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<document id="BA395C42B1613E0453411089CA4EB171" ID-CLB-Dataset="22527" ID-DOI="10.11646/zootaxa.4679.2.7" ID-GBIF-Dataset="128dfed7-a482-46b0-b976-61f73748bc7b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3772595" ID-ZooBank="44D197DB-E48A-4FCC-9B58-0B397D03880D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588057932805" checkinUser="plazi" docAuthor="Tiunova, Tatiana M. &amp; Semenchenko, Alexander A." docDate="2019" docId="5D69DC6F4F576C7549E64AD4FABEF87B" docLanguage="en" docName="zootaxa.4679.2.7.pdf" docOrigin="Zootaxa 4679 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Baetis pentaphyllus Tiunova &amp; Semenchenko 2019, sp. nov." docType="treatment" docUuid="2B453886-A76D-44D0-9FE4-ABBDA273527A" docUuidSource="ZooBank" docVersion="8" lastPageNumber="348" masterDocId="A150A4174F566C7249714F7DFFF0FF9A" masterDocTitle="Baetis pentaphyllus sp. nov., a new species of mayfly (Ephemeroptera: Baetidae) from the Russian Far East" masterLastPageNumber="352" masterPageNumber="341" pageNumber="342" updateTime="1698837707890" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="5A23A54F229020AB6A62CB79CE7F53EB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FA2C34A2F6A97A5C855691A1048029DC">
<mods:title id="18C32927B72736477116843CE4EDA369">Baetis pentaphyllus sp. nov., a new species of mayfly (Ephemeroptera: Baetidae) from the Russian Far East</mods:title>
</mods:titleInfo>
<mods:name id="4BED9B143551924A9FF9FD2DF95DFB1A" type="personal">
<mods:role id="DEB19C49A77A9B76AFCC66838C83F301">
<mods:roleTerm id="5BEA793D5868D0C146CCF07918E39068">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="EA9FB58F7B8A1A391317004A495E488A">Tiunova, Tatiana M.</mods:namePart>
<mods:affiliation id="0943CF90A3DFED51B5B18632BE2A1F19">Federal Scientific Center of the East Asia Terrestrial Biodiversity, Far Eastern Branch, Russian Academy of Sciences, Vladivostok, 690022, Russia.</mods:affiliation>
<mods:nameIdentifier id="C6612E4A6EC9293CC73E055BCFEECD92" type="email">tiunova@biosoil.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="0EA5BDEF2F22F070F1FAFCA13A2428A2" type="personal">
<mods:role id="094DA6A3C84D86AD519C8B68ECF19F1D">
<mods:roleTerm id="C5673A1074240532E72EA1EA4610DC4B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AB6AAC44E1F38903F405C8B697974D89">Semenchenko, Alexander A.</mods:namePart>
<mods:affiliation id="400E6CACF7E059587C5D8F5232968987">Far Eastern Federal University, Vladivostok, 690950, Russia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="C677FC49837B66A4074EFE3622E75D98">text</mods:typeOfResource>
<mods:relatedItem id="3914EE67514A6CCCEF11FC8468AA93ED" type="host">
<mods:titleInfo id="E1C1596A7692F57239DE763BC34C5B9B">
<mods:title id="93115D8713B5717D00C933DB110E99D6">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="25D38F7578D6A336DF60C09DDD5F51C0">
<mods:date id="672E056B61B29CA2C40F70B8F79ABEA1">2019</mods:date>
<mods:detail id="14F27D03F0CAA7F8BE759571D2E0B289" type="pubDate">
<mods:number id="DEF090C42D5C8A5B162330CFFF2C4996">2019-10-02</mods:number>
</mods:detail>
<mods:detail id="A5783C780ABFB3320990073E42E10864" type="volume">
<mods:number id="5659FF926B1E14C0122EC181BB530A71">4679</mods:number>
</mods:detail>
<mods:detail id="DAF8C53F22B294ED6FD8EDB3F22EFB65" type="issue">
<mods:number id="835A59EBBBE93F490B1D123904A93493">2</mods:number>
</mods:detail>
<mods:extent id="3B3D9E54A01FA3C07320B1E9D0E17E27" unit="page">
<mods:start id="9F98D87C53DF179ED8FC14C89CE4A1CF">341</mods:start>
<mods:end id="BC1A7B07A8C3E0B46FF8E520D22A32D1">352</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="D1897BD92272ACD486F234CE379BADE4">journal article</mods:classification>
<mods:identifier id="0710DB0C4F561A0604CD4B0B8B52593B" type="CLB-Dataset">22527</mods:identifier>
<mods:identifier id="51E24B8780F97DFEAECBAF1A7CC01B95" type="DOI">10.11646/zootaxa.4679.2.7</mods:identifier>
<mods:identifier id="2FC306CC233B8DDAC51F4091CEE8D740" type="GBIF-Dataset">128dfed7-a482-46b0-b976-61f73748bc7b</mods:identifier>
<mods:identifier id="0A58B5D2ECB7012D0F1AD21FD9C7B46A" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="3A4E00F8461ECDC565CBA821B92B2C28" type="Zenodo-Dep">3772595</mods:identifier>
<mods:identifier id="AABDC963915323D028EA7059368B940C" type="ZooBank">44D197DB-E48A-4FCC-9B58-0B397D03880D</mods:identifier>
</mods:mods>
<treatment id="5D69DC6F4F576C7549E64AD4FABEF87B" ID-DOI="http://doi.org/10.5281/zenodo.3797701" ID-GBIF-Taxon="163829883" ID-Zenodo-Dep="3797701" LSID="urn:lsid:zoobank.org:act:2B453886-A76D-44D0-9FE4-ABBDA273527A" httpUri="http://treatment.plazi.org/id/5D69DC6F4F576C7549E64AD4FABEF87B" lastPageId="7" lastPageNumber="348" pageId="1" pageNumber="342">
<subSubSection id="9DDA3EF24F576C7349E64AD4FC4AFA7D" pageId="1" pageNumber="342" type="nomenclature">
<paragraph id="D57F6D794F576C7349E64AD4FE19FA59" blockId="1.[151,954,1449,1548]" box="[151,489,1449,1476]" pageId="1" pageNumber="342">
<heading id="8E37DA154F576C7349E64AD4FE19FA59" bold="true" box="[151,489,1449,1476]" fontSize="11" level="1" pageId="1" pageNumber="342" reason="1">
<emphasis id="E7B4B16B4F576C7349E64AD4FE19FA59" bold="true" box="[151,489,1449,1476]" pageId="1" pageNumber="342">
<taxonomicName id="12C016FA4F576C7349E64AD4FE74FA5E" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[151,388,1449,1476]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="342" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F576C7349E64AD4FE74FA5E" bold="true" box="[151,388,1449,1476]" italics="true" pageId="1" pageNumber="342">Baetis pentaphyllus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FC870C104F576C7348FD4AD4FE19FA59" box="[396,489,1449,1475]" pageId="1" pageNumber="342" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D57F6D794F576C7349E64AB1FC4AFA7D" blockId="1.[151,954,1449,1548]" box="[151,954,1484,1511]" pageId="1" pageNumber="342">
<uri id="A151617B4F576C7349E64AB1FC4AFA7D" box="[151,954,1484,1511]" pageId="1" pageNumber="342">
urn:lsid:zoobank.org:pub:
<uuid id="A16657AC4F576C7348C14AB1FC4AFA7D" box="[432,954,1484,1511]" pageId="1" pageNumber="342">32A67029-582F-40C3-9B40-66FEA0881566</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="9DDA3EF24F576C7349E64A8CFEE9F996" box="[151,281,1521,1548]" pageId="1" pageNumber="342" type="description">
