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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.930.47620" ID-GBIF-Dataset="6b7dff3a-b036-4615-ac66-8946bbfd39e2" ID-PMC="PMC7200892" ID-Pensoft-Pub="1313-2970-930-3" ID-Pensoft-UUID="CCE966F157C6546F9D8A0471493C80F6" ID-PubMed="32390746" ID-ZooBank="21AD97D78E864F0A9C205E1E4BFDDB56" ModsDocID="1313-2970-930-3" checkinTime="1588165351563" checkinUser="pensoft" docAuthor="Wesener, Thomas &amp; Anilkumar, Pooja Avinipully" docDate="2020" docId="5D5FAC5DAF07518CA816410B129D7C33" docLanguage="en" docName="ZooKeys 930: 3-35" docOrigin="ZooKeys 930" docSource="http://dx.doi.org/10.3897/zookeys.930.47620" docTitle="Zoosphaerium taitii Anilkumar &amp; Wesener 2020, sp. nov." docType="treatment" docUuid="DA735EC5-8612-4C13-A42E-910718F6ED4C" docUuidSource="ZooBank" docVersion="6" id="CCE966F157C6546F9D8A0471493C80F6" lastPageNumber="3" masterDocId="CCE966F157C6546F9D8A0471493C80F6" masterDocTitle="The millipedes collected by the Museum &quot; La Specola &quot; on Madagascar 1989 / 1991, with the description of three new species of giant pill-millipedes (Diplopoda, Sphaerotheriida, Arthrosphaeridae)" masterLastPageNumber="35" masterPageNumber="3" pageNumber="3" updateTime="1732482069575" updateUser="ExternalLinkService">
<mods:mods id="D3E57237173E03FEDEB43E80CBD0FBB0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="2FFE504857774F6B4631F857265B61C4">
<mods:title id="DAD80873EA0B2AE4D1678FF6E589D8F6">The millipedes collected by the Museum &quot; La Specola &quot; on Madagascar 1989 / 1991, with the description of three new species of giant pill-millipedes (Diplopoda, Sphaerotheriida, Arthrosphaeridae)</mods:title>
</mods:titleInfo>
<mods:name id="1CB4ACAA72FF0E84536EAA633811657E" type="personal">
<mods:role id="B51754EB2AF45CC7FB38FA6375021A4C">
<mods:roleTerm id="CE7DD44CD0D03254291081DB08D23EC0">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F1FC26C29E9A3F6F07879CD29D1A633">Wesener, Thomas</mods:namePart>
<mods:affiliation id="BA8D35B3A5D7765CC5615A1AD91A426E">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany</mods:affiliation>
</mods:name>
<mods:name id="4030EE047D589041FFF3D1475B9F436B" type="personal">
<mods:role id="43153ACF6DD30503564F7BC4850F47F1">
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</mods:role>
<mods:namePart id="32F2EDF74DEE1D6C7B32106D216182B9">Anilkumar, Pooja Avinipully</mods:namePart>
<mods:affiliation id="B91A1D5E86B31EE5BE9307F2B9B9069A">Zoological Research Museum Alexander Koenig (ZFMK), Leibniz Institute for Animal Biodiversity, Adenauerallee 160, D- 53113, Bonn, Germany</mods:affiliation>
<mods:nameIdentifier id="BBB9073E5A13658561DCEBFD07AF3436" type="orcid">https://orcid.org/0000-0001-8473-1457</mods:nameIdentifier>
<mods:nameIdentifier id="5CCB81F9966E86E450F64485E1E053F5" type="email">pooja.anilkumar54@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="38D190D2FB983B9E53B45B0A05EC1D82">text</mods:typeOfResource>
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<mods:titleInfo id="8E7E8EC96B915AB30A3A81B298FE0E7B">
<mods:title id="61CFCE065198552612A8B07ACA771339">ZooKeys</mods:title>
</mods:titleInfo>
<mods:part id="EF986FAC5D194339744D2E2E8E0A03D8">
<mods:date id="0FDD1811F04588D45DDD4BA71E1493C7">2020</mods:date>
<mods:detail id="5C6B5A7D02DAF64CFE036DE4D5156A0F" type="volume">
<mods:number id="E84C3EE389EC7A7872B5AEEBA51EECEE">930</mods:number>
</mods:detail>
<mods:extent id="15B9FB809BB070E95766C7B245ECC2EE" unit="page">
<mods:start id="3EA132BAC23BF42C762C8CA2AE2A6E51">3</mods:start>
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<mods:location id="BEC8BD87F106C1B5D2A7C45DA694DC4B">
<mods:url id="405C8CBBD344AB5867AD34FCB27968EA">http://dx.doi.org/10.3897/zookeys.930.47620</mods:url>
</mods:location>
<mods:classification id="8ECA92DA41B73DA9F3405796F8C8D1B3">journal article</mods:classification>
<mods:identifier id="E2762F22DF9B0E54E3A16347BCECD940" type="DOI">http://dx.doi.org/10.3897/zookeys.930.47620</mods:identifier>
<mods:identifier id="525ADA2082124BF1653B28A35C6D45C3" type="Pensoft-Pub">1313-2970-930-3</mods:identifier>
<mods:identifier id="34E1DBCDC5E140A27C4F5C864905E42D" type="ZooBank">21AD97D78E864F0A9C205E1E4BFDDB56</mods:identifier>
<mods:identifier id="8AA9C2431D4EFA413C8535CA2EB47CED" type="Pensoft-UUID">CCE966F157C6546F9D8A0471493C80F6</mods:identifier>
</mods:mods>
<treatment id="5D5FAC5DAF07518CA816410B129D7C33" ID-GBIF-Taxon="163875355" LSID="urn:lsid:zoobank.org:act:DA735EC5-8612-4C13-A42E-910718F6ED4C" httpUri="http://treatment.plazi.org/id/5D5FAC5DAF07518CA816410B129D7C33" lastPageNumber="3" pageId="0" pageNumber="3" scope_class="Diplopoda" scope_family="Arthrosphaeridae" scope_order="Sphaerotheriida">
<subSubSection id="3C7305C61F22AB0C749AB2A8CECFF323" pageId="0" pageNumber="3" type="nomenclature">
<paragraph id="FC2FC306614F3385536FBB6E6B6D3CD7" pageId="0" pageNumber="3">
<taxonomicName id="81571D555A5E0507D2933B5ACE222063" LSID="5D5FAC5D-AF07-518C-A816-410B129D7C33" authority="Anilkumar &amp; Wesener" authorityName="Anilkumar &amp; Wesener" authorityYear="2020" family="Arthrosphaeridae" genus="Zoosphaerium" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Zoosphaerium taitii" order="Diplopoda" pageId="0" pageNumber="3" rank="species" species="taitii" status="sp. nov.">Zoosphaerium taitii Anilkumar &amp; Wesener</taxonomicName>
<taxonomicNameLabel id="EDA9DF76666D34EB2D2CD6C7C95CE2EC" pageId="0" pageNumber="3">sp. nov.</taxonomicNameLabel>
<figureCitation id="A8F3B749F9E86A31E1CB708384FC11DA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM, Endotergum of mid body tergites, ventral view. A Zoosphaerium mangabe sp. nov., male from Marojejy B Zoosphaerium bartolozzii sp. nov., male holotype C Zoosphaerium taitii sp. nov., male holotype. Abbreviations: IA = inner area; ci = cuticular impressions; mr = marginal ridge; mb = marginal bristles." figureDoi="10.3897/zookeys.930.47620.figure5" httpUri="https://binary.pensoft.net/fig/404587" pageId="0" pageNumber="3">Figures 5C</figureCitation>
