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<document id="B1C3DA01D5C1A4EEA0E728B0C2652664" ID-CLB-Dataset="46575" ID-DOI="10.11646/zootaxa.3755.5.5" ID-GBIF-Dataset="3ffb265e-c119-41d7-bacc-299437eccb43" ID-ISSN="1175-5326" ID-Zenodo-Dep="226483" ID-ZooBank="0BF2489A-8B2D-4414-B7C8-2F0B5C9CFAA6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460375235054" checkinUser="plazi" docAuthor="Kundrata, Robin, Kobieluszova, Lucie &amp; Bocak, Ladislav" docDate="2014" docId="5D1887CEFFCE1D6AAB8403470A32FD06" docLanguage="en" docName="zt03755p469.pdf" docOrigin="Zootaxa 3755 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Drilus rufipes" docType="treatment" docVersion="9" lastPageNumber="467" masterDocId="A121FFB6FFC71D60AB1300030948FFCA" masterDocTitle="A review of Drilini (Coleoptera: Elateridae: Agrypninae) of the Northern Levant, with description of a new species from Syria and a key to Levantine species" masterLastPageNumber="469" masterPageNumber="457" pageNumber="466" updateTime="1698296212971" updateUser="ExternalLinkService">
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<mods:title id="8BEF38164E60FA3A4F3F46D796B6889A">A review of Drilini (Coleoptera: Elateridae: Agrypninae) of the Northern Levant, with description of a new species from Syria and a key to Levantine species</mods:title>
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<mods:namePart id="04E898FD0313A7AEF749BC6EB685F2C3">Kundrata, Robin</mods:namePart>
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<mods:namePart id="1D1C6EE9D49AF041E4F9CE8C51BD5FA7">Kobieluszova, Lucie</mods:namePart>
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<mods:namePart id="A0D0911A2EECD913631664D73A9F3BB5">Bocak, Ladislav</mods:namePart>
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<mods:title id="8B106732040ED461B6A5EFC2D1117F64">Zootaxa</mods:title>
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<mods:date id="B591619B0EA8957E02C25214A14A81E7">2014</mods:date>
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<treatment id="5D1887CEFFCE1D6AAB8403470A32FD06" ID-DOI="http://doi.org/10.5281/zenodo.6133504" ID-GBIF-Taxon="119412929" ID-Zenodo-Dep="6133504" LSID="urn:lsid:plazi:treatment:5D1887CEFFCE1D6AAB8403470A32FD06" httpUri="http://treatment.plazi.org/id/5D1887CEFFCE1D6AAB8403470A32FD06" lastPageId="10" lastPageNumber="467" pageId="9" pageNumber="466">
<subSubSection id="9DAB6553FFCE1D69AB840347083CFCB5" pageId="9" pageNumber="466" type="nomenclature">
<paragraph id="D50E36D8FFCE1D69AB8403470B1EFC94" blockId="9.[151,598,836,895]" box="[151,598,836,862]" pageId="9" pageNumber="466">
<heading id="8E4681B4FFCE1D69AB8403470B1EFC94" bold="true" box="[151,598,836,862]" fontSize="11" level="1" pageId="9" pageNumber="466" reason="1">
<emphasis id="E7C5EACAFFCE1D69AB8403470B1EFC94" bold="true" box="[151,598,836,862]" pageId="9" pageNumber="466">
<taxonomicName id="12B14D5BFFCE1D69AB8403470877FC94" box="[151,319,836,862]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCE1D69AB8403470877FC94" bold="true" box="[151,319,836,862]" italics="true" pageId="9" pageNumber="466">Drilus rufipes</emphasis>
</taxonomicName>
(Baudi di
<bibRefCitation id="B1204B29FFCE1D69AAD103470B06FC94" author="Selve" box="[450,590,836,862]" pageId="9" pageNumber="466" refString="Baudi di Selve, F. (1871) Coleopterorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio: de Europaeis notis quibusdam additis. Pars quarta. Berliner Entomologische Zeitschrift, 15, 49 - 71. http: // dx. doi. org / 10.1002 / mmnd. 18710150113" type="journal article" year="1871">Selve, 1871</bibRefCitation>