<paragraph id="D57F6D794F576C7349E64A8CFEE9F996" blockId="1.[151,954,1449,1548]" box="[151,281,1521,1548]" pageId="1" pageNumber="342">
(
<figureCitation id="4DFB71FC4F576C7349EE4A8CFEE1F996" box="[159,273,1521,1548]" captionStart-0="FIGURES 1–4" captionStart-1="FIGURES 5–10" captionStart-2="FIGURES 11–14" captionStart-3="FIGURES 15" captionStart-4="FIGURES 19–23" captionStart-5="FIGURES 24–26" captionStartId-0="2.[151,264,1968,1993]" captionStartId-1="3.[151,264,1928,1953]" captionStartId-2="4.[151,264,1702,1727]" captionStartId-3="5.[151,264,1448,1473]" captionStartId-4="6.[151,264,1514,1539]" captionStartId-5="7.[151,262,1812,1837]" captionTargetBox-0="[151,1436,194,1868]" captionTargetBox-1="[169,1433,189,1844]" captionTargetBox-2="[151,1428,179,1655]" captionTargetBox-3="[161,1435,194,1398]" captionTargetBox-4="[159,1419,593,1490]" captionTargetBox-5="[248,1339,237,1787]" captionTargetId-0="figure@2.[151,1436,194,1868]" captionTargetId-1="figure@3.[151,1436,181,1844]" captionTargetId-2="figure@4.[151,1433,179,1655]" captionTargetId-3="figure@5.[151,1436,179,1421]" captionTargetId-4="figure@6.[151,1436,589,1495]" captionTargetId-5="figure@7.[248,1339,237,1787]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionTargetPageId-4="6" captionTargetPageId-5="7" captionText-0="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." captionText-1="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." captionText-2="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." captionText-3="FIGURES 15˗18. Baetis pentaphyllus sp. nov., larvae, dorsal view: 15, foreleg; 16, tarsal claw; 17, posterior margin of abdominal tergum VI; 18, paraproct." captionText-4="FIGURES 19–23. Baetis pentaphyllus sp. nov., larvae, tergalii shape, dorsal view, roman numbers belong to the respective tergalius pairs." captionText-5="FIGURES 24–26. Baetis pentaphyllus sp. nov., larvae: 24, posterior margin of abdominal tergum VI; 25, tergalii I; 26, tergalii II." figureDoi-0="http://doi.org/10.5281/zenodo.3772597" figureDoi-1="http://doi.org/10.5281/zenodo.3772599" figureDoi-2="http://doi.org/10.5281/zenodo.3772601" figureDoi-3="http://doi.org/10.5281/zenodo.3772603" figureDoi-4="http://doi.org/10.5281/zenodo.3772605" figureDoi-5="http://doi.org/10.5281/zenodo.3772607" httpUri-0="https://zenodo.org/record/3772597/files/figure.png" httpUri-1="https://zenodo.org/record/3772599/files/figure.png" httpUri-2="https://zenodo.org/record/3772601/files/figure.png" httpUri-3="https://zenodo.org/record/3772603/files/figure.png" httpUri-4="https://zenodo.org/record/3772605/files/figure.png" httpUri-5="https://zenodo.org/record/3772607/files/figure.png" pageId="1" pageNumber="342">Figs 1–26</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9DDA3EF24F576C7349E64945FB60F8B1" pageId="1" pageNumber="342" type="materials_examined">
<paragraph id="D57F6D794F576C7349E64945FB60F8B1" blockId="1.[151,1437,1592,2015]" pageId="1" pageNumber="342">
<emphasis id="E7B4B16B4F576C7349E64945FEF6F9C9" bold="true" box="[151,262,1592,1619]" pageId="1" pageNumber="342">Material.</emphasis>
<materialsCitation id="65A867244F576C7348614944FF07F901" ID-GBIF-Occurrence="2609011863" collectingDate="2015-07-02" collectionCode="T" country="Russia" county="Bolshoi Garmakan River" elevation="300" latitude="53.8858" location="Zeyskii Reserve" longLatPrecision="1" longitude="127.19376" municipality="Zeya Reservoir Basin" pageId="1" pageNumber="342" specimenCount="1" typeStatus="holotype">
<emphasis id="E7B4B16B4F576C7348614944FE8DF9C9" bold="true" box="[272,381,1593,1619]" pageId="1" pageNumber="342">
<typeStatus id="0A7BD3DB4F576C7348614944FE8DF9C9" box="[272,381,1593,1619]" pageId="1" pageNumber="342" type="holotype">Holotype</typeStatus>
</emphasis>
larva.
<emphasis id="E7B4B16B4F576C7348BE4945FCE6F9C9" bold="true" box="[463,790,1592,1619]" pageId="1" pageNumber="342">
<collectingCountry id="ADD72DE94F576C7348BE4945FCE1F9C9" box="[463,785,1592,1619]" name="Russia" pageId="1" pageNumber="342">RUSSIAN FEDERATION</collectingCountry>
:
</emphasis>
AMURSKAYA OBLAST’:
<location id="D01F3BA24F576C734D2E4944FAE9F9C9" LSID="urn:lsid:plazi:treatment:5D69DC6F4F576C7549E64AD4FABEF87B:D01F3BA24F576C734D2E4944FAE9F9C9" box="[1119,1305,1593,1619]" country="Russia" county="Bolshoi Garmakan River" latitude="53.8858" longLatPrecision="1" longitude="127.19376" municipality="Zeya Reservoir Basin" name="Zeyskii Reserve" pageId="1" pageNumber="342">Zeyskii Reserve</location>
,
<collectingMunicipality id="351BF7034F576C734C564944FED7F9ED" pageId="1" pageNumber="342">Zeya Reservoir Basin</collectingMunicipality>
,
<collectingCounty id="3C1E15F54F576C7348444920FDA4F9ED" box="[309,596,1629,1655]" pageId="1" pageNumber="342">Bolshoi Garmakan River</collectingCounty>
, about
<quantity id="1238C09C4F576C734BDA4920FD04F9ED" box="[683,756,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="342" unit="m" value="300.0">
<elevation id="5EED8A4A4F576C734BDA4920FD04F9ED" box="[683,756,1629,1655]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="342" unit="m" value="300.0">300 m</elevation>
</quantity>
above mouth,
<geoCoordinate id="B0F40BBE4F576C734AD54920FBC5F9ED" box="[932,1077,1628,1655]" degrees="53" direction="north" minutes="53.148" orientation="latitude" pageId="1" pageNumber="342" precision="1" value="53.8858">N 53°53.148</geoCoordinate>
<geoCoordinate id="B0F40BBE4F576C734D314920FB2EF9ED" box="[1088,1246,1629,1655]" degrees="127" direction="east" minutes="11.626" orientation="longitude" pageId="1" pageNumber="342" precision="1" value="127.19376">E 127°11.626</geoCoordinate>
,
<date id="A17E4BB94F576C734D9C4921FA88F9ED" box="[1261,1400,1628,1655]" pageId="1" pageNumber="342" value="2015-07-02">
<collectingDate id="B13AB2514F576C734D9C4921FA88F9ED" box="[1261,1400,1628,1655]" pageId="1" pageNumber="342" value="2015-07-02">02.VII.2015</collectingDate>
</date>
,
<collectionCode id="B3D1F5BC4F576C734CF74920FA68F9ED" box="[1414,1432,1629,1655]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="342">T</collectionCode>
. Tiunova
</materialsCitation>
.