<figureCitation id="C0F329B6175BE67B3B6699649424EABD" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Zoosphaerium taitii sp. nov., male holotype, SEM, Right antenna. A lateral view B antennomeres 1 and 2 with sclerotized teeth C apical disc with four sensory cones. Abbreviations: ac = apical cone; ad = apical disc; sb = sensilla basiconica; sc-t = sclerotized teeth." figureDoi="10.3897/zookeys.930.47620.figure8" httpUri="https://binary.pensoft.net/fig/404590" pageId="0" pageNumber="3">, 8</figureCitation>
<figureCitation id="D6449DB02AEA4CAD512538C4D08D346F" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">, 9</figureCitation>
<figureCitation id="303D52FC08ED5DD2D9837EE015D14906" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">, 10</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="13C55193C80A49C036DB6AE2085DEF3E" pageId="0" pageNumber="3" type="etymology">
<paragraph id="6762CBBBE53A361518CFB5CA7E0E2D65" pageId="0" pageNumber="3">Etymology.</paragraph>
<paragraph id="744FB4687C4A67A17C6C236B2A30DA8D" pageId="0" pageNumber="3">Adjective, the species is named after the land isopod expert Dr. Stefano Taiti who collected this species.</paragraph>
</subSubSection>
<subSubSection id="0A9624D45FB215BD6F2F67FBF2CBC20A" pageId="0" pageNumber="3" type="materials_examined">
<paragraph id="4C78ED55A2CCB22C9B4765E7617AE493" pageId="0" pageNumber="3">Material examined.</paragraph>
<paragraph id="EB9A56D5B9BE5B754DC584F5F633C181" pageId="0" pageNumber="3">
<emphasis id="B0BFAB7FAAB242DC598448D9BEB999FD" bold="true" pageId="0" pageNumber="3">1</emphasis>
♂
<emphasis id="BD16968AEC1F0A70F4021ACF79A84F2E" bold="true" pageId="0" pageNumber="3">
<emphasis id="1254CC21744482BF6B816EDAB84FD3A1" italics="true" pageId="0" pageNumber="3">Holotype</emphasis>
(MZUF)
</emphasis>
, Madagascar: SW 17 km Edi Sakaraha,
<normalizedToken id="425265BB7EA5B0BEF138AAB410450BF0" originalValue="forêt">foret</normalizedToken>
de Zombitsy, foresta secca,
<geoCoordinate id="7F3A8804E018B74B1588D2C32EE64822" degrees="22" direction="south" minutes="52" orientation="latitude" precision="1" seconds="47.1" value="-22.87975">22°52'47.1&quot;S</geoCoordinate>
,
<geoCoordinate id="566364AE667A3F19BA20CD43D3D7D56F" degrees="44" direction="east" minutes="36" orientation="longitude" precision="1" seconds="41.1" value="44.611416">44°36'41.1&quot;E</geoCoordinate>
, n. Mag. 1107. Legit: Bartolozzi, S. Taiti, C. Raharimina, 15 May 1991.
</paragraph>
</subSubSection>
<subSubSection id="EB92017F80550B59FB6939C8DDE77E69" pageId="0" pageNumber="3" type="materials_examined">
<paragraph id="4FD6403AD35795F47DB12620A7567CD2" pageId="0" pageNumber="3">Other material.</paragraph>
<paragraph id="AD1057AB0AF4CB7F579A6506675B7272" pageId="0" pageNumber="3">
1 ♂,
<emphasis id="2331F8CCB9A09B06F46C0141F5771604" bold="true" pageId="0" pageNumber="3">CAS BLF Mei-99 Ma-14</emphasis>
, Province Toliara, Zombitse Nature Reserve, 16 km E Sakaraha, 825 m, tropical forest on sand,
<geoCoordinate id="9F58A2E2CCDACE086DA0B779FAA0C595" degrees="22.88231" direction="south" orientation="latitude" precision="1" value="-22.88231">22.88231°S</geoCoordinate>
,
<geoCoordinate id="188A7EFD7645530B8BC1BF01672E502A" degrees="44.70062" direction="east" orientation="longitude" precision="1" value="44.70062">44.70062°E</geoCoordinate>
, coll. E. L. Schlinger, M. E. Irwin, 15-18 Dec 1999.
</paragraph>
</subSubSection>
<subSubSection id="F4EA32AEBF49BF493636087EFC8783C7" pageId="0" pageNumber="3" type="diagnosis">
<paragraph id="9558AD1F4B1B7918D1EBA0B7B0178BDD" pageId="0" pageNumber="3">Diagnosis.</paragraph>
<paragraph id="438A44FFFB65A0A1AF6AA52C944581A1" pageId="0" pageNumber="3">
<taxonomicName id="B3A32E70B0C9D80B39337EF81EE2D79C" authorityName="Anilkumar &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoosphaerium taitii" order="Sphaerotheriida" pageId="0" pageNumber="3" phylum="Arthropoda" rank="species" species="taitii">
<emphasis id="DF405612A2A995C257D93AF6233F4262" italics="true" pageId="0" pageNumber="3">Zoosphaerium taitii</emphasis>
</taxonomicName>
sp. nov. is mostly similar to
<taxonomicName id="DCD36FEF429D371BCA52B8DAFB38A1E4" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="6AFC19FCAAC51BA0010E37FB90CB48EF" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
, both differing from all other species in the anterior telopod where sclerotized teeth are absent on the third podomere.
<taxonomicName id="C809908DDD90904F64A9F068E8B2AA84" authorityName="Anilkumar &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoosphaerium taitii" order="Sphaerotheriida" pageId="0" pageNumber="3" phylum="Arthropoda" rank="species" species="taitii">
<emphasis id="AFE4522F979902772DA7DC0224A5178C" italics="true" pageId="0" pageNumber="3">Zoosphaerium taitii</emphasis>
</taxonomicName>
sp. nov. differs from
<taxonomicName id="59DBFCC55D866915231BE8944D042CFD" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="5FE8E598AE4AA4BA5D1D88F1FD1EC8CB" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
in the shorter marginal bristles of the endotergum (protruding above the tergite margin in
<taxonomicName id="D3207F3AA78C2A2201B9CFBB41A31AB2" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="9F933990D20A09FEE4667EF43BDDC75B" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
), the higher number of ventral spines on leg 2 (four or five versus six or seven) and the slightly differently shaped anal shield (tapering in
<taxonomicName id="693960D9B0E94E1B78260760B9EF4A35" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="200F10B942ECD266661B8EF49FD7C60D" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
, well-rounded in
<taxonomicName id="274F9DF077332D3BECFA50E3AA6F1392" lsidName="Z. taitii" pageId="0" pageNumber="3" rank="species" species="taitii">
<emphasis id="6226DED72DB21F0903959213003E84E1" italics="true" pageId="0" pageNumber="3">Z. taitii</emphasis>
</taxonomicName>
sp. nov.).
