treatments-xml/data/5D/90/89/5D9089D24E10AB68CFF74DFF54F628D7.xml

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<mods:title id="87F785B41E1A14E5AD2A7259AF0EC0EF">Cyphocoleus Chaudoir (Coleoptera, Carabidae, Odacanthini): descriptive taxonomy, phylogenetic relationships, and the Cenozoic history of New Caledonia</mods:title>
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<mods:namePart id="8285F66D6448117211CABF69C1BB9BAE">Liebherr, James K.</mods:namePart>
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<mods:affiliation id="04C8EE7E9E659932B13DED9C02AF3627">Cornell University Insect Collection, John H. and Anna B. Comstock Hall, Cornell University, Ithaca, NY 14853 - 2601, USA</mods:affiliation>
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<mods:date id="5AE3CABDA738CE7DC14655C5D9F56985">2016</mods:date>
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<treatment id="5D9089D24E10AB68CFF74DFF54F628D7" ID-GBIF-Taxon="182223722" LSID="urn:lsid:zoobank.org:act:8894D2DB-88B0-4205-B69F-9C4A8C34245D" httpUri="http://treatment.plazi.org/id/5D9089D24E10AB68CFF74DFF54F628D7" lastPageId="21" lastPageNumber="232" pageId="19" pageNumber="230">
<subSubSection id="5AE8F7F64109C724167C52E6B92A3A52" pageId="19" pageNumber="230" type="nomenclature">
<paragraph id="972BA39DDCBE2A7094318A90AF7941EF" pageId="19" pageNumber="230">
<pageBreakToken id="E88B11C2211EE3E72179C3E85175C8A4" pageId="19" pageNumber="230" start="start">4</pageBreakToken>
. 
<taxonomicName id="7127B60D869949789014E2E7DB681280" ID-CoL="6BWZ3" LSID="http://zoobank.org/8894D2DB-88B0-4205-B69F-9C4A8C34245D" authority="Liebherr, 2016" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus parovicollis" order="Coleoptera" pageId="19" pageNumber="230" phylum="Arthropoda" rank="species" species="parovicollis" status="sp. n.">Cyphocoleus parovicollis</taxonomicName>
<taxonomicNameLabel id="19AF2E4E1F25FCE5E44F0D7CABE15ED7" pageId="19" pageNumber="230">sp. n.</taxonomicNameLabel>
<figureCitation id="294F6B76CC56E43C1A05BA846401D7D0" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="19" pageNumber="230">Figures 50</figureCitation>
<figureCitation id="95BE5A9EE9BD436D719D710F7F125E2D" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="19" pageNumber="230">, 55-56</figureCitation>
<figureCitation id="BEB3BCC414AC05F436562362544B1585" captionStart="Figures 60–65" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="19" pageNumber="230">, 63</figureCitation>
<figureCitation id="93A96AEC7E56447D073DD693DBE03AEC" captionStart="Figures 66–75" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="19" pageNumber="230">, 69</figureCitation>
<figureCitation id="94012AAC0472D0527A89A6AAD2811286" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="19" pageNumber="230">, 78</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="9E21741F3184C61745E0B9C4CE5903CF" pageId="20" pageNumber="231" type="diagnosis">
<paragraph id="C78B41475D0C67EE92AA70E77FF7CC4E" pageId="20" pageNumber="231">
<pageBreakToken id="041836282A25A85335AC906D83AD2DA8" pageId="20" pageNumber="231" start="start">Diagnosis</pageBreakToken>
.
</paragraph>
<paragraph id="36C3502F314CE0575996AD305A5F6155" pageId="20" pageNumber="231">
This species and its following adelphotaxon are difficult to diagnose practically (Figs 
<figureCitation id="62C3D29EB859250FCC42E6E15AD8E96F" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="20" pageNumber="231">50</figureCitation>
, 
<figureCitation id="17DA551AF9E5914A86BFDB39AF030B4B" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="20" pageNumber="231">82</figureCitation>
), yet are clearly separate species. They can be diagnosed by the more convex eyes in beetles of this species, ocular ratio 1.47-1.55, and presence of four apical abdominal setae in males - two setae each side of apical visible abdominal ventrite - versus two apical abdominal setae - one each side - in males of 
<taxonomicName id="715D062BE17B74C5962E0C5B9BEE5B62" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus ovicollis" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="species" species="ovicollis">
<emphasis id="E113B095A3A5A1F7387FC8E4BAB85DE9" italics="true" pageId="20" pageNumber="231">Cyphocoleus ovicollis</emphasis>
</taxonomicName>
. The pronotum is generally narrower basally among individuals of this species - MPW/BPW = 1.87-2.10 - but the values overlap in the two species, with individuals of 
<taxonomicName id="2188E34147D380C8332BBC4CFA7EC8BB" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus ovicollis" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="species" species="ovicollis">
<emphasis id="50A1A44C634D227EA2A8D099899313E1" italics="true" pageId="20" pageNumber="231">Cyphocoleus ovicollis</emphasis>
</taxonomicName>
ranging in values 2.04-2.29 for the same ratio. If a male is available, the identity based on abdominal setation can be confirmed based on the very different aedeagal internal sacs: that of 
<taxonomicName id="07A5B6527C61041C1514ED62AB7D9FD0" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus parovicollis" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="species" species="parovicollis">
<emphasis id="B129097E43B6531D2EBBBE1DFECAA950" italics="true" pageId="20" pageNumber="231">Cyphocoleus parovicollis</emphasis>
</taxonomicName>
exhibiting an apical 
<normalizedToken id="62E4ABE47E8C8C6B214A35ED07ADF528" originalValue="“cockscomb”">&quot;cockscomb&quot;</normalizedToken>
of large, densely pack macrotrichia (Fig. 