<materialsCitation id="65A867244F576C73487249FCFB7CF8B1" ID-GBIF-Occurrence="2609011864" collectingDate="2014-07-07" collectingDateMax="2016-08-03" collectingDateMin="2014-07-07" collectionCode="T" collectorName="Tiunova. &amp; Kolyma River Basin &amp; A. Semenchenko" country="Russia" county="Tiunova" elevation="500" latitude="68.38585" location="Malyi Garmakan River" longLatPrecision="1" longitude="127.17432" municipality="Zeya Reservoir Basin" pageId="1" pageNumber="342" specimenCount="18" specimenCount-larva="18" stateProvince="Chukotka" typeStatus="paratype">
<emphasis id="E7B4B16B4F576C73487249FCFE8BF901" bold="true" box="[259,379,1665,1691]" pageId="1" pageNumber="342">
<typeStatus id="0A7BD3DB4F576C73487249FCFE8BF901" box="[259,379,1665,1691]" pageId="1" pageNumber="342" type="paratype">Paratypes</typeStatus>
</emphasis>
: collected the same data and place as
<typeStatus id="0A7BD3DB4F576C734A4249FCFC69F901" box="[819,921,1665,1691]" pageId="1" pageNumber="342" type="holotype">holotype</typeStatus>
:
<specimenCount id="C3C6A6F04F576C734AD449FDFBF3F901" box="[933,1027,1664,1691]" pageId="1" pageNumber="342" type="larva">5 larvae</specimenCount>
,
<specimenCount id="C3C6A6F04F576C734D6149FCFBAFF901" box="[1040,1119,1665,1691]" pageId="1" pageNumber="342" type="larva">1 larva</specimenCount>
(
<emphasis id="E7B4B16B4F576C734D0049FDFAB9F901" bold="true" box="[1137,1353,1664,1691]" pageId="1" pageNumber="342">BAE-BGAR1_110</emphasis>
); same locality,
<date id="A17E4BB94F576C73498A49D9FE77F925" box="[251,391,1700,1727]" pageId="1" pageNumber="342" value="2015-07-08">
<collectingDate id="B13AB2514F576C73498A49D9FE77F925" box="[251,391,1700,1727]" pageId="1" pageNumber="342" value="2015-07-08">08.VII.2015</collectingDate>
</date>
,
<specimenCount id="C3C6A6F04F576C7348E749D8FE05F925" box="[406,501,1701,1727]" pageId="1" pageNumber="342" type="larva">4 larvae</specimenCount>
,
<collectionCode id="B3D1F5BC4F576C734B7549D8FDE6F925" box="[516,534,1701,1727]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="342">T</collectionCode>
.
<collectingCounty id="3C1E15F54F576C734B5549D8FD74F925" box="[548,644,1701,1727]" pageId="1" pageNumber="342">Tiunova</collectingCounty>
;
<collectingMunicipality id="351BF7034F576C734BE249D8FC61F925" box="[659,913,1701,1727]" pageId="1" pageNumber="342">Zeya Reservoir Basin</collectingMunicipality>
:
<location id="D01F3BA24F576C734AD149D8FB5CF925" LSID="urn:lsid:plazi:treatment:5D69DC6F4F576C7549E64AD4FABEF87B:D01F3BA24F576C734AD149D8FB5CF925" box="[928,1196,1701,1727]" county="Tiunova" latitude="68.38585" longLatPrecision="1" longitude="127.17432" municipality="Zeya Reservoir Basin" name="Malyi Garmakan River" pageId="1" pageNumber="342" stateProvince="Chukotka">Malyi Garmakan River</location>
, about
<quantity id="1238C09C4F576C734C7549D9FABFF925" box="[1284,1359,1700,1727]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="342" unit="m" value="500.0">
<elevation id="5EED8A4A4F576C734C7549D9FABFF925" box="[1284,1359,1700,1727]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="342" unit="m" value="500.0">500 m</elevation>
</quantity>
above mouth,
<geoCoordinate id="B0F40BBE4F576C73499D49B4FE89F979" box="[236,377,1736,1763]" degrees="53" direction="north" minutes="52.295" orientation="latitude" pageId="1" pageNumber="342" precision="1" value="53.871582">N 53°52.295</geoCoordinate>
<geoCoordinate id="B0F40BBE4F576C7348F149B4FDECF979" box="[384,540,1736,1763]" degrees="127" direction="east" minutes="10.459" orientation="longitude" pageId="1" pageNumber="342" precision="1" value="127.17432">E 127°10.459</geoCoordinate>
,
<date id="A17E4BB94F576C734B5449B5FD40F979" box="[549,688,1736,1763]" pageId="1" pageNumber="342" value="2015-07-06">
<collectingDate id="B13AB2514F576C734B5449B5FD40F979" box="[549,688,1736,1763]" pageId="1" pageNumber="342" value="2015-07-06">06.VII.2015</collectingDate>
</date>
,
<specimenCount id="C3C6A6F04F576C734BCA49B4FCE6F979" box="[699,790,1737,1763]" pageId="1" pageNumber="342" type="larva">3 larvae</specimenCount>
,
<collectionCode id="B3D1F5BC4F576C734A5149B4FCC2F979" box="[800,818,1737,1763]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="342">T</collectionCode>
. Tiunova;
<location id="D01F3BA24F576C734AD449B4FB9CF979" LSID="urn:lsid:plazi:treatment:5D69DC6F4F576C7549E64AD4FABEF87B:D01F3BA24F576C734AD449B4FB9CF979" box="[933,1132,1737,1763]" county="Tiunova" latitude="68.38585" longLatPrecision="1" longitude="127.17432" municipality="Zeya Reservoir Basin" name="Shirokovka River" pageId="1" pageNumber="342" stateProvince="Chukotka">Shirokovka River</location>
, about
<quantity id="1238C09C4F576C734DCB49B4FB0FF979" box="[1210,1279,1737,1763]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="342" unit="m" value="700.0">
<elevation id="5EED8A4A4F576C734DCB49B4FB0FF979" box="[1210,1279,1737,1763]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="342" unit="m" value="700.0">700 m</elevation>
</quantity>
above mouth,
<date id="A17E4BB94F576C7349E64990FED2F89D" box="[151,290,1773,1799]" pageId="1" pageNumber="342" value="2014-07-07">
<collectingDate id="B13AB2514F576C7349E64990FED2F89D" box="[151,290,1773,1799]" pageId="1" pageNumber="342" value="2014-07-07">07.VII.2014</collectingDate>
</date>
,
<specimenCount id="C3C6A6F04F576C73485F4991FE79F89D" box="[302,393,1772,1799]" pageId="1" pageNumber="342" type="larva">5 larvae</specimenCount>
,
<collectionCode id="B3D1F5BC4F576C7348E54990FE56F89D" box="[404,422,1773,1799]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="342">T</collectionCode>
.