</paragraph>
</subSubSection>
<subSubSection id="A24511AB8CA1F761C71326232871E30E" pageId="0" pageNumber="3" type="description">
<paragraph id="9895B0466960E603821EDF3BB0F9D995" pageId="0" pageNumber="3">Description</paragraph>
<paragraph id="8F8C27C4ABFA327D0500B9E3741D4951" pageId="0" pageNumber="3">(all measurements in mm):</paragraph>
<paragraph id="6FBB0BB17D0195E13391ACCADB31BAB6" pageId="0" pageNumber="3">Body length: holotype male: length 20.4, width 9.4 (2nd) up to 9.9 (tergite 9 = widest), height 5.4 (2nd = highest).</paragraph>
<paragraph id="ED7DCECED955ABB3C6DB2F2394F3BE0F" pageId="0" pageNumber="3">
<emphasis id="BB8B6E3AC1D6E2E1BA36CDB4BDEC0018" italics="true" pageId="0" pageNumber="3">Coloration</emphasis>
: Strongly faded due to exposure to alcohol. Antennae dark green. Legs basally brown and apically green. Head and collum light green. Tergites and anal shield faded light brown.
</paragraph>
<paragraph id="A720CAAFD6E351318A0BE90B53BAE80E" pageId="0" pageNumber="3">
<emphasis id="545C31514EE6FAFCA0D899DB45A845CA" italics="true" pageId="0" pageNumber="3">Head</emphasis>
: Eyes with 60-70 ommatidia. Antennae short, protruding up to leg 3 or 4. Size of antennomeres 1&gt;2&lt;3&gt;4&lt;5&lt;6 (Fig.
<figureCitation id="DF86A6880F8FB476815A9DB891581DA9" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Zoosphaerium taitii sp. nov., male holotype, SEM, Right antenna. A lateral view B antennomeres 1 and 2 with sclerotized teeth C apical disc with four sensory cones. Abbreviations: ac = apical cone; ad = apical disc; sb = sensilla basiconica; sc-t = sclerotized teeth." figureDoi="10.3897/zookeys.930.47620.figure8" httpUri="https://binary.pensoft.net/fig/404590" pageId="0" pageNumber="3">8A</figureCitation>
). Antennomeres 1-3 with sclerotized teeth. Antennomeres 1 and 6 with a single row of sensilla basiconica (Fig.
<figureCitation id="BE417ED751807CD876FC7BA53E5ECF4B" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Zoosphaerium taitii sp. nov., male holotype, SEM, Right antenna. A lateral view B antennomeres 1 and 2 with sclerotized teeth C apical disc with four sensory cones. Abbreviations: ac = apical cone; ad = apical disc; sb = sensilla basiconica; sc-t = sclerotized teeth." figureDoi="10.3897/zookeys.930.47620.figure8" httpUri="https://binary.pensoft.net/fig/404590" pageId="0" pageNumber="3">8B, C</figureCitation>
). Antennomeres 3-6 with long setae. Antennomere 6 with an apical disc containing four apical sensory cones (Fig.
<figureCitation id="D9F5B2B99E891725D7F5C53BB80FAB74" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Zoosphaerium taitii sp. nov., male holotype, SEM, Right antenna. A lateral view B antennomeres 1 and 2 with sclerotized teeth C apical disc with four sensory cones. Abbreviations: ac = apical cone; ad = apical disc; sb = sensilla basiconica; sc-t = sclerotized teeth." figureDoi="10.3897/zookeys.930.47620.figure8" httpUri="https://binary.pensoft.net/fig/404590" pageId="0" pageNumber="3">8B</figureCitation>
).
</paragraph>
<caption id="89BEBBAC810A8EC4F24BFAA5373C2CF8" doi="10.3897/zookeys.930.47620.figure8" httpUri="https://binary.pensoft.net/fig/404590" pageId="0" pageNumber="3" start="Figure 8" startId="F8">
<paragraph id="02E523E92AC0AF6F2EAB765CF8E48C9C" pageId="0" pageNumber="3">
<emphasis id="0397E2F53BDD3B2FD0A94D2A474FC3FF" bold="true" pageId="0" pageNumber="3">Figure 8.</emphasis>
<taxonomicName id="9468C22D9A32A990E81197A3995BF578" authorityName="Anilkumar &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoosphaerium taitii" order="Sphaerotheriida" pageId="0" pageNumber="3" phylum="Arthropoda" rank="species" species="taitii">
<emphasis id="FAB0567D0B62E47AEDD8FC27B2A2ECBD" italics="true" pageId="0" pageNumber="3">Zoosphaerium taitii</emphasis>
</taxonomicName>
sp. nov., male holotype, SEM, Right antenna.
<emphasis id="F1008E45B9789AEB9AAE0E37BF4C166A" bold="true" pageId="0" pageNumber="3">A</emphasis>
lateral view
<emphasis id="5F8931A1AF6B8BD50083AE442E19381B" bold="true" pageId="0" pageNumber="3">B</emphasis>
antennomeres 1 and 2 with sclerotized teeth
<emphasis id="2195284EB875D8EFE6A143170CAC290B" bold="true" pageId="0" pageNumber="3">C</emphasis>
apical disc with four sensory cones. Abbreviations: ac = apical cone; ad = apical disc; sb = sensilla basiconica; sc-t = sclerotized teeth.
</paragraph>
</caption>
<paragraph id="0844338E4B07F255C7434AA69864F50B" pageId="0" pageNumber="3">
<emphasis id="B725506AE6BC62BC90EE368425A78DC8" italics="true" pageId="0" pageNumber="3">Gnathochilarium</emphasis>
: Stipites and central mentum with long setae, setae absent at center of lamellae linguales. Inner palpi protruding to medial side of gnathochilarium, bearing single field of sensory cones. Rudimentary lateral palpi sharing a well-developed base bearing four sensory cones. Hypopharynx with one row of marginal teeth. Central pads apically protruding from lamellae linguales, with a median triangular incision on each pad. Posterior half of underside with single field of large sensory cones interspersed with longer, slimmer structures.
</paragraph>
<paragraph id="64430C34CBC25CFFF9C7F34A7E702E7A" pageId="0" pageNumber="3">
<emphasis id="DB4F6ED0B01AEFD0B021E60DBA0B140E" italics="true" pageId="0" pageNumber="3">Mandible</emphasis>
not dissected.
</paragraph>
<paragraph id="C2C3DA8D1E62B3F01577A341E4DCFAC0" pageId="0" pageNumber="3">
<emphasis id="406AA2A712542CA4F332D7D3FB81297E" italics="true" pageId="0" pageNumber="3">Stigmatic plates</emphasis>
: First stigmatic plate apically elliptical with marginal setae, lateral end pointed (Fig.
<figureCitation id="C0AB62D6173F9D4362D77A7130ADD28D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9A</figureCitation>
). Second stigmatic plate trapezoidal with nine spines near tracheal opening, covered with tiny setae inside and few long marginal setae (Fig.