<figureCitation id="42E697F9755FAAC64D7C4E1A5CCB1EBC" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="20" pageNumber="231">56</figureCitation>
). The allopatric geographic distributions can assist in the sorting of these two species, with 
<taxonomicName id="383D1A18610530B646E1327C3FC66DB7" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus parovicollis" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="species" species="parovicollis">
<emphasis id="6C68FF47667479E610F003ADFE9B47F2" italics="true" pageId="20" pageNumber="231">Cyphocoleus parovicollis</emphasis>
</taxonomicName>
distributed to the north of 
<taxonomicName id="D5E2FD8BA380C1161B89021396F414B5" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus ovicollis" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="species" species="ovicollis">
<emphasis id="879194E0CF98BC8CCC188F525EE7A0C8" italics="true" pageId="20" pageNumber="231">Cyphocoleus ovicollis</emphasis>
</taxonomicName>
(Fig. 
<figureCitation id="2798F7751F9DDE92ED06C1F58CF94ECB" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="20" pageNumber="231">78</figureCitation>
). Standardized body length 9.4-12.1 mm.
</paragraph>
</subSubSection>
<subSubSection id="8685820CC5A78E241D2F9B4400238E52" pageId="20" pageNumber="231" type="description">
<paragraph id="C62A2E1311E02D2C239774BBA2F88CA7" pageId="20" pageNumber="231">Description</paragraph>
<paragraph id="5F3BFFD9DA37E0D83C7E76980D79D39F" pageId="20" pageNumber="231">
(n = 5). Head capsule moderately broad, gena elongate anterad constricted neck (Fig. 
<figureCitation id="1A64D0CB419878721D2BE128920ECCBF" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="20" pageNumber="231">50</figureCitation>
); frons broadly convex, but with variably developed chevron-shaped depression medially between eyes; supraorbital setae absent; mandibles elongate, length 2.5 
<normalizedToken id="8BDC7BE1320191406437CD0126E9D2B4" originalValue="×">x</normalizedToken>
distance from antennal articulatory socket to anterolateral margin of labrum; antennae very elongate, scape length 4.33 
<normalizedToken id="93BA344A574A10B82BA52AA69B3D5D51" originalValue="×">x</normalizedToken>
maximal breadth; supraorbital setae absent; gena glabrous, subgenal seta absent. Pronotum narrowly obovoid, MPW/PL = 0.78-0.88; front angles slightly protruded, right to slightly obtuse; lateral marginal bead continuous to obtuse-rounded basal angles and across straight median base; proepisternum bulging outward and so visible in dorsal view for middle half of pronotal length; median longitudinal impression deep, finely incised, terminated posteriorly in variably developed transverse depression; anterior transverse impression very shallow, obliquely paralleling anterior pronotal margin; prosternal process indistinctly depressed medially on ventral surface. Elytra narrowly subellipsoid, MEW/EL = 0.65-0.70; disc broadly and moderately convex; elytral striae deep, impunctate, associated intervals broadly convex; parascutellar seta present, socket not upraised above elytral surface; humeri very narrow, distinctly obtuse-angulate at base of fifth stria; elytral lateral margin convex laterad anterior series of lateral elytral setae; two to three dorsal elytral setae present (setae at anterior and middle positions always present, posterior seta present or absent); lateral elytral setae arranged as 6+ 1(0) + 7; subapical and apical setae present; subapical sinuation broadly and slightly concave, joining lateral margin in a broad curve; elytral apices tightly rounded, slightly separated from suture. Mesepisternum impunctate; metepisternum dorsal length 0.93 
<normalizedToken id="EE5698B4FA277EA3A415AC1BADBC63EF" originalValue="×">x</normalizedToken>
diagonal width. Abdomen of males with distinct, narrow notch medially on apical margin of apical ventrite; females with two setae each side on apical margin of apical ventrite. Legs elongate, gracile, mt1 length/tibial length 0.26; metacoxae bisetose; tarsomeres with short, sparse seta on dorsum; mt4 outer lobe 1.5 
<normalizedToken id="FC147EB96D3D04A087010E8DE4F4C149" originalValue="×">x</normalizedToken>
median length, 4-5 lateroventral setae each side. Microsculpture of frons transversely stretched isodiametric mesh to distinctly transverse mesh on vertex; pronotal disc with dense transverse-line microsculpture loosely organized into a mesh; elytral disc with dense transverse lines resulting in cyanotic to cupreous iridescence; pro-, meso-, and metasternum with sparse pelage of short microsetae, abdominal ventrites and dorsal body surface apparently glabrous. Coloration of head capsule rufopiceous, antennae rufobrunneous with piceous cast on antennomeres 1-3; pronotum piceous, proepipleuron and proepisternum rufopiceous; elytra rufopiceous; elytral epipleuron rufous, metepisternum rufopiceous; femora and tibiae rufobrunneous, tarsi rufoflavous.