<collectorName id="783508AF4F576C7348C14990FDE4F89D" box="[432,532,1773,1799]" pageId="1" pageNumber="342">Tiunova.</collectorName>
<collectingRegion id="1704A39B4F576C734B6B4990FD46F89D" box="[538,694,1773,1799]" country="Russia" name="Chukotka" pageId="1" pageNumber="342">CHUKOTKA</collectingRegion>
AUTONOMOUS DISTRICT,
<collectorName id="783508AF4F576C734D634990FB0BF89D" box="[1042,1275,1773,1799]" pageId="1" pageNumber="342">
<location id="D01F3BA24F576C734D634990FB41F89D" LSID="urn:lsid:plazi:treatment:5D69DC6F4F576C7549E64AD4FABEF87B:D01F3BA24F576C734D634990FB41F89D" box="[1042,1201,1773,1799]" county="Tiunova" latitude="68.38585" longLatPrecision="1" longitude="127.17432" municipality="Zeya Reservoir Basin" name="Kolyma River" pageId="1" pageNumber="342" stateProvince="Chukotka">
Kolyma
<location id="D01F3BA24F576C734D024990FB41F89D" LSID="urn:lsid:plazi:treatment:5D69DC6F4F576C7549E64AD4FABEF87B:D01F3BA24F576C734D024990FB41F89D" box="[1139,1201,1773,1799]" county="Tiunova" latitude="68.38585" longLatPrecision="1" longitude="127.17432" municipality="Zeya Reservoir Basin" name="River Basin" pageId="1" pageNumber="342" stateProvince="Chukotka">River</location>
</location>
Basin
</collectorName>
,
<location id="D01F3BA24F576C734C744990FEECF8B1" LSID="urn:lsid:plazi:treatment:5D69DC6F4F576C7549E64AD4FABEF87B:D01F3BA24F576C734C744990FEECF8B1" county="Tiunova" latitude="68.38585" longLatPrecision="1" longitude="127.17432" municipality="Zeya Reservoir Basin" name="Malyi Keperweim River" pageId="1" pageNumber="342" stateProvince="Chukotka">Malyi Keperweim River</location>
, 2 mature larvae,
<geoCoordinate id="B0F40BBE4F576C734895486CFD86F8B1" box="[484,630,1808,1835]" degrees="68" direction="north" minutes="23.151" orientation="latitude" pageId="1" pageNumber="342" precision="1" value="68.38585">N 68°23.151</geoCoordinate>
; E 166°80.348,
<date id="A17E4BB94F576C734A5B486CFC4FF8B1" box="[810,959,1809,1835]" pageId="1" pageNumber="342" value="2016-08-03">
<collectingDate id="B13AB2514F576C734A5B486CFC4FF8B1" box="[810,959,1809,1835]" pageId="1" pageNumber="342" value="2016-08-03">03.VIII.2016</collectingDate>
</date>
,
<collectorName id="783508AF4F576C734AB8486CFB7CF8B1" box="[969,1164,1809,1835]" pageId="1" pageNumber="342">A. Semenchenko</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9DDA3EF24F576C7449B64849FDCAFD81" lastPageId="6" lastPageNumber="347" pageId="1" pageNumber="342" type="description">
<paragraph id="D57F6D794F576C7449B64849FDCAFD81" blockId="1.[151,1437,1592,2015]" lastBlockId="6.[151,1437,153,539]" lastPageId="6" lastPageNumber="347" pageId="1" pageNumber="342">
<emphasis id="E7B4B16B4F576C7349B64849FE40F8D5" bold="true" box="[199,432,1844,1871]" pageId="1" pageNumber="342">
Description.
<emphasis id="E7B4B16B4F576C73482E4848FE40F8D5" bold="true" box="[351,432,1845,1871]" italics="true" pageId="1" pageNumber="342">Larvae</emphasis>
</emphasis>
(in alcohol). Length (mm): body 4.6–5.7; cerci 2.5–3.1. General body colour brown (
<figureCitation id="4DFB71FC4F576C734C1B4848FF32F8E9" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="1" pageNumber="342">Figs 1–4</figureCitation>
).
<emphasis id="E7B4B16B4F576C7349AA4824FED4F8E9" bold="true" box="[219,292,1881,1907]" pageId="1" pageNumber="342">Head:</emphasis>
brown. Antennae brownish, slightly longer than ½ of body length. Scape and pedicel with hair-like setae only. Labrum distinctly wider than long (the width/length ratio 1.69–1.70); dorsal surface with 1 + 6–7 long submarginal setae, arranged in uneven row and row long pointed setae laterally on both margins (
<figureCitation id="4DFB71FC4F576C734DBC48DCFAE0F821" box="[1229,1296,1952,1979]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="1" pageNumber="342">Fig. 5</figureCitation>
). Canines of left mandible with eight teeth divided into two groups (
<figureCitation id="4DFB71FC4F576C734A7048B8FCBAF845" box="[769,842,1989,2015]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="1" pageNumber="342">Figs 6</figureCitation>
,
<figureCitation id="4DFB71FC4F576C734A2448B8FC84F845" box="[853,884,1989,2015]" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="1" pageNumber="342">12</figureCitation>
). Incisor (outer group) with four teeth; outermost tooth broadened and rounded apically; kinetodontium (inner group) with four small teeth. Both groups separated by a shallow incision. Left prostheca toothbrush-like (
<figureCitation id="4DFB71FC4F526C764B9C4853FCC4F8D2" box="[749,820,1838,1864]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="4" pageNumber="345">Figs 6</figureCitation>
,
<figureCitation id="4DFB71FC4F526C764A314852FCAEF8D3" box="[832,862,1839,1865]" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="4" pageNumber="345">12</figureCitation>
). The right mandible canines are also separated into two groups of teeth (
<figureCitation id="4DFB71FC4F526C764806482FFE4DF8F7" box="[375,445,1874,1901]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="4" pageNumber="345">Figs 7</figureCitation>
,
<figureCitation id="4DFB71FC4F526C7648B9482EFE15F8F7" box="[456,485,1875,1901]" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="4" pageNumber="345">11</figureCitation>
). Incisor with three teeth, first tooth widened and rounded apically; kinetodontium with four teeth, from which second largest. Between the kinetodontium and incisor there is a groove. Right prostheca with four non-pointed teeth (
<figureCitation id="4DFB71FC4F526C7648E648E7FE2CF82F" box="[407,476,1946,1973]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="4" pageNumber="345">Figs 7</figureCitation>
,
<figureCitation id="4DFB71FC4F526C76489648E6FDF4F82F" box="[487,516,1947,1973]" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="4" pageNumber="345">11</figureCitation>
). Maxillary palp two-segmented, individual segments approximately equal in length; surface of both segments covered with hair-like seta, which occur more densely on distal part of second segment (
<figureCitation id="4DFB71FC4F526C764C0148C3FF45F867" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="4" pageNumber="345">Fig. 13</figureCitation>
). Second segment of labial palpus with rounded apicomedial projection, its width 1.1 times wider than the base of third segment; third segment almost symmetrical rounded, relatively wide (
<figureCitation id="4DFB71FC4F536C774AA8494AFBEDF9C8" box="[985,1053,1591,1618]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="5" pageNumber="346">Fig. 9</figureCitation>
); dorsal surface with a row of 4–5 long pointed setae on second segment and long pointed stout setae in apical margin of third segment; ventral surface of third segment covered numerous stout setae accompanied by hair-like seta (
<figureCitation id="4DFB71FC4F536C774AA34902FBE4F900" box="[978,1044,1663,1690]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="5" pageNumber="346">Fig. 9</figureCitation>
). Labium with paraglossae concave in middle, nearly two times wider than glossae; with 7–8 pairs of long stout bristles form three irregular rows on tip and long stout setae near apex; 5–6 long bristles located along outer margin and two stout setae near apex of inner margin (
<figureCitation id="4DFB71FC4F536C7749864996FEBFF89C" box="[247,335,1771,1798]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="5" pageNumber="346">Figs 10</figureCitation>
,
<figureCitation id="4DFB71FC4F536C77482C4991FE8CF89C" box="[349,380,1772,1798]" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="5" pageNumber="346">14</figureCitation>
). Glossae relatively slender, with row 6–7 of long setae on inner margin and 5–6 bristles on outer margin and by one long stout seta located near apex (
<figureCitation id="4DFB71FC4F536C774A5B4872FC70F8B0" box="[810,896,1807,1834]" captionStart="FIGURES 5–10" captionStartId="3.[151,264,1928,1953]" captionTargetBox="[169,1433,189,1844]" captionTargetId="figure@3.[151,1436,181,1844]" captionTargetPageId="3" captionText="FIGURES 5–10. Baetis pentaphyllus sp. nov., larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="5" pageNumber="346">Figs 10</figureCitation>
,
<figureCitation id="4DFB71FC4F536C774AFD486DFC5BF8B0" box="[908,939,1808,1834]" captionStart="FIGURES 11–14" captionStartId="4.[151,264,1702,1727]" captionTargetBox="[151,1428,179,1655]" captionTargetId="figure@4.[151,1433,179,1655]" captionTargetPageId="4" captionText="FIGURES 11–14. Baetis pentaphyllus sp. nov., larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view." figureDoi="http://doi.org/10.5281/zenodo.3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="5" pageNumber="346">14</figureCitation>
).