<figureCitation id="D274537AA50DCA13F5866034316D3857" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9B</figureCitation>
).
</paragraph>
<caption id="C8AD407AACF470B3D9D7EE8E4850B516" doi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3" start="Figure 9" startId="F9">
<paragraph id="BFB4EF45B8AFF5D500E3BD2D1FD95445" pageId="0" pageNumber="3">
<emphasis id="AEAC151929313873AFC2296029C5E91C" bold="true" pageId="0" pageNumber="3">Figure 9.</emphasis>
<taxonomicName id="B3E91AFFD2665886FBE42E4E50DA20E3" authorityName="Anilkumar &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoosphaerium taitii" order="Sphaerotheriida" pageId="0" pageNumber="3" phylum="Arthropoda" rank="species" species="taitii">
<emphasis id="88CD2A9993347D20ACFE4E1445B61046" italics="true" pageId="0" pageNumber="3">Zoosphaerium taitii</emphasis>
</taxonomicName>
sp. nov., male holotype.
<emphasis id="BCD56322E8590E78F5CA5234067AE0CB" bold="true" pageId="0" pageNumber="3">A</emphasis>
,
<emphasis id="A593506AA40309C3889E5EA93F73873E" bold="true" pageId="0" pageNumber="3">B</emphasis>
Coxae of first and second right legs,
<emphasis id="EC8FB2AA42AF10D3535B3CFFD88A02C0" bold="true" pageId="0" pageNumber="3">D</emphasis>
-
<emphasis id="63FE7475747F00F8FD0AFD2DC2E23285" bold="true" pageId="0" pageNumber="3">F</emphasis>
Left anterior telopod.
<emphasis id="6FBFE18A0EFE3B223C67038FE5F04D75" bold="true" pageId="0" pageNumber="3">A</emphasis>
first stigmatic plate
<emphasis id="F07FE7B139154F31FF5D0C21A58000A4" bold="true" pageId="0" pageNumber="3">B</emphasis>
second stigmatic plate
<emphasis id="F45149E7F7C30E0C3477B58AF0207A2F" bold="true" pageId="0" pageNumber="3">C</emphasis>
9th left leg
<emphasis id="943CA42EA3FCB98353DBBDFB516B5C09" bold="true" pageId="0" pageNumber="3">D</emphasis>
anterior view
<emphasis id="1477CFA0CA2E3C1417A9A10FBFE5B976" bold="true" pageId="0" pageNumber="3">E</emphasis>
posterior view
<emphasis id="935ECF9DB44EAE092DF8332C6257114F" bold="true" pageId="0" pageNumber="3">F</emphasis>
mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="5162F22F6772F09010B82C1E54E4AE4F" pageId="0" pageNumber="3">
<emphasis id="E27336FE2E55D2BC9A2D4FEA8A3BF5F1" italics="true" pageId="0" pageNumber="3">Pleurite</emphasis>
: First pleurite weakly extending posteriorly with a well-rounded tip.
</paragraph>
<paragraph id="67A06168E24F079A5EE40A0941ABF5B1" pageId="0" pageNumber="3">
<emphasis id="460153A5049B2FBC5036DC932EF9EDB7" italics="true" pageId="0" pageNumber="3">Collum</emphasis>
: Glabrous, anterior and posterior margin with sparse rows of isolated setae.
</paragraph>
<paragraph id="AE15720161EE5BEFACC9FA035D39AF2D" pageId="0" pageNumber="3">
<emphasis id="AD2FD6906AC7BA04CEB2BD9F0A0E63DB" italics="true" pageId="0" pageNumber="3">Thoracic shield</emphasis>
: Glabrous expect for narrow lateral grooves.
</paragraph>
<paragraph id="709E0EF5A2F48FC6F9097BB73311BBC4" pageId="0" pageNumber="3">
<emphasis id="D89FF7118B3143A2E129502953458CD7" italics="true" pageId="0" pageNumber="3">Tergites</emphasis>
: Surface glabrous and shiny, chagrined. Paratergite tips not projecting.
</paragraph>
<paragraph id="775A04B92FE1F59551E92700DAD2A7F8" pageId="0" pageNumber="3">
<emphasis id="9E6301CD18431654F17838093F7205F2" italics="true" pageId="0" pageNumber="3">Endotergum</emphasis>
: Inner area with narrow conical spines, very few isolated setae. A single row of rounded-elliptical cuticular impressions. Broad smooth marginal ridge. Two rows of marginal bristle protruding towards marginal brim, few reaching tip, other few reaching 1/4-3/4 of distance to margin (Fig.
<figureCitation id="2B90C343A9AF7DD4E4C993AD204E760F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. SEM, Endotergum of mid body tergites, ventral view. A Zoosphaerium mangabe sp. nov., male from Marojejy B Zoosphaerium bartolozzii sp. nov., male holotype C Zoosphaerium taitii sp. nov., male holotype. Abbreviations: IA = inner area; ci = cuticular impressions; mr = marginal ridge; mb = marginal bristles." figureDoi="10.3897/zookeys.930.47620.figure5" httpUri="https://binary.pensoft.net/fig/404587" pageId="0" pageNumber="3">5C</figureCitation>
).
</paragraph>
<paragraph id="5C18E50E329FBDA2C5992C85DC2AA778" pageId="0" pageNumber="3">
<emphasis id="0B41F3E9B9B9CEAC2286FE5A2528BD87" italics="true" pageId="0" pageNumber="3">Anal shield</emphasis>
: Large and well rounded, surface glabrous. Two locking carinae, second carina 2.3 times longer than first, close to anal shield margin.
</paragraph>
<paragraph id="6828B76E5CFFC56A8AD4909691839EB1" pageId="0" pageNumber="3">
<emphasis id="5C57801085E2EA99F9B9175CE4F8D4C2" italics="true" pageId="0" pageNumber="3">Legs</emphasis>
: Leg 1 with four or five spines, leg 2 with six or seven spines, leg 3 with seven or eight ventral spines and an apical spine, legs 4-21 with nine ventral spines and one apical spine. In leg 9 femur 2.0, tarsus 4.7 times longer than wide. Uniform distribution of setae on all podomeres. Prefemur and femur with few long setae. Femur ridge length reaching 1/4 of femur length (Fig.
<figureCitation id="933075CB76E571DD4F5CE3BCD2D9247D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9C</figureCitation>
).
</paragraph>
<paragraph id="507CE5B1C0010B0B67E3A8C744ECF9E4" pageId="0" pageNumber="3">
<emphasis id="0A84759CF705B5F44B3C5870C79465B3" italics="true" pageId="0" pageNumber="3">Female unknown</emphasis>
.
</paragraph>
<paragraph id="0A95F6B5A99393EAE273008297F4AFE4" pageId="0" pageNumber="3">
<emphasis id="2CDC4891DE7EED5699C01D1CF4F6722A" italics="true" pageId="0" pageNumber="3">Male sexual characters</emphasis>
: Gonopore slightly oval, rounded apically, divided, reaching 1/2 length and 1/4 width of coxa, covered by 1/4 semicircular membranous plate apically and 3/4 sclerotized plate basally with few setae (Fig.
