145 lines
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145 lines
18 KiB
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<document id="F2CD35DEA22204A99127E75F98E21D75" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="5D58FCF498BFD8D6B55F23E0AE560AA5" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus propinquus Omer-Cooper 1958" docType="treatment" docVersion="5" lastPageNumber="72" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="71" updateTime="1701314941532" updateUser="plazi">
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<mods:titleInfo id="1BEBFF170FE11C89B86CB89B0970CCE7">
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<mods:title id="1BDAE59E2C788BF597E59128F551E519">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="AFDE015C7634A7BC8A49CB82FB5E3793">Bistroem, Olof</mods:namePart>
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<mods:namePart id="C0ED17EE162446E9B317394134D0AECE">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="0032005DBFE4B92623487E20448B8286">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="590020154DB80E8E2132B3A6BCA06656">text</mods:typeOfResource>
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<mods:date id="E6DC141690A7AF6948CF9ECB206494D2">2015</mods:date>
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<mods:detail id="E7C1CBB68ABA4DBA48820392AF05FE80" type="volume">
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<mods:number id="BB7D1744459C7E30014BA2FB9F6F8CED">542</mods:number>
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<mods:url id="831A11D5730F982FF05B45F635D67447">http://dx.doi.org/10.3897/zookeys.542.5975</mods:url>
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<mods:classification id="E73A93ED7C88E400FA1846D5AB0D232F">journal article</mods:classification>
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<mods:identifier id="0C41242EE3EF6CC36B53AC66C2317FAE" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="5D58FCF498BFD8D6B55F23E0AE560AA5" ID-GBIF-Taxon="127862794" LSID="urn:lsid:plazi:treatment:5D58FCF498BFD8D6B55F23E0AE560AA5" httpUri="http://treatment.plazi.org/id/5D58FCF498BFD8D6B55F23E0AE560AA5" lastPageId="71" lastPageNumber="72" pageId="70" pageNumber="71">
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<subSubSection id="DA105E258D81B2A437393DD1B27C7F12" pageId="70" pageNumber="71" type="multiple">
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<paragraph id="60C611E713C05914C06D60F12ED00337" pageId="70" pageNumber="71">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="8D7C0DF9BEDAB58BF486A52764837F4B" pageId="70" pageNumber="71" type="nomenclature">
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<paragraph id="91D8EC9957868936070D1D37F3EACEFF" pageId="70" pageNumber="71">
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<taxonomicName id="0068880EEBE4654895DFEA73E7B1D218" ID-CoL="6NRNP" authority="Omer-Cooper, 1958" authorityName="Omer-Cooper" authorityYear="1958" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="70" pageNumber="71" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus Omer-Cooper, 1958</taxonomicName>
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Figs 57-58, 254, 413, 540
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</paragraph>
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</subSubSection>
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<subSubSection id="56509AE6519C0572E2C4A782AF7651BF" pageId="71" pageNumber="72" type="reference_group">
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<paragraph id="2D0A24C6E4B890BA1050643BD91C35A9" pageId="71" pageNumber="72">
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<taxonomicName id="35538B9BB540B9F19E79A76CA138078A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propinquus">
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<pageBreakToken id="15E2AC9EF6CB0B9472ADE5C53B8A4AEF" pageId="71" pageNumber="72" start="start">Laccophilus</pageBreakToken>
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propinquus
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</taxonomicName>
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<bibRefCitation id="ECC46E4CE5AF3A1AF2822621720CE880" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="260" pageNumber="261" pagination="36 - 55" title="Dytiscidae from Nyasaland and Southern Rhodesia. III. Laccophilinae." volume="21" year="1958 b">Omer-Cooper 1958b</bibRefCitation>