<emphasis id="E7B4B16B4F536C774AB1486DFBE9F8B0" bold="true" box="[960,1049,1808,1834]" pageId="5" pageNumber="346">Thorax</emphasis>
: brown with whitish diffuse spots. Anterior margins of pro- and mesonotum darker (
<figureCitation id="4DFB71FC4F536C774BB9484EFCDEF8D4" box="[712,814,1843,1870]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="5" pageNumber="346">Figs 1–2</figureCitation>
). Legs whitish, joints of leg segments brown (
<figureCitation id="4DFB71FC4F536C774C4A484EFA7FF8D4" box="[1339,1423,1843,1870]" captionStart="FIGURES 15" captionStartId="5.[151,264,1448,1473]" captionTargetBox="[161,1435,194,1398]" captionTargetId="figure@5.[151,1436,179,1421]" captionTargetPageId="5" captionText="FIGURES 15˗18. Baetis pentaphyllus sp. nov., larvae, dorsal view: 15, foreleg; 16, tarsal claw; 17, posterior margin of abdominal tergum VI; 18, paraproct." figureDoi="http://doi.org/10.5281/zenodo.3772603" httpUri="https://zenodo.org/record/3772603/files/figure.png" pageId="5" pageNumber="346">Fig. 15</figureCitation>
). Femora pale, at edges darker; outer margin with row of long and pointed bristles and short setae located irregular; inner margin with villopore and small pointed setae located irregularly (
<figureCitation id="4DFB71FC4F536C774AB64806FBEBF80C" box="[967,1051,1915,1942]" captionStart="FIGURES 15" captionStartId="5.[151,264,1448,1473]" captionTargetBox="[161,1435,194,1398]" captionTargetId="figure@5.[151,1436,179,1421]" captionTargetPageId="5" captionText="FIGURES 15˗18. Baetis pentaphyllus sp. nov., larvae, dorsal view: 15, foreleg; 16, tarsal claw; 17, posterior margin of abdominal tergum VI; 18, paraproct." figureDoi="http://doi.org/10.5281/zenodo.3772603" httpUri="https://zenodo.org/record/3772603/files/figure.png" pageId="5" pageNumber="346">Fig. 15</figureCitation>
). Tibia and tarsus brown in apical part. Stout setae arranged in regular row along inner margin of tibia and tarsi and setae of different sizes located along outer margin. Claws brown, with row of 10–11 teeth and a pair of subapical setae (Fig. 16).
<emphasis id="E7B4B16B4F536C774D9048BEFAA6F844" bold="true" box="[1249,1366,1987,2014]" pageId="5" pageNumber="346">Abdomen</emphasis>
: terga I–VIII brown, lateral area lighter; a pair of dark brown spots located near anterior margin; lateral and posterior margins dark brown (
<figureCitation id="4DFB71FC4F506C7448F64FC0FE38FF4D" box="[391,456,189,215]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Fig. 1</figureCitation>
); terga I–VI with median light longitudinal stripe; terga IX–X brown, lighter than other terga; posterior margin of terga with broad, rounded apically teeth (Figs 17, 24). Sterna generally light brown with brown diffused spots near anterior margins (
<figureCitation id="4DFB71FC4F506C744BE44E78FD2CFE85" box="[661,732,261,287]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Fig. 3</figureCitation>
.) Paraproct serrated on posterior part of inner margin (10–13 large pointed teeth of different size) and outer part of posterior part (approximately 19–22 teeth diminishing proximally) (Fig. 18). Five pairs of elongated, oval-shaped tergalii located on III–VII tergites (
<figureCitation id="4DFB71FC4F506C744D094E30FB4CFEFD" box="[1144,1212,333,359]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Fig. 4</figureCitation>
), approximately 2.5 times longer than broad, rounded (
<figureCitation id="4DFB71FC4F506C744B6D4E0CFD55FE11" box="[540,677,369,395]" captionStart="FIGURES 19–23" captionStartId="6.[151,264,1514,1539]" captionTargetBox="[159,1419,593,1490]" captionTargetId="figure@6.[151,1436,589,1495]" captionTargetPageId="6" captionText="FIGURES 19–23. Baetis pentaphyllus sp. nov., larvae, tergalii shape, dorsal view, roman numbers belong to the respective tergalius pairs." figureDoi="http://doi.org/10.5281/zenodo.3772605" httpUri="https://zenodo.org/record/3772605/files/figure.png" pageId="6" pageNumber="347">Figs. 19–23</figureCitation>
), and 1.8–2.0 times longer than corresponding segment (
<figureCitation id="4DFB71FC4F506C744C544E0CFA9BFE11" box="[1317,1387,369,395]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Fig. 4</figureCitation>
); all tergalii white, tracheation poorly visible; margins hairy; hairs inserted in small teeth bases. Tergalii I and III almost equal length (
<figureCitation id="4DFB71FC4F506C74485F4EC4FE8CFE49" box="[302,380,441,467]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Figs. 4</figureCitation>
,
<figureCitation id="4DFB71FC4F506C7448F64EC4FE3EFE49" box="[391,462,441,467]" captionStart="FIGURES 19–23" captionStartId="6.[151,264,1514,1539]" captionTargetBox="[159,1419,593,1490]" captionTargetId="figure@6.[151,1436,589,1495]" captionTargetPageId="6" captionText="FIGURES 19–23. Baetis pentaphyllus sp. nov., larvae, tergalii shape, dorsal view, roman numbers belong to the respective tergalius pairs." figureDoi="http://doi.org/10.5281/zenodo.3772605" httpUri="https://zenodo.org/record/3772605/files/figure.png" pageId="6" pageNumber="347">19, 20</figureCitation>
), tergalii IV slightly shorter than III (
<figureCitation id="4DFB71FC4F506C744A174EC4FC44FE49" box="[870,948,441,467]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Figs. 4</figureCitation>
,
<figureCitation id="4DFB71FC4F506C744ACE4EC4FBF8FE49" box="[959,1032,441,467]" captionStart="FIGURES 19–23" captionStartId="6.[151,264,1514,1539]" captionTargetBox="[159,1419,593,1490]" captionTargetId="figure@6.[151,1436,589,1495]" captionTargetPageId="6" captionText="FIGURES 19–23. Baetis pentaphyllus sp. nov., larvae, tergalii shape, dorsal view, roman numbers belong to the respective tergalius pairs." figureDoi="http://doi.org/10.5281/zenodo.3772605" httpUri="https://zenodo.org/record/3772605/files/figure.png" pageId="6" pageNumber="347">22, 23</figureCitation>
). Tergalius II is the largest, 1.2 times longer than the first and 1.6 times longer than V (
<figureCitation id="4DFB71FC4F506C744BBB4EA0FCEAFE6D" box="[714,794,477,503]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Figs. 4</figureCitation>
,
<figureCitation id="4DFB71FC4F506C744A564EA0FCB7FE62" box="[807,839,477,504]" captionStart="FIGURES 24–26" captionStartId="7.[151,262,1812,1837]" captionTargetBox="[248,1339,237,1787]" captionTargetId="figure@7.[248,1339,237,1787]" captionTargetPageId="7" captionText="FIGURES 24–26. Baetis pentaphyllus sp. nov., larvae: 24, posterior margin of abdominal tergum VI; 25, tergalii I; 26, tergalii II." figureDoi="http://doi.org/10.5281/zenodo.3772607" httpUri="https://zenodo.org/record/3772607/files/figure.png" pageId="6" pageNumber="347">26</figureCitation>
). Caudal filaments brownish (
<figureCitation id="4DFB71FC4F506C744DED4EA0FAF9FE6D" box="[1180,1289,477,503]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Figs. 1, 3</figureCitation>
). Cerci three times longer than paracercus (
<figureCitation id="4DFB71FC4F506C7448994D7CFDDDFD81" box="[488,557,513,539]" captionStart="FIGURES 1–4" captionStartId="2.[151,264,1968,1993]" captionTargetBox="[151,1436,194,1868]" captionTargetId="figure@2.[151,1436,194,1868]" captionTargetPageId="2" captionText="FIGURES 1–4. Baetis pentaphyllus sp. nov., larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III˗VII, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="6" pageNumber="347">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="81BF3DF14F546C7049E648CDFBC9F877" ID-DOI="http://doi.org/10.5281/zenodo.3772597" ID-Zenodo-Dep="3772597" httpUri="https://zenodo.org/record/3772597/files/figure.png" pageId="2" pageNumber="343" startId="2.[151,264,1968,1993]" targetBox="[151,1436,194,1868]" targetPageId="2">
<paragraph id="D57F6D794F546C7049E648CDFBC9F877" blockId="2.[151,1436,1968,2029]" pageId="2" pageNumber="343">
<emphasis id="E7B4B16B4F546C7049E648CDFECBF853" bold="true" box="[151,315,1968,1993]" pageId="2" pageNumber="343">FIGURES 1–4.</emphasis>
<taxonomicName id="12C016FA4F546C70483348CCFDF9F852" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[322,521,1968,1992]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F546C70483348CCFDF9F852" box="[322,521,1968,1992]" italics="true" pageId="2" pageNumber="343">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F546C704B6148CCFD98F853" bold="true" box="[528,616,1969,1993]" pageId="2" pageNumber="343">
<taxonomicNameLabel id="FC870C104F546C704B6148CCFD94F853" box="[528,612,1969,1993]" pageId="2" pageNumber="343" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
larvae, dorsal view: 1, (material from type locality); 2, larva from Malyi Keperweim River (Chukotka); 3, ventral view; 4, location of tergalii on terga III
<emphasis id="E7B4B16B4F546C704AF748A8FC7EF877" bold="true" box="[902,910,2005,2029]" pageId="2" pageNumber="343">˗</emphasis>
VII, dorsal view.