</paragraph>
<paragraph id="16901AE4A92503296377A3102095C729" pageId="20" pageNumber="231">
<emphasis id="DA6042C62D994479179CB672459BDF38" italics="true" pageId="20" pageNumber="231">Male genitalia</emphasis>
(n = 4). Male aedeagal median lobe robust, broadly parallel sided to near distal margin of ostial opening, dorsoventral breadth at midlength 0.2 
<normalizedToken id="57F63B2F8F377CCCF2FFCC6DB1F5EBB4" originalValue="×">x</normalizedToken>
distance from tip to base of closed basal bulb (Fig. 
<figureCitation id="3F0817D6375618977F99E6F6725054C2" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="20" pageNumber="231">55</figureCitation>
); lobe shaft brunneous, the internal sac obscured in uneverted position by the moderately melanized lobe wall; lobe apex with broadly rounded tip, tip skewed toward the left side of lobe, length distad ostial opening only slightly more than breadth; internal sac with apical 
<normalizedToken id="92484436B48C53172429F7F41F9FBBAD" originalValue="“cockscomb”">&quot;cockscomb&quot;</normalizedToken>
composed of densely packed, elongate, melanized, spike-like macrotrichia, the sac broadest at midlength (Fig. 
<figureCitation id="D6F920D431E4EB775BBC35D0EC2AC231" captionStart="Figures 51–59" captionStartId="F11" captionText="Figures 51 - 59. Male aedeagal median lobe and associated parameres of Cyphocoleus spp., dextral view: 51, Cyphocoleus lissus, Mt. Humboldt; 52, Cyphocoleus prolixus, Mt. Humboldt; 53, Cyphocoleus heterogenus, Yahoue; 54, Cyphocoleus heterogenus, internal sac everted, La Foa; 55, Cyphocoleus parovicollis, Me Maoya; 56, Cyphocoleus parovicollis, internal sac everted, Me Maoya; 57, Cyphocoleus ovicollis, Ningua Res.; 58 - 59, Cyphocoleus ovicollis, internal sac everted, Mt. Do; 58, laevoventral view; 59, dextrodorsal view. Scale bars, 1.0 mm: horizontal scale for figures with transverse orientation; vertical scale for figures with vertical orientation." figureDoi="10.3897/dez.63.10241.figures51-59" httpUri="https://binary.pensoft.net/fig/111548" pageId="20" pageNumber="231">56</figureCitation>
).
</paragraph>
<paragraph id="8F21D2C995D55E3A4427CC30FD45F3FE" pageId="20" pageNumber="231">
<emphasis id="AD9BFF470E60183F475B4FCE40F4D53A" italics="true" pageId="20" pageNumber="231">Female reproductive tract</emphasis>
(n = 2). Bursa copulatrix vase-shaped, basally stalked, distance from base of gonocoxites to spermathecal basal sclerite 1.8 
<normalizedToken id="2954A1A191A4AABA4375FB079A0BC661" originalValue="×">x</normalizedToken>
maximum breadth (dissection compressed under cover slip) (Fig. 
<figureCitation id="FD0F9E55AA318A031A2A786C321C031E" captionStart="Figures 60–65" captionStartId="F12" captionText="Figures 60 - 65. Female reproductive tract, ventral view, for Cyphocoleus spp.; 60, Cyphocoleus lissus, Mt. Humboldt; 61, Cyphocoleus prolixus, Mt. Humboldt; 62, Cyphocoleus heterogenus, Yahoue; 63, Cyphocoleus parovicollis, Mandjelia; 64, Cyphocoleus monteithi, Mt. Mou; 65, Cyphocoleus miricollis, Foret Nord. For abbreviations see Table 1." figureDoi="10.3897/dez.63.10241.figures60-65" httpUri="https://binary.pensoft.net/fig/111549" pageId="20" pageNumber="231">63</figureCitation>
); bursal walls thick, heavily pigmented by Chlorazol Black stain, with broad band of thick, densely packed spicules lining lumen over broadest part of bursa; basal gonocoxite with apical fringe of six setae, an additional small seventh seta observed in one individual (Fig. 