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: 37, 43, 45, 46 (original description, faunistics);
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<bibRefCitation id="DF5CCE8DD10EB43AB6DD648ACC774F03" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper 1965</bibRefCitation>
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: 77, 82 (description, faunistics);
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<bibRefCitation id="3215E9DEC523390F2323DECA3FC41DA1" author="Omer-Cooper, Joyce" journalOrPublisher="Journal of the Entomological Society of Southern Africa" pageId="261" pageNumber="262" pagination="285 - 294" title="New and little known species of African Laccophilus (Coleoptera: Dytiscidae)." volume="33" year="1970">Omer-Cooper 1970</bibRefCitation>
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: 288, 289, 290 (discussion, description, faunistics);
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<bibRefCitation id="82600E7F67B77077F07E4D2D8CDCCD62" author="Pederzani, F" journalOrPublisher="Atti dell'Accademia Roveretana degli Agiati, Contribute della Classe di Scienze Matematiche, Fisiche e Naturali" pageId="261" pageNumber="262" pagination="103 - 118" title="Dytiscidae collected in Zambia and description of Hyphydrusfluviatilis sp. n. and Hyphydruszambiensis sp. n. (ColeopteraDytiscidae)." volume="26" year="1988">Pederzani 1988</bibRefCitation>
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: 107 (faunistics, biology);
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<bibRefCitation id="E5FE15B9A27F9EBF9DC0EC51AEB5BAB7" author="Nilsson, AN" journalOrPublisher="World catalogue of Insects" pageId="260" pageNumber="261" pagination="1 - 395" title="Dytiscidae." volume="3" year="2001">Nilsson 2001</bibRefCitation>
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: 249 (catalogue, faunistics);
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<bibRefCitation id="05AE0A7FEAB9FA72A5B220780E3949A2" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 216 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="7A70E6B2E09450193687D210B47AA24E" pageId="71" pageNumber="72" type="type locality">
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<paragraph id="3A527C28E37458F4D754386FB83E0B22" pageId="71" pageNumber="72">Type locality.</paragraph>
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<paragraph id="21DFDE5B7805699DA8A207BB63DF2543" pageId="71" pageNumber="72">Malawi: Mwanza.</paragraph>
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</subSubSection>
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<subSubSection id="E716E1D69D265DAD2D163714D07C9089" pageId="71" pageNumber="72" type="type material">
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<paragraph id="02FD864D6CA4E545C729D3E6F37FBBF0" pageId="71" pageNumber="72">Type material studied</paragraph>
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<paragraph id="5EC06070A4CB999A58F997715B7BE771" pageId="71" pageNumber="72">
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(14 exs.). Holotype: male: "Type / male, female (symbols) Types / River near portuguese border, near Mwanza 9.II. (11?) 1948 / Brit. Mus. 1957-660 /
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<taxonomicName id="D16F586FF7CB7D6CB325338E19589C57" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
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O-C."(BMNH; habitus in Fig. 413). - Paratype: female: Pinned together with holotype but on separate label (1 ex. BMNH); "Paratype / Nyasaland Zomba Plateau Reservoir 7.11. 1948" (1 ex. AMGS); "Paratype / Nyasaland Reservoir Mwanza 9.11. 1948" (2exs. AMGS); "Paratype / Nyasaland stream longer Lilongwe rd. 20 miles from Dedza 30.9. 1948" (1 ex. AMGS); "Paratype / Nyasaland dam on lower Lilongwe rd. 29.9. 1948" (2exs. AMGS); "Paratype / Nyasaland Cisaiti R. nr. Dedza 28.9. 1948" (1 ex. AMGS); "S. Rhodesia Inganyi River 17.IX. 1948 /
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<taxonomicName id="3E0E027C4C965D0FE697A7C6705C63D0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
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sp. n. det. J. Omer-Cooper" (4 exs. AMGS); Nyasaland Dambo below Livingstonea Lake shore 21.9. 1945 / Paratype /
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<taxonomicName id="A6F79DE4C63EC78E1C82C40272D5AE17" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
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O.-C., O. Cooper det." (1 ex. IRSNB).