</paragraph>
</caption>
<caption id="81BF3DF14F556C7149E648F5FCB4F873" ID-DOI="http://doi.org/10.5281/zenodo.3772599" ID-Zenodo-Dep="3772599" httpUri="https://zenodo.org/record/3772599/files/figure.png" pageId="3" pageNumber="344" startId="3.[151,264,1928,1953]" targetBox="[169,1433,189,1844]" targetPageId="3">
<paragraph id="D57F6D794F556C7149E648F5FCB4F873" blockId="3.[151,1436,1928,2025]" pageId="3" pageNumber="344">
<emphasis id="E7B4B16B4F556C7149E648F5FEB6F83B" bold="true" box="[151,326,1928,1953]" pageId="3" pageNumber="344">FIGURES 5–10.</emphasis>
<taxonomicName id="12C016FA4F556C71483D48F4FDE2F83B" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[332,530,1929,1953]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F556C71483D48F4FDE2F83B" box="[332,530,1929,1953]" italics="true" pageId="3" pageNumber="344">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F556C714B6948F4FD9EF83B" bold="true" box="[536,622,1929,1953]" pageId="3" pageNumber="344">
<taxonomicNameLabel id="FC870C104F556C714B6948F4FD9AF83B" box="[536,618,1929,1953]" pageId="3" pageNumber="344" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
larvae, details of mouthparts: 5, labrum, dorsal view; 6, canines and prostheca of right mandible, dorsal view; 7, canines and prostheca of left mandible, dorsal view; 8, partial view of dorsal surface of maxilla; 9, labial palp, ventral view; 10, glossa and paraglossa, ventral view.
</paragraph>
</caption>
<caption id="81BF3DF14F526C7649E649DBFC12F979" ID-DOI="http://doi.org/10.5281/zenodo.3772601" ID-Zenodo-Dep="3772601" httpUri="https://zenodo.org/record/3772601/files/figure.png" pageId="4" pageNumber="345" startId="4.[151,264,1702,1727]" targetBox="[151,1428,179,1655]" targetPageId="4">
<paragraph id="D57F6D794F526C7649E649DBFC12F979" blockId="4.[151,1437,1702,1763]" pageId="4" pageNumber="345">
<emphasis id="E7B4B16B4F526C7649E649DBFEA5F925" bold="true" box="[151,341,1702,1727]" pageId="4" pageNumber="345">FIGURES 11–14.</emphasis>
<taxonomicName id="12C016FA4F526C76482F49DAFDD7F925" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[350,551,1703,1727]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="345" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F526C76482F49DAFDD7F925" box="[350,551,1703,1727]" italics="true" pageId="4" pageNumber="345">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F526C764B4149DAFD79F925" bold="true" box="[560,649,1703,1727]" pageId="4" pageNumber="345">
<taxonomicNameLabel id="FC870C104F526C764B4149DAFD75F925" box="[560,645,1703,1727]" pageId="4" pageNumber="345" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
larvae: 11, right mandible, dorsal view; 12, left mandible, dorsal view; 13, partial view of dorsal surface of maxilla; 14, glossae and paraglossae, ventral view.
</paragraph>
</caption>
<caption id="81BF3DF14F536C7749E64AD5FE4FFA7F" ID-DOI="http://doi.org/10.5281/zenodo.3772603" ID-Zenodo-Dep="3772603" httpUri="https://zenodo.org/record/3772603/files/figure.png" pageId="5" pageNumber="346" startId="5.[151,264,1448,1473]" targetBox="[161,1435,194,1398]" targetPageId="5">
<paragraph id="D57F6D794F536C7749E64AD5FE4FFA7F" blockId="5.[151,1436,1448,1510]" pageId="5" pageNumber="346">
<emphasis id="E7B4B16B4F536C7749E64AD5FEBFFA5B" bold="true" box="[151,335,1448,1473]" pageId="5" pageNumber="346">FIGURES 15˗18.</emphasis>
<taxonomicName id="12C016FA4F536C7748254AD7FDEAFA5B" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[340,538,1449,1473]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="346" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F536C7748254AD7FDEAFA5B" box="[340,538,1449,1473]" italics="true" pageId="5" pageNumber="346">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F536C774B6E4AD4FD86FA5B" bold="true" box="[543,630,1449,1473]" pageId="5" pageNumber="346">
<taxonomicNameLabel id="FC870C104F536C774B6E4AD4FD81FA5B" box="[543,625,1449,1473]" pageId="5" pageNumber="346" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
larvae, dorsal view: 15, foreleg; 16, tarsal claw; 17, posterior margin of abdominal tergum VI; 18, paraproct.
</paragraph>
</caption>
<caption id="81BF3DF14F506C7449E64A97FEDDF9BD" ID-DOI="http://doi.org/10.5281/zenodo.3772605" ID-Zenodo-Dep="3772605" httpUri="https://zenodo.org/record/3772605/files/figure.png" pageId="6" pageNumber="347" startId="6.[151,264,1514,1539]" targetBox="[159,1419,593,1490]" targetPageId="6">
<paragraph id="D57F6D794F506C7449E64A97FEDDF9BD" blockId="6.[151,1437,1514,1575]" pageId="6" pageNumber="347">
<emphasis id="E7B4B16B4F506C7449E64A97FEA6F999" bold="true" box="[151,342,1514,1539]" pageId="6" pageNumber="347">FIGURES 19–23.</emphasis>
<taxonomicName id="12C016FA4F506C74482F4A91FDD7F999" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[350,551,1515,1539]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F506C74482F4A91FDD7F999" box="[350,551,1515,1539]" italics="true" pageId="6" pageNumber="347">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F506C744B414A96FD7AF999" bold="true" box="[560,650,1515,1539]" pageId="6" pageNumber="347">
<taxonomicNameLabel id="FC870C104F506C744B414A96FD75F999" box="[560,645,1515,1539]" pageId="6" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
larvae, tergalii shape, dorsal view, roman numbers belong to the respective tergalius pairs.