<figureCitation id="E82D9781B5241995218BF88F733E1EE9" captionStart="Figures 66–75" captionStartId="F13" captionText="Figures 66 - 75. Female right gonocoxae, ventral view, for Cyphocoleus spp. illustrating apical fringe setae of basal gonocoxite 1, and two or three lateral ensiform setae, one dorsal ensiform seta, and two apical nematiform setae on apical gonocoxite 2; 66, Cyphocoleus lissus, Mt. Humboldt; 67, Cyphocoleus prolixus, Mt. Humboldt; 68, Cyphocoleus heterogenus, Yahoue; 69, Cyphocoleus parovicollis, Mandjelia; 70, Cyphocoleus ovicollis, Ningua Res.; 71, Cyphocoleus burwelli, Mt. Humboldt; 72, Cyphocoleus monteithi, Mt. Mou; 73, Cyphocoleus miricollis, Foret Nord; 74, Cyphocoleus mirabilis, Mt. Koghi; 75, Cyphocoleus lescheni, apical gonocoxite damaged, Aoupinie. Scale bar, 0.10 mm." figureDoi="10.3897/dez.63.10241.figures66-75" httpUri="https://binary.pensoft.net/fig/111550" pageId="20" pageNumber="231">69</figureCitation>
); apical gonocoxite acuminate with narrowly rounded apex and three lateral ensiform setae.
</paragraph>
</subSubSection>
<subSubSection id="84FD54E60F02C9595F3D26BFA248E27E" lastPageId="21" lastPageNumber="232" pageId="20" pageNumber="231" type="materials_examined">
<paragraph id="13ECBA3E916FEB4CD231580BECA55175" pageId="20" pageNumber="231">
<typeStatus id="EF442FFE469A49777C91B9E5CF7EDB9A">Type</typeStatus>
.
</paragraph>
<paragraph id="42D291315D93DC0A2C1F87B602161DA8" pageId="20" pageNumber="231">
<materialsCitation id="9976968E390BA49C21B4A063E98B8876" collectingDate="2001-11-24" collectingDateMax="2003-11" collectingDateMin="2001-11-24" collectorName="Pic d'Amoa, N, Pyrethrum, Date, J. K. Liebherr" country="NEW CALEDONIA" elevation="500" location="Holotype" longitude="165.28334" municipality="Sx" specimenCount="1" typeStatus="Holotype">
<typeStatus id="94E517A470E2DA0F132467D8B875D7DC">
<location id="DBC083204272EAB46825451415287AD0" LSID="urn:lsid:plazi:treatment:5D9089D24E10AB68CFF74DFF54F628D7:DBC083204272EAB46825451415287AD0" country="NEW CALEDONIA" longitude="165.28334" municipality="Sx" name="Holotype">Holotype</location>
</typeStatus>
male (QMB deposited in MNHN): 
<collectingCountry id="992DEC9822E0E4D14A230AE02466B5FF" name="New Caledonia">NEW CALEDONIA</collectingCountry>
<emphasis id="98599B86B8441246D5D2BB3477467B6D" bold="true" pageId="20" pageNumber="231">8682</emphasis>
/ 20°58 
<normalizedToken id="8DE0AC84776A7EC603AFBFD1FE2D56A1" originalValue="’Sx165°17’">
' 
<collectingMunicipality id="71879F7872F9B1D8B8FF4D2D24866DC3">Sx</collectingMunicipality>
<geoCoordinate id="15D3B3E3A0658F9F1A0154C2EB9139A9" degrees="165" direction="east" minutes="17" orientation="longitude" precision="925" value="165.28334">165°17'E</geoCoordinate>
</normalizedToken>
. 
<quantity id="93D1D189CEAE60F7BF18972E2DEB8373" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation id="4B9658B52EEDB1B53ED3E52C393F1E79" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500m</elevation>
</quantity>
/ 
<collectorName id="DC6F45926B8852D6DF34F952214E53C1">
Pic 
<normalizedToken id="45552DF7B281B5D20B3B08DF442323FE" originalValue="d’Amoa">d'Amoa</normalizedToken>
, N
</collectorName>
slopes / 
<collectingDate id="54A158FFDA00852B73978745E0B2449E" value="2001-11-24">24Nov2001</collectingDate>
. GBMonteith / 
<collectorName id="B7C80561DA534856B842B1E7AF1C143D">Pyrethrum</collectorName>
, trees &amp; logs // 
<emphasis id="571F4D5EB3A8A83D06AC6B2184CE9187" bold="true" pageId="20" pageNumber="231">QUEENSLAND</emphasis>
/ 
<emphasis id="D9C214A8787EE689CD9764CDC884AAB3" bold="true" pageId="20" pageNumber="231">MUSEUM LOAN</emphasis>
/ 
<collectorName id="F6D7280783F69F3289CE49DFC3DCE153">Date</collectorName>
: 
<collectingDate id="01F4F4007EE252738CE910EFB0152776" value="2003-11">Nov. 2003</collectingDate>
/ no. LEN-1688 (green label) // 
<taxonomicName id="F6885806DBC23CE71708AC717475DF05" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="genus">
<emphasis id="5E097E27D98034641409CF1D976E3E76" italics="true" pageId="20" pageNumber="231">Cyphocoleus</emphasis>
</taxonomicName>
/ n. sp. / 
<taxonomicName id="89B25C039A5E44FE42B728A5D2310768" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="ovicollis" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="species" species="ovicollis">
<emphasis id="5CB36CE959E86FE925741B16929C3528" italics="true" pageId="20" pageNumber="231">ovicollis</emphasis>
</taxonomicName>
male 9 / det. J.K. 