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</paragraph>
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</subSubSection>
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<subSubSection id="7A0AF08082C0F9B77880E992A772C1DB" pageId="71" pageNumber="72" type="materials_examined">
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<paragraph id="E2550B4B4E15F0D7D6B8CE322B95C063" pageId="71" pageNumber="72">Additional material studied</paragraph>
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<paragraph id="7843E78F68B8971666D454167F8115A9" pageId="71" pageNumber="72">
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(20 exx.). Tanzania: "Ruvu North Forest Reserve, waterholes, 3 km SE of Base
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<geoCoordinate id="50D31E6AC56C2DC4A7507EA8760B4019" direction="south" orientation="latitude" precision="15" value="-6.6222224">6°37'20" S</geoCoordinate>
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,
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<geoCoordinate id="15BFD828EE237975C08FCC05DDFB7D09" direction="east" orientation="longitude" precision="15" value="38.916668">38°55'00" E</geoCoordinate>
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alt. 250 ft 1.11. 1992 / Hynd Collection" (4 exs. BMNH, 1 ex. MZH); "Ruvu North Forest Reserve, Base Camp
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<geoCoordinate id="3B11B11EA66EF0EB10FB5071731A90ED" direction="south" orientation="latitude" precision="15" value="-6.627778">6°37'40" S</geoCoordinate>
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,
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<geoCoordinate id="C08C935781B71137D0A0A2427B603220" direction="east" orientation="longitude" precision="15" value="38.853886">38°51'14" E</geoCoordinate>
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alt. 200 ft 30.10. 1992 / Hynd Collection" (1 ex. MZH); "Zanzibar Pemba Sept. 1955 Fowler" (3 exs. AMGS); "Zanzibar Mangapwani Rd. Sept. 1955 JOC." (5 exs. AMGS). - Malawi: "Dam, Dedza on lower Lilongwe rd. 30.9. 1948" (2exs. AMGS); "Mtiti R. 1.10. 1948" (3exs. AMGS).
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</paragraph>
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</subSubSection>
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<subSubSection id="2C4878CCB1A66B1AD7AB6142E606C5B4" pageId="71" pageNumber="72" type="diagnosis">
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<paragraph id="A71DFFB57650DACCD5F8A052506D50C7" pageId="71" pageNumber="72">Diagnosis.</paragraph>
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<paragraph id="140F0E59A92C84B6679E22AF19F1BC02" pageId="71" pageNumber="72">
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<taxonomicName id="022F83C84A43D5E3B4B27EAB1B20D714" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
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is very closely related to
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<taxonomicName id="F5F92F106DBD2FCFF3F065ED3F0E5C37" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
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and externally very similar to this species. Small but distinct differences in shape of penis allow confident separation of the two species; penis short and apical process vague in
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<taxonomicName id="C9AC65095BC4BAFDC5A1300792FB0C2E" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus propinquus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="propinquus">Laccophilus propinquus</taxonomicName>
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(see also diagnosis of
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<taxonomicName id="E2F33D961427BD44B6387588A0D52E12" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus taeniolatus" order="Coleoptera" pageId="71" pageNumber="72" phylum="Arthropoda" rank="species" species="taeniolatus">Laccophilus taeniolatus</taxonomicName>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="113E474E8E285EF8D3592CDB21648FB9" pageId="71" pageNumber="72" type="description">
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<paragraph id="75A70D6E14931740C388A47A39672CD2" pageId="71" pageNumber="72">Description.</paragraph>
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<paragraph id="B069B2946D4FC4D4C3DF1338F8E07228" pageId="71" pageNumber="72">Body length 3.8-3.9 mm, width 1.9-2.0 mm. Pale ferrugineous; dorsal, dark ferrugineous colour pattern of body quite distinct (Fig. 413).</paragraph>