<figureCitation id="36E0601CA4AF451F58B84B6FC072C4D4" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9B</figureCitation>
).
</paragraph>
<paragraph id="22250E891DCA1EB5442D24B731E7477A" pageId="0" pageNumber="3">
<emphasis id="26B396042B3E0F05CFB54EAC5868B65C" italics="true" pageId="0" pageNumber="3">Anterior telopod</emphasis>
: Harp carrying one stridulation rib positioned medially with one end pointing laterad. Podomere 1 broad with marginal setae, few setae above stridulation rib (Fig.
<figureCitation id="477872E0647CCB9DBEEB7BE56DCB4C70" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9D</figureCitation>
) and a shallow mesal cavity laterally (Fig.
<figureCitation id="A0E6017845782ED4D8D52BE2892E6966" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9E</figureCitation>
). Podomere 2, process visible in anterior view, reaching 2/3rd of length of podomere 3 (Fig.
<figureCitation id="C02BD20894BC92B63A0C77BA135AD237" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9D</figureCitation>
), with sclerotized nubs along apical-mesal margin and one basal spine present below field of spots (Fig.
<figureCitation id="D7DD8D1ED8A477FE64881DAB7BAD6A01" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9F</figureCitation>
). Podomere 3 apically broad, rounded, longer than podomere 2 (Figs
<figureCitation id="BD134484538F297B66394B03107A7036" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9F</figureCitation>
,
<figureCitation id="3631EDB1671FEF81B1C359118A1C2878" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10G</figureCitation>
), and carrying one dark sclerotized spot at apical tip (Fig.
<figureCitation id="4AE28C276ED672688F07CF30CC204F9B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9F</figureCitation>
). Field of sclerotized spots run along apical-basal margin with four spines out of six visible (Figs
<figureCitation id="F30A43501AA01E295B6081129C42E89D" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Zoosphaerium taitii sp. nov., male holotype. A, B Coxae of first and second right legs, D - F Left anterior telopod. A first stigmatic plate B second stigmatic plate C 9 th left leg D anterior view E posterior view F mesal view. Abbreviations: as = apical spine; cx = coxa; ds-p = dark sclerotized spot; fe = femur; f-rdg = femur ridge; gp = gonopore; mem-p = apical membranous part of plate covering gonopore; pf = prefemur; po = postfemur; scl-p = sclerotized plate; sp-p = second podomere process; sr = stridulation rib; ss = sclerotized spot; st = stigmatic plate; syn = syncoxite; ta = tarsus; ti = tibia; to = tracheal opening. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure9" httpUri="https://binary.pensoft.net/fig/404591" pageId="0" pageNumber="3">9F</figureCitation>
,
<figureCitation id="B8AB20B51220B4972BA58DC89330BBA4" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10G</figureCitation>
). Three spines positioned at apical margin above field of spots, one in middle of cavity with tip protruding to sclerotized spot, one small single spine in lateral projected area slightly towards base, and one spine at base of field of spots (Fig.
<figureCitation id="9F0CF0A931CBBECFC467594C573A5844" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10G</figureCitation>
). Podomeres 2 and 3 covered with setae except mesally (Fig.
<figureCitation id="E78BD8CC033239D646FB5776456A8B97" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10G</figureCitation>
).
</paragraph>
<caption id="1472EC2D5F5ED8286024F5A94B533B6D" doi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3" start="Figure 10" startId="F10">
<paragraph id="1AFAC4EB29F9AD82643BA4AB09BD01FA" pageId="0" pageNumber="3">
<emphasis id="A8EDD5973E563CA7B3A55D397DF210E8" bold="true" pageId="0" pageNumber="3">Figure 10.</emphasis>
<taxonomicName id="093AA51E15315515BBC9B1F619BC1AEE" authorityName="Anilkumar &amp; Wesener" authorityYear="2020" class="Diplopoda" family="Arthrosphaeridae" genus="Zoosphaerium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoosphaerium taitii" order="Sphaerotheriida" pageId="0" pageNumber="3" phylum="Arthropoda" rank="species" species="taitii">
<emphasis id="ED194C4C92992BC8A451C3C51E23C831" italics="true" pageId="0" pageNumber="3">Zoosphaerium taitii</emphasis>
</taxonomicName>
sp. nov., male holotype, left anterior telopod.
<emphasis id="DFF5F1071E19D9F3DDE2CB6C18975E43" bold="true" pageId="0" pageNumber="3">G</emphasis>
lateral view.
<emphasis id="8026D4D0175A6A6EFB88FAA1D9DB5D61" bold="true" pageId="0" pageNumber="3">H</emphasis>
,
<emphasis id="75DC1304E3F32027D4726682BDD7BE47" bold="true" pageId="0" pageNumber="3">I</emphasis>
Left posterior telopod.
<emphasis id="C97FEDD7FAE20740CC375B5136C12CE8" bold="true" pageId="0" pageNumber="3">H</emphasis>
posterior view
<emphasis id="76C199FFEE910A1F6DDACC0D492F8B50" bold="true" pageId="0" pageNumber="3">I</emphasis>
anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm.
</paragraph>
</caption>
<paragraph id="65E9EE7DBFE809FFA2EAE7FEA2A1B55A" pageId="0" pageNumber="3">
<emphasis id="EA32C451A931F6B1E091628EEAF3310C" italics="true" pageId="0" pageNumber="3">Posterior telopod</emphasis>
: Movable finger 2.5 times longer than wide with tip slightly curving towards immovable finger. Apical tip with a dark sclerotized spot, eight sclerotized crenulated teeth (arranged in three groups), three mesal spines (two merged at tip), with very few setae at base and a shallow mesal cavity with two membranous lobes (Fig.
<figureCitation id="3B89AC5575FD0A3FA2F71B4364E550D3" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10H, I</figureCitation>
). First five teeth positioned below apically merged spines, other two teeth below middle spine, located in middle of first membranous lobe, two isolated single tooth located between both membranous lobes (Fig.
<figureCitation id="28188A774D4B7851799A10D6861EEC43" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10H</figureCitation>
). Immovable finger 4.3 times longer than wide, reaching 3/4th of length of movable finger, slender, tip curved towards movable finger with a row of small sclerotized spots along 1/4 apical mesal margin, covered with setae in posterior aspect (Fig.
<figureCitation id="8881770B6B803489D9752F870B741B07" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Zoosphaerium taitii sp. nov., male holotype, left anterior telopod. G lateral view. H, I Left posterior telopod. H posterior view I anterior view. Abbreviations: cr-t = crenulated teeth; ds-p = dark sclerotized spot; imm-f = immovable finger; mf = movable finger; ml = membranous lobes; s-ca = shallow cavity sn = sclerotized nubs; ss = sclerotized spot. Scale bars: 1 mm." figureDoi="10.3897/zookeys.930.47620.figure10" httpUri="https://binary.pensoft.net/fig/404592" pageId="0" pageNumber="3">10H</figureCitation>
). Podomere 1 glabrous except for very few marginal setae.