<bibRefCitation id="58353CDA218BD07759BF73EDB0486EBD" author="Liebherr, JK" journalOrPublisher="ZooKeys" pageId="49" pageNumber="260" pagination="1 - 407" publicationUrl="10.3897/zookeys.544.6074" refId="B33" refString="Liebherr, JK, 2015. The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation. ZooKeys 544: 1 - 407, DOI: 10.3897/zookeys.544.6074" title="The Mecyclothorax beetles (Coleoptera, Carabidae, Moriomorphini) of Haleakala, Maui: Keystone of a hyperdiverse Hawaiian radiation." url="10.3897/zookeys.544.6074" volume="544" year="2015">Liebherr 2015</bibRefCitation>
// genitalia vial // 
<typeStatus id="31D07992E99F4DAC4FE44923405BF93D">HOLOTYPE</typeStatus>
/ 
<taxonomicName id="8A4C849CDD7FBFB730EC968B2367AEC0" authorityName="Chaudoir" authorityYear="1877" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus" order="Diptera" pageId="20" pageNumber="231" phylum="Arthropoda" rank="genus">Cyphocoleus</taxonomicName>
/ parovipennis / 
<collectorName id="C23D5040AD5D7397935E038FEA1836F3">J.K. Liebherr</collectorName>
2016 (black-bordered red label)
</materialsCitation>
.
</paragraph>
<paragraph id="1124D1546D6000E4709976CBC578F2B6" lastPageId="21" lastPageNumber="232" pageId="20" pageNumber="231">
<materialsCitation id="2AA3FB269B2F6C4666B952103295C734" collectingDate="1984-05-11" collectingDateMax="2004-10-02" collectingDateMin="1984-05-11" collectorName="Monteith, Cook, Burwell, Pic d'Amoa, N, Wright" country="NEW CALEDONIA" county="Mt. Panie" elevation="675" latitude="-21.366667" location="Mandjelia" longLatPrecision="1264" longitude="165.33333" municipality="Pouebo" specimenCount="30" typeStatus="Paratypes">
<typeStatus id="9B2C1B5A1E896823F25922D12F2BF02C">Paratypes</typeStatus>
(
<specimenCount id="F23F6C36CAF9C610F6C230FA03BC6E84" type="generic">30 specimens</specimenCount>
). 
<collectingCountry id="BF7975CBBC39BE9A4A76D3E0D9DACE13" name="New Caledonia">NEW CALEDONIA</collectingCountry>
: 
<normalizedToken id="C7D02451E3AA4A2839EF2A8040BB6512" originalValue="Mandjélia">
<location id="A8E534B18091266B1801DB4013D5696E" LSID="urn:lsid:plazi:treatment:5D9089D24E10AB68CFF74DFF54F628D7:A8E534B18091266B1801DB4013D5696E" country="NEW CALEDONIA" county="Mt. Panie" latitude="-21.366667" longLatPrecision="1264" longitude="165.33333" municipality="Pouebo" name="Mandjelia">Mandjelia</location>
</normalizedToken>
, above 
<normalizedToken id="8C845567B60FB90EB293EAB22D29BF1E" originalValue="Pouébo">
<collectingMunicipality id="E5075D0932C13948A5499F9BD9A57240">Pouebo</collectingMunicipality>
</normalizedToken>
<quantity id="6E53831A7373410AC11A88EF846B4F17" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" unit="m" value="675.0" valueMax="750.0" valueMin="600.0">
<elevation id="03C42D20D8C1EC8BE99A7E4A8E088F7A" metricMagnitude="2" metricUnit="m" metricValue="6.75" metricValueMax="7.5" metricValueMin="6.0" unit="m" value="675.0" valueMax="750.0" valueMin="600.0">600-750 m</elevation>
</quantity>
el, 
<geoCoordinate id="AA6F0F9441596E5AC378145C01D94D88" degrees="20" direction="south" minutes="24" orientation="latitude" precision="925" value="-20.4">20°24'S</geoCoordinate>
, 
<geoCoordinate id="04F91EF41A8BEDFD5C34FB6C052AA112" degrees="164" direction="east" minutes="32" orientation="longitude" precision="925" value="164.53334">164°32'E</geoCoordinate>
, 
<collectingDate id="E4EEDF75AA37A1706BC9F4E0537F7333" value="1984-05-11" valueMax="1984-05-13" valueMin="1984-05-11">11-13-v-1984</collectingDate>
, 
<collectorName id="13A31B007DA5BEFBA581250BBFBD5A01">Monteith</collectorName>
&amp; 
<collectorName id="7B58F82A0B5C4AF8C77D92851B780D3E">Cook</collectorName>
(QMB,1); 
<collectingCounty id="B45A2916CD91487D9F6D092A2953F1AF">
Mt. 
