to our friend and colleague, Dr. Andolalao Rakotoarison, in recognition of her valuable contributions to the systematics and taxonomy of the Malagasy microhylid fauna. The name is to be treated as an invariable noun in the nominative singular.</paragraph>
</subSubSection>
<subSubSection id="E78F4980286FB884C9E00D1B4B7C2F90" pageId="0" pageNumber="75" type="distribution">
<paragraph id="F73639C71BFA592492D4E225582D9608" pageId="0" pageNumber="75">Distribution.</paragraph>
<paragraph id="987F85FA4E3130CEF9612E029A7C5544" pageId="0" pageNumber="75">
The new species is reliably known only from the type locality Ambodivoangy, but the species is likely to be more widespread in low altitude forest of the adjacent Makira Natural Park. 
<bibRefCitation id="B9E11B0D39993B4D06FF912498A497F8" author="Glaw, F" journalOrPublisher="Vences &amp; Glaw Verlags GbR, Cologne" pageId="0" pageNumber="75" title="A fieldguide to the amphibians and reptiles of Madagascar." year="1992">Glaw and Vences (1992)</bibRefCitation>
found a small 
<taxonomicName id="D5D0DDC4706622028416D65E1F19B926" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="genus">Platypelis</taxonomicName>
species (assigned to and figured as 
<taxonomicName id="33E2C668EA1EFF71D6E6F26F6697F97B" lsidName="P. occultans" pageId="0" pageNumber="75" rank="species" species="occultans">P. occultans</taxonomicName>
) near Voloina (
<geoCoordinate id="B2252B355FC1F9E965518A30B4B6C44E" direction="south" orientation="latitude" precision="5" value="-15.5775">15.5775S</geoCoordinate>
, 
<geoCoordinate id="6AB4957E14396A512B6BF620A13E4C0C" direction="south" orientation="latitude" precision="5" value="-49.6042">49.6042S</geoCoordinate>
; voucher specimens ZFMK 52777-52779), ca. 30 km south of the type locality with similar calls and morphology, which is possibly conspecific with 
<taxonomicName id="18B4E565F54B74D7C559A0CCD9357D76" class="Amphibia" family="Microhylidae" genus="Platypelis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Platypelis ando" order="Anura" pageId="0" pageNumber="75" phylum="Chordata" rank="species" species="ando">Platypelis ando</taxonomicName>
, but further studies are necessary to confirm its identity.
</paragraph>
</subSubSection>
</treatment>
</document>