</paragraph>
</subSubSection>
<subSubSection id="D7A2FFF4AA0DEE37E427984A32594D24" pageId="0" pageNumber="3" type="remarks">
<paragraph id="AD4C84D3DAD10317556C736C2C9FB2CA" pageId="0" pageNumber="3">Remarks.</paragraph>
<paragraph id="C152F288ED86F5F9D2963EB798122543" pageId="0" pageNumber="3">
This species was described as a population of
<taxonomicName id="853804BC68ABBA9C29DAD4B4AF88364E" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="8AC093A88DCDE98BE9E6CC4DBF8EA9C9" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
in a previous study, already with a remark that the status of the population should be evaluated when more male specimens become available (
<bibRefCitation id="7A50B4A34C52BEA4F6523D78B3E9043D" DOI="https://doi.org/10.11646/zootaxa.2097.1.1" author="Wesener, T" journalOrPublisher="Zootaxa" pageId="0" pageNumber="3" pagination="1 - 131" refId="B55" refString="Wesener, T, 2009. Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on micro-endemism (Diplopoda, Sphaerotheriida). Zootaxa 2097: 1 - 131, DOI: https://doi.org/10.11646/zootaxa.2097.1.1" title="Unexplored richness: discovery of 31 new species of Giant Pill-Millipedes endemic to Madagascar, with a special emphasis on micro-endemism (Diplopoda, Sphaerotheriida)." url="https://doi.org/10.11646/zootaxa.2097.1.1" volume="2097" year="2009">Wesener 2009</bibRefCitation>
). With the additional male specimen available from the collections of the &quot;La Specola&quot; Museum, we feel more confident in describing the Zombitse specimens as a species separate from
<taxonomicName id="C4EC7C87A3F26A51D990718F3322A1B1" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="9BC2609773613A4880991E401B69FF8B" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
. The species lives in sympatry with
<taxonomicName id="51CCDD85CBCA936F9FE89D18A94859E9" lsidName="Z. album" pageId="0" pageNumber="3" rank="species" species="album">
<emphasis id="030D71ADB8CD6DD86AFEED9D0F7E3370" italics="true" pageId="0" pageNumber="3">Z. album</emphasis>
</taxonomicName>
Wesener, 2009, a species belonging to a different species group (
<bibRefCitation id="882A7C784CE18CBFA7B14B68CA13496A" author="Wesener, T" journalOrPublisher="Bonn Zoological Bulletin Supplementum" pageId="0" pageNumber="3" pagination="1 - 104" refId="B58" refString="Wesener, T, 2016. The Giant Pill-Millipedes, order Sphaerotheriida - An annotated species catalogue with morphological atlas and list of apomorphies (Arthropoda: Diplopoda). Bonn Zoological Bulletin Supplementum 63: 1 - 104" title="The Giant Pill-Millipedes, order Sphaerotheriida - An annotated species catalogue with morphological atlas and list of apomorphies (Arthropoda: Diplopoda)." volume="63" year="2016">Wesener 2016</bibRefCitation>
).
</paragraph>
<caption id="09FF8801A769263A849B0ACE01C8C169" ID-Table-UUID="09FF8801A769263A849B0ACE01C8C169" httpUri="http://table.plazi.org/id/09FF8801A769263A849B0ACE01C8C169" pageId="0" pageNumber="3" start="Table 2" startId="T2">
<paragraph id="9C6FA5C7FE14EAB0978A9CE937A642CF" pageId="0" pageNumber="3">
<emphasis id="6E34CB9CA315BAAEF94EEC39FFA2C131" bold="true" pageId="0" pageNumber="3">Table 2.</emphasis>
Comparison of
<taxonomicName id="D8AD9E3316677F48B0BECDA899A2FAD0" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="CCAED6E11EFAB6FA92C752EAF375880C" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
Wesener, 2009,
<taxonomicName id="DB9C5B820D45D2CF471DC7163F8BEAA3" lsidName="Z. bilobum" pageId="0" pageNumber="3" rank="species" species="bilobum">
<emphasis id="FEF6A85620F2C3B3E02999CDE41FD456" italics="true" pageId="0" pageNumber="3">Z. bilobum</emphasis>
</taxonomicName>
Wesener, 2009,
<taxonomicName id="4313DC9F40675027FF3446088A0300D0" lsidName="Z. tigrioculatum" pageId="0" pageNumber="3" rank="species" species="tigrioculatum">
<emphasis id="DB67BEE75F2EACED2EF1FCDDEBEDEE02" italics="true" pageId="0" pageNumber="3">Z. tigrioculatum</emphasis>
</taxonomicName>
Wesener &amp; Bespalova, 2010,
<taxonomicName id="862E5DFE4C157F671D249B62A676FA3E" lsidName="Z. bartolozzii" pageId="0" pageNumber="3" rank="species" species="bartolozzii">
<emphasis id="76B0F95F89B5182EF5CF51BD1A5198CB" italics="true" pageId="0" pageNumber="3">Z. bartolozzii</emphasis>
</taxonomicName>
sp. nov., and
<taxonomicName id="6ECDE1F6F57276C8D44920118C89B7B9" lsidName="Z. taitii" pageId="0" pageNumber="3" rank="species" species="taitii">
<emphasis id="510FC65E6E3D7C2B48697DA2078032BA" italics="true" pageId="0" pageNumber="3">Z. taitii</emphasis>
</taxonomicName>
sp. nov. Abbreviations: ANT - Antenna; aT - anterior telopod, bas - basiconica, Endo - endotergum. Modified after
<bibRefCitation id="32903872E341636FF5C1710D1EAA376E" DOI="https://doi.org/10.5733/afin.051.0102" author="Wesener, T" journalOrPublisher="African Invertebrates" pageId="0" pageNumber="3" pagination="133 - 161" refId="B68" refString="Wesener, T, Bespalova, I, Sierwald, P, 2010b. Madagascar's living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium). African Invertebrates 51 (1): 133 - 161, DOI: https://doi.org/10.5733/afin.051.0102" title="Madagascar's living giants: discovery of five new species of endemic giant pill-millipedes from Madagascar (Diplopoda: Sphaerotheriida: Arthrosphaeridae: Zoosphaerium)." url="https://doi.org/10.5733/afin.051.0102" volume="51" year="2010 b">Wesener et al. 2010b</bibRefCitation>
.