<normalizedToken id="F769E14BDF639C1FD94F1082A2162674" originalValue="Panié">Panie</normalizedToken>
</collectingCounty>
refuge, 
<quantity id="3D9E3470E6B8EEC54368B93FCA53C177" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation id="12608528831A4CCFA7542A8F79384401" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
el., 
<geoCoordinate id="A5995208E45DFD82149CA5B490A91651" degrees="20" direction="south" minutes="34" orientation="latitude" precision="925" value="-20.566668">20°34'S</geoCoordinate>
, 
<geoCoordinate id="77154C9BA12C8C98717297CDD25FA244" degrees="164" direction="east" minutes="46" orientation="longitude" precision="925" value="164.76666">164°46'E</geoCoordinate>
, 
<collectingDate id="CE2DFAFF95E0FB1489DA4BC25E5908C5" value="2000-11-16" valueMax="2000-11-18" valueMin="2000-11-16">16-18-xi-2000</collectingDate>
, Bouchard, 
<collectorName id="0365A4E089B46E9F483FDAB8A6065E85">Burwell</collectorName>
&amp; 
<collectorName id="4C02EE9CEF1CB6C7499B43F1369CBDFB">Monteith</collectorName>
(QMB, 1; lot no. 9938); 
<collectorName id="8ED7C312E93CC58FBA3AC80890DC2C6E">
Pic 
<normalizedToken id="39D6201DD2458E8239F8ABA1B6C4A081" originalValue="d’Amoa">d'Amoa</normalizedToken>
, N
</collectorName>
slopes, 
<quantity id="49A0DA5B840517F62816B5534A5D46F7" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">
<elevation id="4527626D74425016E36B8673E8A379A4" metricMagnitude="2" metricUnit="m" metricValue="5.0" unit="m" value="500.0">500 m</elevation>
</quantity>
el., hand collecting, 
<geoCoordinate id="9D98AB71E080199C0FEDB866DD3B570F" degrees="20" direction="south" minutes="58" orientation="latitude" precision="925" value="-20.966667">20°58'S</geoCoordinate>
, 
<geoCoordinate id="78D95168FEEBD194B570A457CDDD9EC6" degrees="165" direction="east" minutes="17" orientation="longitude" precision="925" value="165.28334">165°17'E</geoCoordinate>
, 
<collectingDate id="D8FE3A8027E5C3F8AB43708DE2317070" value="2001-11-10" valueMax="2001-11-11" valueMin="2001-11-10">10-11-xi- 2001</collectingDate>
, 
<collectorName id="92205582E441AB6785E06103A72C6790">Burwell</collectorName>
&amp; 
<collectorName id="5529D97C34A431F189F119DDE6584871">Monteith</collectorName>
(QMB, 1; lot no. 8687); 
<normalizedToken id="8B14961002C99F8857AFA62213FED71F" originalValue="Aoupinié">Aoupinie</normalizedToken>
, 
<quantity id="6BDE5972F35D13C987D7938A4941D80B" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">
<elevation id="1D459EFB95E9EB664BCD93B4AF8DAD34" metricMagnitude="2" metricUnit="m" metricValue="8.5" unit="m" value="850.0">850 m</elevation>
</quantity>
el., 
<geoCoordinate id="BC221329B754A4C5026552B76104591D" degrees="21" direction="south" minutes="11" orientation="latitude" precision="925" value="-21.183332">21°11'S</geoCoordinate>
, 
<geoCoordinate id="9AD41F2170B4ABB3C00D3960D4BEDB6C" degrees="165" direction="east" minutes="19" orientation="longitude" precision="925" value="165.31667">165°19'E</geoCoordinate>
, 
<collectingDate id="1B7537E7A3FE92621D10B539CF6FA996" value="2000-11-20" valueMax="2000-11-21" valueMin="2000-11-20">20-21-xi-2000</collectingDate>
, Bouchard, 
<collectorName id="AA085D2925A41ADC8F53C404EEE71DEB">Burwell</collectorName>
&amp; 
<collectorName id="BCCCC45DF4FBBE0CDAE7DD7E8AC78A0F">Monteith</collectorName>
(QMB, 2; lot no. 9930), summit, 
<quantity id="5319DAC4198CBDADD9B1FBEB1EB35FAF" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">
<elevation id="4C4A749A0B114DCF7A792499551D8234" metricMagnitude="3" metricUnit="m" metricValue="1.0" unit="m" value="1000.0">1000 m</elevation>
</quantity>
el., pyrethrum trees &amp; logs, 
<geoCoordinate id="6C6DAED4517819BF296405862FCD87CB" degrees="21" direction="south" minutes="11" orientation="latitude" precision="925" value="-21.183332">21°11'S</geoCoordinate>
, 
<geoCoordinate id="D73565D74823C39ABF57074F54D0CD37" degrees="165" direction="east" minutes="16" orientation="longitude" precision="925" value="165.26666">165°16'E</geoCoordinate>
, 
<collectingDate id="CF9270591B4E7050A9F10D1063680496" value="2004-10-02">02-x-2004</collectingDate>
, Monteith (QMB, 1; lot no. 11665), 
<geoCoordinate id="850C41EFE6E0F14530F5A2939917E7F8" degrees="21" direction="south" minutes="11" orientation="latitude" precision="925" value="-21.183332">21°11'S</geoCoordinate>