</paragraph>
</caption>
<subSubSection id="9DDA3EF24F506C7549B64905FEDDFF4D" lastPageId="7" lastPageNumber="348" pageId="6" pageNumber="347" type="diagnosis">
<paragraph id="D57F6D794F506C7549B64905FEDDFF4D" blockId="6.[151,1437,1656,2044]" lastBlockId="7.[151,1437,153,215]" lastPageId="7" lastPageNumber="348" pageId="6" pageNumber="347">
<emphasis id="E7B4B16B4F506C7449B64905FECAF909" bold="true" box="[199,314,1656,1683]" pageId="6" pageNumber="347">Diagnosis</emphasis>
. By the morphological characteristics, the larvae of
<taxonomicName id="12C016FA4F506C744AF44907FB91F909" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[901,1121,1657,1683]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F506C744AF44907FB91F909" box="[901,1121,1657,1683]" italics="true" pageId="6" pageNumber="347">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F506C744D194904FB32F909" bold="true" box="[1128,1218,1657,1683]" pageId="6" pageNumber="347">
<taxonomicNameLabel id="FC870C104F506C744D194904FB32F909" box="[1128,1218,1657,1683]" pageId="6" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguishable from those of other
<taxonomicName id="12C016FA4F506C7448DA49E3FE01F92D" authorityName="Leach" authorityYear="1815" box="[427,497,1694,1719]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="E7B4B16B4F506C7448DA49E3FE01F92D" box="[427,497,1694,1719]" italics="true" pageId="6" pageNumber="347">Baetis</emphasis>
</taxonomicName>
species by the presence of only five pairs of tergalii on segments III–VII. Larvae of
<taxonomicName id="12C016FA4F506C7449E649BFFEB7F941" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[151,327,1729,1755]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F506C7449E649BFFEB7F941" box="[151,327,1729,1755]" italics="true" pageId="6" pageNumber="347">B. pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F506C74482149BCFE5BF941" bold="true" box="[336,427,1729,1755]" pageId="6" pageNumber="347">
<taxonomicNameLabel id="FC870C104F506C74482149BCFE5BF941" box="[336,427,1729,1755]" pageId="6" pageNumber="347" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be distinguished from all Palaearctic representatives of the
<taxonomicName id="12C016FA4F506C744DF649BFFB3DF941" authorityName="Leach" authorityYear="1815" box="[1159,1229,1730,1755]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="E7B4B16B4F506C744DF649BFFB3DF941" box="[1159,1229,1730,1755]" italics="true" pageId="6" pageNumber="347">Baetis</emphasis>
</taxonomicName>
species-groups as defined by
<bibRefCitation id="B15110884F506C7448694998FDD9F89A" author="Muller-Liebenau, I." box="[280,553,1765,1792]" pageId="6" pageNumber="347" pagination="1 - 214" refId="ref4497" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815. (Insecta, Ephemeroptera). Gewass- er und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">Müller-Liebenau (1969)</bibRefCitation>
and
<bibRefCitation id="B15110884F506C744B134998FD05F965" author="Jacob U." box="[610,757,1765,1791]" pageId="6" pageNumber="347" pagination="59 - 129" refId="ref3978" refString="Jacob U. (2003) Baetis Leach 1815, sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimmungsschlusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob (2003)</bibRefCitation>
by the following combination of morphological characters: from the
<emphasis id="E7B4B16B4F506C7448734874FE78F8B9" box="[258,392,1801,1827]" italics="true" pageId="6" pageNumber="347">propingulus</emphasis>
- and
<taxonomicName id="12C016FA4F506C7448BC4874FDBCF8B9" authorityName="Eaton" authorityYear="1870" box="[461,588,1801,1827]" class="Insecta" family="Baetidae" genus="Labiobaetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="atrebatinus">
<emphasis id="E7B4B16B4F506C7448BC4874FDBCF8B9" box="[461,588,1801,1827]" italics="true" pageId="6" pageNumber="347">atrebatinus</emphasis>
</taxonomicName>
-groups by absence lateroapical protuberance on scape and incurvation on distal segment of maxillary palp, subapical bristles of claw, medioapical projection paraproct; from
<emphasis id="E7B4B16B4F506C744D884850FAA1F8DD" box="[1273,1361,1837,1863]" italics="true" pageId="6" pageNumber="347">rhodani</emphasis>
-group absence of robust setae on the antennal scape and pedicel and medioapical projection paraproct, subapical bristles of claw; from
<taxonomicName id="12C016FA4F506C74484B480BFE75F815" authorityName="Curtis" authorityYear="1834" box="[314,389,1910,1935]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="vernus">
<emphasis id="E7B4B16B4F506C74484B480BFE75F815" box="[314,389,1910,1935]" italics="true" pageId="6" pageNumber="347">vernus</emphasis>
</taxonomicName>
- and
<taxonomicName id="12C016FA4F506C7448B64808FDE2F815" authorityName="Muller-Liebenau" authorityYear="1967" box="[455,530,1909,1935]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="lutheri">
<emphasis id="E7B4B16B4F506C7448B64808FDE2F815" box="[455,530,1909,1935]" italics="true" pageId="6" pageNumber="347">lutheri</emphasis>
</taxonomicName>
- groups by the extent of medioapical protuberance on segment 2 of labial palp, form of femoral setae, which apically rounded, absent subapical bristles of claw, medioapical projection paraproct and thick, spine-like setae on gill margins; from
<taxonomicName id="12C016FA4F506C744BC748C0FCD4F84D" authorityName="Eaton" authorityYear="1870" box="[694,804,1981,2007]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="buceratus">
<emphasis id="E7B4B16B4F506C744BC748C0FCD4F84D" box="[694,804,1981,2007]" italics="true" pageId="6" pageNumber="347">buceratus</emphasis>
</taxonomicName>
-,
<taxonomicName id="12C016FA4F506C744A4D48C0FC68F84D" baseAuthorityName="Linnaeus" baseAuthorityYear="1761" box="[828,920,1981,2007]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="347" phylum="Arthropoda" rank="species" species="fuscatus">
<emphasis id="E7B4B16B4F506C744A4D48C0FC68F84D" box="[828,920,1981,2007]" italics="true" pageId="6" pageNumber="347">fuscatus</emphasis>
</taxonomicName>
- and
<emphasis id="E7B4B16B4F506C744AA648C0FBDFF84D" box="[983,1071,1981,2007]" italics="true" pageId="6" pageNumber="347">rhodani</emphasis>
-groups absence of dark band on caudal filaments and medioapical projection on paraproct (
<bibRefCitation id="B15110884F506C744A55489CFC2AF861" author="Ikonomov, P." box="[804,986,2017,2043]" pageId="6" pageNumber="347" pagination="83 - 140" refId="ref4014" refString="Ikonomov, P. (1962) Baetidae (Ephemeroptera) нa Мakidoнija. Faculte des Sciences Naturelles de lUniversite de Skopje, Biologie, 13, 83 - 140." type="journal article" year="1962">Ikonomov 1962</bibRefCitation>
;
<bibRefCitation id="B15110884F506C744A94489CFB14F861" author="Muller-Liebenau, I." box="[997,1252,2017,2043]" pageId="6" pageNumber="347" pagination="48 - 64" refId="ref4453" refString="Muller-Liebenau, I. (1967) Zur Frage der Baetis &quot; venustulus &quot;, zugleich Beschreibung der neuen Art Baetis lutheri n. sp. (Insecta, Ephemeroptera). Gewasser und Abwasser, 44 / 45, 48 - 64." type="journal article" year="1967">Müller-Liebenau 1967</bibRefCitation>