</paragraph>
</caption>
<paragraph id="7C18EE1CA5D3088B0FD52E049C8F207E" pageId="0" pageNumber="3">
<table id="330047E45B85E506B43C56C0E5D6F460" pageId="0" pageNumber="3">
<tr id="504F2F35A4124C067F918887E9253064" pageId="0" pageNumber="3">
<td id="AE01FD9CF70E2DE434791E285EF711C8" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="A8E5A0B27D58BFCFD6B48F642697CB45" bold="true" pageId="0" pageNumber="3">Character</emphasis>
</td>
<td id="24B7538D9802DA536B8511C1E58B81F5" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<taxonomicName id="6CDAC01971FF31B19509F213D8B0E312" lsidName="Z. isalo" pageId="0" pageNumber="3" rank="species" species="isalo">
<emphasis id="3ABD15619DE7A4276AB1D257DAA69248" bold="true" italics="true" pageId="0" pageNumber="3">Z. isalo</emphasis>
</taxonomicName>
</td>
<td id="A76A807096E4076F83F91A8148F5C8ED" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<taxonomicName id="948A9A2DD06FC3B9D924943AE57FA60E" lsidName="Z. bilobum" pageId="0" pageNumber="3" rank="species" species="bilobum">
<emphasis id="49B7BAEC6C3D8153BD27EE626B6BDD89" bold="true" italics="true" pageId="0" pageNumber="3">Z. bilobum</emphasis>
</taxonomicName>
</td>
<td id="EFC8C6B68FB2C32F427549BFC0974C2D" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<taxonomicName id="F6FE1103F1EB1024D986432749C49AA3" lsidName="Z. tigrioculatum" pageId="0" pageNumber="3" rank="species" species="tigrioculatum">
<emphasis id="4BBB9ADDC0B9906C272A9B28B5CB96B1" bold="true" italics="true" pageId="0" pageNumber="3">Z. tigrioculatum</emphasis>
</taxonomicName>
</td>
<td id="4DCD3E3F4840F08E1F8A0C3F8A85D941" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="3CD7CCA5A2F42BCE416236670DDD4748" bold="true" pageId="0" pageNumber="3">
<taxonomicName id="AB7C09658B1CC8F84BD7F07D14C736D5" lsidName="Z. bartolozzii" pageId="0" pageNumber="3" rank="species" species="bartolozzii">
<emphasis id="5F59EE523252B58C878CDC55460F7297" italics="true" pageId="0" pageNumber="3">Z. bartolozzii</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
<td id="37461335224D6FF93C70E60DD68AF1C1" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="55C4F9DE66203A033F3833DF2ED847E6" bold="true" pageId="0" pageNumber="3">
<taxonomicName id="8589F91FC30821100A59666DA259B84D" lsidName="Z. taitii" pageId="0" pageNumber="3" rank="species" species="taitii">
<emphasis id="ED8D5362A4962F7D92CC2AAD7C032EC3" italics="true" pageId="0" pageNumber="3">Z. taitii</emphasis>
</taxonomicName>
sp. nov.
</emphasis>
</td>
</tr>
<tr id="CE4A4F09D574C8E60D9B1063B06DCDAE" pageId="0" pageNumber="3">
<td id="19C9D8908E61F5D062448267E12699D7" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="15E1E5C7F6E5BFD2BA7F7D69E2DDE4E5" bold="true" pageId="0" pageNumber="3">Shape of anal shield</emphasis>
</td>
<td id="340038FDDB746842FF3CC43AA8B777E1" colspan="1" pageId="0" pageNumber="3" rowspan="1">Tapering</td>
<td id="284FFD42666BED6D1AD0A82E23F9002B" colspan="1" pageId="0" pageNumber="3" rowspan="1">Tapering</td>
<td id="FCE46C43036219C61D57EE2C52C9F7F8" colspan="1" pageId="0" pageNumber="3" rowspan="1">Weakly bell shaped</td>
<td id="C7AF1B32B2BAC64AE196F891EA2529DB" colspan="1" pageId="0" pageNumber="3" rowspan="1">Well rounded</td>
<td id="413621A95045AF8E0E837C6338D87B5B" colspan="1" pageId="0" pageNumber="3" rowspan="1">Well rounded</td>
</tr>
<tr id="785C0E7C18040012F3BF9D91EDC261E6" pageId="0" pageNumber="3">
<td id="65A847FF446B9DD469C54B36BFCCC4A8" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="2EE3063AF99A5C4190961A1692491512" bold="true" pageId="0" pageNumber="3">Locking carinae</emphasis>
</td>
<td id="81FE8EF0D17D6EEF0208A8DE94B31718" colspan="1" pageId="0" pageNumber="3" rowspan="1">
2nd 3
<normalizedToken id="5E86A1ECAE5369A222549C9EB10E2A4E" originalValue="×">x</normalizedToken>
1st
</td>
<td id="8C8CBC9A44D0DCD7CF29CEC908EA2974" colspan="1" pageId="0" pageNumber="3" rowspan="1">
2nd 2.5
<normalizedToken id="89DA6676282B66346CEC38C6FD4F66FD" originalValue="×">x</normalizedToken>
1st
</td>
<td id="BB4C22A4DD633EDF6A4772BCCF80DD72" colspan="1" pageId="0" pageNumber="3" rowspan="1">
2nd 4
<normalizedToken id="D314E7DDD672EE596EB83872EF795726" originalValue="×">x</normalizedToken>
1st
</td>
<td id="9B8C528D9628C7FF7187EE0295A95517" colspan="1" pageId="0" pageNumber="3" rowspan="1">
2nd 3.5
<normalizedToken id="6014E3022DD356F4C56952771B054F82" originalValue="×">x</normalizedToken>
1st
</td>
<td id="EE5BDEC20DEC704C282A4B2C84D12DAA" colspan="1" pageId="0" pageNumber="3" rowspan="1">
2nd 2.3
<normalizedToken id="379DC357E463DE3AF5350677E297C21C" originalValue="×">x</normalizedToken>
1st
</td>
</tr>
<tr id="3BDDB94F39BA1FA84EE786C9A008B3DE" pageId="0" pageNumber="3">
<td id="FB3075BC1444DFFAF5F834F48FB3BE3E" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="B1B8C6BA5FE97211D5B793EE7830D191" bold="true" pageId="0" pageNumber="3">1st leg no. of ventral spines</emphasis>
</td>
<td id="42BB496EF11F1BAEB4FA21449C6F2242" colspan="1" pageId="0" pageNumber="3" rowspan="1">3 or 4</td>
<td id="06B051791F57284A66853C32F99C9D28" colspan="1" pageId="0" pageNumber="3" rowspan="1">6 or 7</td>
<td id="27A2F21AEAB4618FF1393D48BD5D611D" colspan="1" pageId="0" pageNumber="3" rowspan="1">4 or 5</td>
<td id="2E7BF2031D1BB7AC88DAB083744EFA45" colspan="1" pageId="0" pageNumber="3" rowspan="1">3 or 4</td>
<td id="A33AB0B32D766CED17A20C5212B13F01" colspan="1" pageId="0" pageNumber="3" rowspan="1">4 or 5</td>
</tr>
<tr id="204E4577F43D5B403AE8554E8581E5E0" pageId="0" pageNumber="3">
<td id="3F9B49755092558DBF939B924E58CB11" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="55EFCF3EB57E3A1BDD2D23EAC8E46329" bold="true" pageId="0" pageNumber="3">2nd leg no. of ventral spines</emphasis>
</td>