, 
<geoCoordinate id="A20927312605CC6BA0567236F151A6D4" degrees="165" direction="east" minutes="19" orientation="longitude" precision="925" value="165.31667">165°19'E</geoCoordinate>
, 
<collectingDate id="61886991CAAC135ABB1A01F305041377" value="1993-12-12">
<pageBreakToken id="B5253DEA91E2699693EE4314377C59EA" pageId="21" pageNumber="232" start="start">12</pageBreakToken>
-xii-1993
</collectingDate>
, Raven (QMB, 1); Me Maoya, near summit, 
<quantity id="D9ABF56A1C51C45DF1804CADF8A1D4B7" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">
<elevation id="30E0F3259DF7C77C46817517AE44B67A" metricMagnitude="3" metricUnit="m" metricValue="1.4" unit="m" value="1400.0">1400 m</elevation>
</quantity>
el., pyrethrum trees &amp; logs, 
<geoCoordinate id="E8D4E217DFD63D2A155D4EB8285FCEAD" degrees="21" direction="south" minutes="22" orientation="latitude" precision="925" value="-21.366667">21°22'S</geoCoordinate>
, 
<geoCoordinate id="CD02F0EB1DFE70B63F9C13EEFF0673A1" degrees="165" direction="east" minutes="20" orientation="longitude" precision="925" value="165.33333">165°20'E</geoCoordinate>
, 
<collectingDate id="A114D0BF02412B81AEB3A3CCCA256B00" value="2002-11-12">12-xi-2002</collectingDate>
, 
<collectorName id="9C9404FC9297C3AFFE4506E820258EB3">Burwell</collectorName>
&amp; 
<collectorName id="373D8C825145F00E727A63A45CA902B3">Monteith</collectorName>
(QMB, 1; lot no. 11163); Me Maoya camp, 
<quantity id="D9F55C1F59F3BB105C53C36D06BC5608" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">
<elevation id="354D553E7D5ADE67D82B33E99F1D3998" metricMagnitude="3" metricUnit="m" metricValue="1.15" unit="m" value="1150.0">1150 m</elevation>
</quantity>
el., night collecting, 
<geoCoordinate id="67C584FF1F8A9657C0ECD515854DC3D1" degrees="21" direction="south" minutes="22" orientation="latitude" precision="925" value="-21.366667">21°22'S</geoCoordinate>
, 
<geoCoordinate id="27C3F388545A98BA1A95299578B94D52" degrees="165" direction="east" minutes="20" orientation="longitude" precision="925" value="165.33333">165°20'E</geoCoordinate>
, 
<collectingDate id="0F152922B5DC2064D19003414BBDE20F" value="2002-11-11" valueMax="2002-11-12" valueMin="2002-11-11">11-12-xi- 2002</collectingDate>
, Burwell, 
<collectorName id="DC9525AF93A618442EE261F556619E8E">Monteith</collectorName>
&amp; 
<collectorName id="4A54CCABE6E66022DE76B3EFEA102C64">Wright</collectorName>
(QMB, 15; lot no. 11159), pyrethrum trees &amp; logs, 
<collectingDate id="DEACBBBE45D74BA47BB0574064A12528" value="2002-11-11" valueMax="2002-11-13" valueMin="2002-11-11">11-13-xi-2002</collectingDate>
, 
<collectorName id="763B9F21EAF2D2F28E0CB553B0B22C84">Monteith</collectorName>
&amp; 
<collectorName id="C8167D25DE3B41D9AE7415DD80686D10">Burwell</collectorName>
(QMB, 5; lot no. 11158); Me Maoya, near summit, 
<quantity id="AB034916906AA683767157C666F89B9E" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">
<elevation id="4061C1E8CA26BE332C7E42F6D2848067" metricMagnitude="3" metricUnit="m" metricValue="1.3" unit="m" value="1300.0">1300 m</elevation>
</quantity>
el., pyrethrum trees &amp; logs, 
<geoCoordinate id="E8F03D1E695ABA93FF0D6034F4BFBDFC" degrees="21" direction="south" minutes="22" orientation="latitude" precision="925" value="-21.366667">21°22'S</geoCoordinate>
, 
<geoCoordinate id="05A76A36E6F906FE3AA2B1914D157920" degrees="165" direction="east" minutes="20" orientation="longitude" precision="925" value="165.33333">165°20'E</geoCoordinate>
, 
<collectingDate id="D26C63DB7004EFAC69575AABFCB6CC65" value="2002-11-12">12-xi-2002</collectingDate>
, 
<collectorName id="398FFEEBA13D14010D7462970F1EA86A">Monteith</collectorName>
&amp; 
<collectorName id="DE071A978C515DEAF8F6F427F79B3B64">Burwell</collectorName>
(QMB, 2; lot no. 11169)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9A949D7371069B07B9A169EEEC7888C4" pageId="21" pageNumber="232" type="etymology">
<paragraph id="E49D0920ED6EDEB7836B5BC828DFBB81" pageId="21" pageNumber="232">Etymology.</paragraph>
<paragraph id="2F4064C4E589D3E261C6D5926DDF8749" pageId="21" pageNumber="232">
The adelphotaxon status and great similarity of this species and 