,
<bibRefCitation id="B15110884F506C744D9F489CFAD9F861" author="Muller-Liebenau, I." box="[1262,1321,2017,2043]" pageId="6" pageNumber="347" pagination="1 - 214" refId="ref4497" refString="Muller-Liebenau, I. (1969) Revision der europaischen Arten der Gattung Baetis Leach, 1815. (Insecta, Ephemeroptera). Gewass- er und Abwasser, 48 / 49, 1 - 214." type="journal article" year="1969">1969</bibRefCitation>
;
<bibRefCitation id="B15110884F506C754C45489CFE8BFF29" author="Morihara, D. K. &amp; McCafferty, W. P." lastPageId="7" lastPageNumber="348" pageId="6" pageNumber="347" pagination="130 - 135" refId="ref4275" refString="Morihara, D. K. &amp; McCafferty, W. P. (1979) Systematics of the propinquus group of Baetis species (Ephemeroptera: Baetidae). Annals of the Entomological Society of America, 72 (1), 130 - 135. https: // doi. org / 10.1093 / aesa / 72.1.130" type="journal article" year="1979">Morihara &amp; McCafferty 1979</bibRefCitation>
;
<bibRefCitation id="B15110884F516C7548F94FE4FDFCFF29" author="Jacob U." box="[392,524,153,179]" pageId="7" pageNumber="348" pagination="59 - 129" refId="ref3978" refString="Jacob U. (2003) Baetis Leach 1815, sensu stricto oder sensu lato. Ein Beitrag zum Gattungskonzept auf der Grundlage von Artengruppen mit Bestimmungsschlusseln. Lauterbornia, 47, 59 - 129." type="journal article" year="2003">Jacob 2003</bibRefCitation>
;
<bibRefCitation id="B15110884F516C754B684FE4FCF6FF29" author="Godunko, R. J. &amp; Prokopov, G. A. &amp; Soldan, T." box="[537,774,153,179]" pageId="7" pageNumber="348" pagination="231 - 248" refId="ref3801" refString="Godunko, R. J., Prokopov, G. A. &amp; Soldan, T. (2004) Mayflies of the Crimean Peninsula. III. The description of Baetis milani sp. n. with notes on taxonomy of the subgenus Rhodobaetis Jacob, 2003 (Ephemeroptera: Baetidae). Acta Zoologica Cracoviensia, 47 (3 - 4), 231 - 248. https: // doi. org / 10.3409 / 173491504783995799" type="journal article" year="2004">Godunko et al. 2004</bibRefCitation>
;
<bibRefCitation id="B15110884F516C754A624FE4FBA6FF29" author="Bauernfeind, E. &amp; Soldan, T." box="[787,1110,153,179]" pageId="7" pageNumber="348" refId="ref3642" refString="Bauernfeind, E. &amp; Soldan, T. (2012) The Mayflies of Europe (Ephemeroptera). Apollo Books, Ollerup, 781 pp. https: // doi. org / 10.1163 / 9789004260887" type="book" year="2012">Bauernfeind &amp; Soldán 2012</bibRefCitation>
;
<bibRefCitation id="B15110884F516C754D124FE4FB00FF29" author="Sroka, P." box="[1123,1264,153,179]" pageId="7" pageNumber="348" pagination="23 - 53" refId="ref4819" refString="Sroka, P. (2012) Systematics and phylogeny of the West Palaearctic representatives of subfamily Baetinae (Insecta: Ephemeroptera): Combined analysis of mitochondrial DNA sequences and morphology. Aquatic Insects, 34 (1), 23 - 53. https: // doi. org / 10.1080 / 01650424.2012.718081" type="journal article" year="2012">Sroka, 2012</bibRefCitation>
; Soldan &amp; Godunko 2012).
</paragraph>
</subSubSection>
<caption id="81BF3DF14F516C7549E64869FA6BF8B6" ID-DOI="http://doi.org/10.5281/zenodo.3772607" ID-Zenodo-Dep="3772607" box="[151,1435,1812,1837]" httpUri="https://zenodo.org/record/3772607/files/figure.png" pageId="7" pageNumber="348" startId="7.[151,262,1812,1837]" targetBox="[248,1339,237,1787]" targetPageId="7">
<paragraph id="D57F6D794F516C7549E64869FA6BF8B6" blockId="7.[151,1435,1812,1837]" box="[151,1435,1812,1837]" pageId="7" pageNumber="348">
<emphasis id="E7B4B16B4F516C7549E64869FEA1F8B7" bold="true" box="[151,337,1812,1837]" pageId="7" pageNumber="348">FIGURES 24–26.</emphasis>
<taxonomicName id="12C016FA4F516C7548274868FDE8F8B7" authority="Tiunova &amp; Semenchenko, 2019" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[342,536,1813,1837]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="pentaphyllus" status="sp. nov.">
<emphasis id="E7B4B16B4F516C7548274868FDE8F8B7" box="[342,536,1813,1837]" italics="true" pageId="7" pageNumber="348">Baetis pentaphyllus</emphasis>
</taxonomicName>
<emphasis id="E7B4B16B4F516C754B6E4868FD84F8B7" bold="true" box="[543,628,1813,1837]" pageId="7" pageNumber="348">
<taxonomicNameLabel id="FC870C104F516C754B6E4868FD9EF8B7" box="[543,622,1813,1837]" pageId="7" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
,
</emphasis>
larvae: 24, posterior margin of abdominal tergum VI; 25, tergalii I; 26, tergalii II.
</paragraph>
</caption>
<subSubSection id="9DDA3EF24F516C7549B64827FABEF87B" pageId="7" pageNumber="348" type="distribution">
<paragraph id="D57F6D794F516C7549B64827FABEF87B" blockId="7.[151,1437,1882,2018]" pageId="7" pageNumber="348">
<emphasis id="E7B4B16B4F516C7549B64827FE91F8EF" bold="true" box="[199,353,1882,1909]" pageId="7" pageNumber="348">Distribution.</emphasis>
Known from two habitats in the Far East of
<collectingCountry id="ADD72DE94F516C754A3F4826FC6DF8EF" box="[846,925,1883,1909]" name="Russia" pageId="7" pageNumber="348">Russia</collectingCountry>
: Amurskaya oblast’ (
<figureCitation id="4DFB71FC4F516C754DF54826FAF4F8EF" box="[1156,1284,1883,1909]" captionStart="FIGURES 27–28" captionStartId="8.[151,264,1856,1881]" captionTargetBox="[207,1381,181,1830]" captionTargetId="figure@8.[206,1381,181,1830]" captionTargetPageId="8" captionText="FIGURES 27–28. Baetis pentaphyllus sp. nov., localities. 28, Bolshoi Garmakan River, about 300 m above mouth, (Amurskiy Region); 29, Malyi. Garmakan River, about 700 m above mouth, (Amurskiy Region)." figureDoi="http://doi.org/10.5281/zenodo.3772609" httpUri="https://zenodo.org/record/3772609/files/figure.png" pageId="7" pageNumber="348">Figs 27–28</figureCitation>
) and
<collectingRegion id="1704A39B4F516C754C314826FF41F803" country="Russia" name="Chukotka" pageId="7" pageNumber="348">Chukotka</collectingRegion>
Autonomous District. The larvae of the new species were found in swift riffles with cobble and gravel substrate. Water temperature was 6–9 °C, depth 10–20 сm.
<emphasis id="E7B4B16B4F516C7549B648BAFEB8F87B" bold="true" box="[199,328,1991,2017]" pageId="7" pageNumber="348">Etymology</emphasis>
. The species name is derived from the Latin word
<taxonomicName id="12C016FA4F516C754AF348BAFBE1F87B" authorityName="Tiunova &amp; Semenchenko" authorityYear="2019" box="[898,1041,1991,2017]" class="Insecta" family="Baetidae" genus="Baetis" kingdom="Animalia" order="Ephemeroptera" pageId="7" pageNumber="348" phylum="Arthropoda" rank="species" species="pentaphyllus">
<emphasis id="E7B4B16B4F516C754AF348BAFBE1F87B" box="[898,1041,1991,2017]" italics="true" pageId="7" pageNumber="348">pentaphyllus</emphasis>
</taxonomicName>
, which means five-petalled.
</paragraph>
</subSubSection>
</treatment>
</document>
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