<td id="07E103E137330658A8C35C6B1CD0083D" colspan="1" pageId="0" pageNumber="3" rowspan="1">4 or 5</td>
<td id="6162D68A18511FB57AED710B8B5BA5A1" colspan="1" pageId="0" pageNumber="3" rowspan="1">8 or 9</td>
<td id="3F86CCFE0788108C4E9A7253BC0CC5D5" colspan="1" pageId="0" pageNumber="3" rowspan="1">6 or 7</td>
<td id="8A38DCA8362E2B2770978552E83A2D64" colspan="1" pageId="0" pageNumber="3" rowspan="1">6 or 7</td>
<td id="F1EBB8F3308FE05C31DC814D4952FCCB" colspan="1" pageId="0" pageNumber="3" rowspan="1">6 or 7</td>
</tr>
<tr id="A3868925E9191A6BB10DB4F1FA10CAA4" pageId="0" pageNumber="3">
<td id="AAFF60FCB7A297FE39CE49F38F2546AC" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="8B8128C5267216832ACDE35F07EEBD60" bold="true" pageId="0" pageNumber="3">aT, 2nd podomere in av</emphasis>
</td>
<td id="EA07DAA546CB1636CDA8A0165C35434D" colspan="1" pageId="0" pageNumber="3" rowspan="1">visible</td>
<td id="05BB1C50EF0F259F0E01A022D8356647" colspan="1" pageId="0" pageNumber="3" rowspan="1">Not visible</td>
<td id="1F964568B4DAED52877D88F4AC450228" colspan="1" pageId="0" pageNumber="3" rowspan="1">visible</td>
<td id="E105D436AD8EA832F47249AC91E73DD1" colspan="1" pageId="0" pageNumber="3" rowspan="1">visible</td>
<td id="47982B91003F4F95A17F8DFD2C27C477" colspan="1" pageId="0" pageNumber="3" rowspan="1">visible</td>
</tr>
<tr id="99F335FF1BF585F72C5FE34FD5800694" pageId="0" pageNumber="3">
<td id="70913382F5AC36217EA59131E1B1ACF5" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="EB0B88970994EFF6A7388B40F6496FBE" bold="true" pageId="0" pageNumber="3">aT, 3rd podomere in av</emphasis>
</td>
<td id="C0F6DE46FB714748EFBCB00E00B7325E" colspan="1" pageId="0" pageNumber="3" rowspan="1">without crenulated teeth</td>
<td id="FB8D446A9A24DC6C4D3FBDB47D3C9CD1" colspan="1" pageId="0" pageNumber="3" rowspan="1">without crenulated teeth</td>
<td id="CBAE5072BAAA415C15D60216360B69A1" colspan="1" pageId="0" pageNumber="3" rowspan="1">with crenulated teeth</td>
<td id="1DEB6454D737A1B7B707D20B5E63478B" colspan="1" pageId="0" pageNumber="3" rowspan="1">with crenulated teeth</td>
<td id="44B562679F24EEE8031F3FE4F1837CBE" colspan="1" pageId="0" pageNumber="3" rowspan="1">without crenulated teeth</td>
</tr>
<tr id="4EDA1468ACDE552F8DDE5F1389BC7105" pageId="0" pageNumber="3">
<td id="66E574DC670DE96D7B159FDB77FEE367" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="E610DB86AA9ED2EE1419735DC0A5D706" bold="true" pageId="0" pageNumber="3">ANT, sclerotized teeth</emphasis>
</td>
<td id="9BE1C6C13B69AF80732A7562C66F3245" colspan="1" pageId="0" pageNumber="3" rowspan="1">on antennomeres 1-4</td>
<td id="299F9CCC21A19399963ECEAAD8C9A286" colspan="1" pageId="0" pageNumber="3" rowspan="1">on antennomeres 1-5</td>
<td id="3729497A0A45D36B34BD5FBC8EF095FC" colspan="1" pageId="0" pageNumber="3" rowspan="1">on antennomeres 1-3</td>
<td id="B4F90DCF7934B50FC61E8964D590046E" colspan="1" pageId="0" pageNumber="3" rowspan="1">on antennomeres 1-3</td>
<td id="554630651BB447394187156CD8D34EDF" colspan="1" pageId="0" pageNumber="3" rowspan="1">on antennomeres 1-3</td>
</tr>
<tr id="E4957EBAD27CDB1440C8D7348117B57F" pageId="0" pageNumber="3">
<td id="302CB8BC1A8D0B3C5B5F19350CB9C255" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="B73A112F547CCCF1C062D1A36DA88E7D" bold="true" pageId="0" pageNumber="3">ANT, sensilla bas.</emphasis>
</td>
<td id="79ADE6C509EDC402506AE5A575B0F6FC" colspan="1" pageId="0" pageNumber="3" rowspan="1">only on 1st</td>
<td id="D5EEBB06B95AF502BAEE77DB88B7AD7D" colspan="1" pageId="0" pageNumber="3" rowspan="1">only on 1st and 5th</td>
<td id="8D1F355BE07DFCC36CA95C3C80509073" colspan="1" pageId="0" pageNumber="3" rowspan="1">on 1st and 2nd</td>
<td id="B383B1E0F5E3273C6EE9506DA866DD75" colspan="1" pageId="0" pageNumber="3" rowspan="1">absent</td>
<td id="891F74CAC27F1B3EB1F1852340E09FE0" colspan="1" pageId="0" pageNumber="3" rowspan="1">only on 1st</td>
</tr>
<tr id="F4D41C89145B53A89ED15A38BC8734F8" pageId="0" pageNumber="3">
<td id="A27655F9139B4A0B3744EC78097FDE2A" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="929B27339B8D09CF7C93395D8C2A2127" bold="true" pageId="0" pageNumber="3">Endo, marginal bristle</emphasis>
</td>
<td id="412A5EF1D970DE6750892A2F194A24E5" colspan="1" pageId="0" pageNumber="3" rowspan="1">protruding slightly above margin</td>
<td id="65EED7811B21F21B85DBCD4A28A651DB" colspan="1" pageId="0" pageNumber="3" rowspan="1">extending beyond margin</td>
<td id="B8A845B527EEFE6C5FB66182AD943AF1" colspan="1" pageId="0" pageNumber="3" rowspan="1">extending beyond</td>
<td id="10653D67B62FA22D52429CD57F2E3BFD" colspan="1" pageId="0" pageNumber="3" rowspan="1">protruding to margin</td>
<td id="AC1FEE66906D8CC0704CC4FBFE1149CE" colspan="1" pageId="0" pageNumber="3" rowspan="1">protruding to margin</td>
</tr>
<tr id="70DFDDE3C5257125946AC572B1E52714" pageId="0" pageNumber="3">
<td id="6FF289C9CC8E9EBBFE476C14ED1590A7" colspan="1" pageId="0" pageNumber="3" rowspan="1">
<emphasis id="15E77483E7DC30E3CEF65A732177FB06" bold="true" pageId="0" pageNumber="3">Endo, cuticular patterns</emphasis>
</td>
<td id="F95791181DDE01FA8966075F74C92A0A" colspan="1" pageId="0" pageNumber="3" rowspan="1">single row</td>
<td id="5EC968BF2108F1347E6A616950394315" colspan="1" pageId="0" pageNumber="3" rowspan="1">two rows</td>
<td id="2D199C261B3A966FD5E1404F1A70F95D" colspan="1" pageId="0" pageNumber="3" rowspan="1">two rows</td>
<td id="3C41942FD3D731F9B8F1D88C07C81B00" colspan="1" pageId="0" pageNumber="3" rowspan="1">single row</td>
<td id="9E754EDBE4FB81DEFB77D18FF0A71500" colspan="1" pageId="0" pageNumber="3" rowspan="1">single row</td>
</tr>
</table>
</paragraph>
</subSubSection>
</treatment>
</document>
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