<taxonomicName id="C21B7B25D554AF687B066E8014DC9964" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus ovicollis" order="Diptera" pageId="21" pageNumber="232" phylum="Arthropoda" rank="species" species="ovicollis">
<emphasis id="935327A376D7F3D0349100F2B7717E6A" italics="true" pageId="21" pageNumber="232">Cyphocoleus ovicollis</emphasis>
</taxonomicName>
(Figs 
<figureCitation id="F13CAEA14CB9E90895B5CD94D2C7EE1F" captionStart="Figure 46" captionStartId="F9" captionText="Figure 46. Parsimony cladogram of subtribe Homethina with characters indicated; character numbers above cladogram stems, state numbers below stems [cladogram divided into two parts joined at &quot; A &quot;]; geographic abbreviations as in Figs 44 - 45." figureDoi="10.3897/dez.63.10241.figure46" httpUri="https://binary.pensoft.net/fig/111546" pageId="21" pageNumber="232">46</figureCitation>
, 
<figureCitation id="0F2F004BA65548FF183716C4B5D05346" captionStart="Figures 47–50" captionStartId="F10" captionText="Figures 47 - 50. Dorsal habitus photographs of Cyphocoleus spp.: 47, Cyphocoleus lissus female, Mt. Humboldt; 48, Cyphocoleus prolixus male, Mt. Humboldt; 49, Cyphocoleus heterogenus male, Yahoue; 50, Cyphocoleus parovicollis male, Pic d'Amoa." figureDoi="10.3897/dez.63.10241.figures47-50" httpUri="https://binary.pensoft.net/fig/111547" pageId="21" pageNumber="232">50</figureCitation>
, 
<figureCitation id="D407BB8A845EA2126C04A484AE74A2EF" captionStart="Figures 82–85" captionStartId="F15" captionText="Figures 82 - 85. Dorsal habitus photographs of Cyphocoleus spp.: 82, Cyphocoleus ovicollis female, Ningua Res.; 83, Cyphocoleus burwelli female, Mt. Canala; 84, Cyphocoleus angustatus male, Mt. Panie; 85, Cyphocoleus monteithi male, Dzumac Rd. jct." figureDoi="10.3897/dez.63.10241.figures82-85" httpUri="https://binary.pensoft.net/fig/111552" pageId="21" pageNumber="232">82</figureCitation>
) is signified through use of 
<taxonomicName id="CB2104B64B1D44735BA7A36339015161" authorityName="Liebherr" authorityYear="2016" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="parovicollis" order="Diptera" pageId="21" pageNumber="232" phylum="Arthropoda" rank="species" species="parovicollis">parovicollis</taxonomicName>
as this 
<normalizedToken id="D394E33EBBDE1EF3101CD89B758A358A" originalValue="species’">species'</normalizedToken>
epithet: i.e. equal to 
<taxonomicName id="1B0C330C88EFDC42981CAF96E0BD2BE7" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="ovicollis" order="Diptera" pageId="21" pageNumber="232" phylum="Arthropoda" rank="species" species="ovicollis">ovicollis</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="58EE58A619A23C43BB34798B763049DE" pageId="21" pageNumber="232" type="distribution">
<paragraph id="4CCC621A29C769F212B6EC8211B2A371" pageId="21" pageNumber="232">Distribution and habitat.</paragraph>
<paragraph id="EA653F3959636EAEE73ED8218B6DD944" pageId="21" pageNumber="232">
This species is distributed in the northern half of Grande Terre, allopatric with its adelphotaxon, 
<taxonomicName id="F6EA879758BF6E8067A4631BA64B3868" authorityName="Fauvel" authorityYear="1882" class="Insecta" family="Carabidae" genus="Cyphocoleus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cyphocoleus ovicollis" order="Diptera" pageId="21" pageNumber="232" phylum="Arthropoda" rank="species" species="ovicollis">
<emphasis id="F71C6752BC394152DE978EA12B4C100A" italics="true" pageId="21" pageNumber="232">Cyphocoleus ovicollis</emphasis>
</taxonomicName>
, which is distributed to the south (Fig. 
<figureCitation id="AF64B57DF9B5D01265A07DF00B11E40F" captionStart="Figures 76–81" captionStartId="F14" captionText="Figures 76 - 81. Distribution maps of Cyphocoleus spp.: 76, Cyphocoleus lissus; 77, Cyphocoleus heterogenus, Cyphocoleus prolixus; 78, Cyphocoleus parovicollis, Cyphocoleus ovicollis; 79, Cyphocoleus burwelli, Cyphocoleus angustatus, Cyphocoleus monteithi; 80, Cyphocoleus miricollis, Cyphocoleus fasciatus; 81, Cyphocoleus mirabilis, Cyphocoleus moorei, Cyphocoleus lescheni." figureDoi="10.3897/dez.63.10241.figures76-81" httpUri="https://binary.pensoft.net/fig/111551" pageId="21" pageNumber="232">78</figureCitation>
). Beetles have been collected from tree trunks and downed logs via pyrethrin fog, and also by hand collecting at night.
</paragraph>
</subSubSection>
</treatment>
</document>