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<paragraph id="0B4F97482A7DA58582C80D9B3F86BEBB" pageId="71" pageNumber="72">Head: Pale ferrugineous, posteriorly with vague dark ferrugineous area. Submat, finely microsculptured. Reticulation almost of one kind, simple; double reticulation indistinct, weakly developed and difficult to distinguish. At eyes with irregularly distributed, sparse punctures.</paragraph>
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<paragraph id="A40F021800B53704F023833BAF7F26D3" pageId="71" pageNumber="72">Pronotum: Pale ferrugineous, anteriorly and at base with rather narrow, vague blackish ferrugineous to dark ferrugineous markings which are medially connected by a vague ferrugineous area. Submat, finely microsculptured; reticulation indistinctly double but size classes difficult to separate. Laterally and anteriorly with indistinct and sparse punctures.</paragraph>
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<paragraph id="D4A2ACFC03908CC843CECF6900BF06B5" pageId="71" pageNumber="72">Elytra: Pale ferrugineous, with rather distinct dark ferrugineous markings formed as irrorations (Fig. 413). Rather shiny, although finely microsculptured. Reticulation indistinctly double; clear size-classes difficult to discern. Discal, dorsolateral and lateral rows of fine and irregular punctures discernible but weakly developed.</paragraph>
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<paragraph id="B33CC67E5F1020DDFA43800617AAFAD6" pageId="71" pageNumber="72">Ventral aspect: Pale ferrugineous. Rather shiny and very finely (partly indistinctly) microsculptured. Abdomen with fine to very fine striae. Metacoxal plates with fine and shallow transverse furrows. Almost impunctate. Prosternal process slender, pointed. Apical ventrite almost symmetric, lacks lateral knob; apex more angle-shaped than in female (Fig. 57).</paragraph>
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<paragraph id="B6D4BB886ACCC4B57448C7955C1C827D" pageId="71" pageNumber="72">Legs: Pro- and mesotarsus long, slender, and with suckers.</paragraph>
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<paragraph id="CD405EF333FF96E026B9CF7D0DB4D1DB" pageId="71" pageNumber="72">Male genitalia: Penis In lateral aspect comparatively short, evenly curved and apical process not prominent (Fig. 254).</paragraph>
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<paragraph id="6FE243BBDB6422BD875D5F7B927D9896" pageId="71" pageNumber="72">Female: Externally as male but apex of apical ventrite broader and more rounded (Fig. 58).</paragraph>
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</subSubSection>
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<subSubSection id="83EF5EF763E50F2523D78F53FCEEC1C7" pageId="71" pageNumber="72" type="distribution">
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<paragraph id="1344467ACF2000FB8FA877A8462F7270" pageId="71" pageNumber="72">Distribution.</paragraph>
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<paragraph id="2AD6A129A4C096C416411713F9C6CB61" pageId="71" pageNumber="72">
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Malawi, Zimbabwe, Tanzania and Zanzibar (Fig. 540).
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<bibRefCitation id="6BAD9E0D41ACA21F5E34E0674F873394" author="Omer-Cooper, Joyce" journalOrPublisher="South African Animal Life" pageId="261" pageNumber="262" pagination="59 - 214" title="Chapter 2. Coleoptera: Dytiscidae." volume="11" year="1965">Omer-Cooper (1965)</bibRefCitation>
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adds Mozambique.
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</paragraph>
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</subSubSection>
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<subSubSection id="1B7B870E8C0404416A68A72BF0E02FFE" pageId="71" pageNumber="72" type="collecting circumstances">
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<paragraph id="C1C1C46AE8FBD25AA674DF49E67A0DCE" pageId="71" pageNumber="72">Collecting circumstances.</paragraph>
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<paragraph id="9E8EE5DD9179A6F4EB0164F26B2CE429" pageId="71" pageNumber="72">The species has been collected in streams e.g. with rocks, sand and some vegetation. It is also recorded from standing water in a swamp and a reservoir.</paragraph>
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</subSubSection>
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</treatment>
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</document> |