)
</emphasis>
</heading>
</paragraph>
<paragraph id="D50E36D8FFCE1D69AB8D0365083CFCB5" blockId="9.[151,598,836,895]" box="[158,372,870,895]" pageId="9" pageNumber="466">
(
<figureCitation id="4D8A2A5DFFCE1D69ABB5036509A6FCB5" box="[166,238,870,895]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[170,1419,200,1853]" captionTargetId="figure@3.[156,1430,195,1858]" captionTargetPageId="3" captionText="FIGURES 1 – 6. General appearance. 1 — Drilus adustus; 2 — D. akbesianus; 3 — D. bicolor; 4 — D. nemethi sp. nov.; 5 — D. rectus; 6 — D. rufipes. Scale 2.0 mm." httpUri="https://zenodo.org/record/226484/files/figure.png" pageId="9" pageNumber="466">Figs 6</figureCitation>
,
<figureCitation id="4D8A2A5DFFCE1D69ABE90364080AFCB5" box="[250,322,871,895]" captionStart="FIGURES 7 – 18" captionStartId="4.[151,264,1576,1599]" captionTargetBox="[154,1423,199,1547]" captionTargetId="figure@4.[151,1436,193,1555]" captionTargetPageId="4" captionText="FIGURES 7 – 18. Antenna. 7 — Drilus adustus; 8 — D. akbesianus; 9 — D. bicolor; 10 — D. nemethi sp. nov.; 11 — D. rectus; 12 — D. rufipes. Pronotum. 13 — Drilus adustus; 14 — D. akbesianus; 15 — D. bicolor; 16 — D. nemethi sp. nov.; 17 — D. rectus; 18 — D. rufipes. Scale 1.0 mm (Figs 7 – 12), 0.5 mm (Figs 13 – 18)." httpUri="https://zenodo.org/record/226485/files/figure.png" pageId="9" pageNumber="466">12, 18</figureCitation>
,
<figureCitation id="4D8A2A5DFFCE1D69AA5D03640823FCB5" box="[334,363,871,895]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1605,1628]" captionTargetBox="[162,1421,342,1577]" captionTargetId="figure@5.[151,1436,337,1584]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Male genitalia. 19 — Drilus adustus; 20 — D. akbesianus; 21 — D. bicolor; 22 — D. nemethi sp. nov.; 23 — D. rectus; 24 — D. rufipes. Scale 0.25 mm." httpUri="https://zenodo.org/record/226486/files/figure.png" pageId="9" pageNumber="466">24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9DAB6553FFCE1D69AB8403AD0B9BFBC8" pageId="9" pageNumber="466" type="reference_group">
<paragraph id="D50E36D8FFCE1D69AB8403AD08BCFC29" blockId="9.[151,723,942,1026]" pageId="9" pageNumber="466">
<treatmentCitationGroup id="F5A111F6FFCE1D69AB8403AD08BCFC29" pageId="9" pageNumber="466">
<taxonomicName id="12B14D5BFFCE1D69AB8403AD08FAFC0E" authority="Baudi" authorityName="Baudi" box="[151,434,942,964]" class="Insecta" family="Drilidae" genus="Malacogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCE1D69AB8403AD0838FC0E" box="[151,368,942,964]" italics="true" pageId="9" pageNumber="466">Malacogaster rufipes</emphasis>
Baudi
</taxonomicName>
di
<treatmentCitation id="541010C9FFCE1D69AAC003AD0B3CFC0E" author="Selve" box="[467,628,942,964]" page="62" pageId="9" pageNumber="466" year="1871">
<bibRefCitation id="B1204B29FFCE1D69AAC003AD0B06FC0E" author="Selve" box="[467,590,942,964]" pageId="9" pageNumber="466" refString="Baudi di Selve, F. (1871) Coleopterorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio: de Europaeis notis quibusdam additis. Pars quarta. Berliner Entomologische Zeitschrift, 15, 49 - 71. http: // dx. doi. org / 10.1002 / mmnd. 18710150113" type="journal article" year="1871">Selve, 1871</bibRefCitation>
: 62
</treatmentCitation>
.
<taxonomicName id="12B14D5BFFCE1D69AB8403CE08B8FC29" authority="Zurcher 1911: 243" authorityName="Zurcher" authorityPageNumber="243" authorityYear="1911" box="[151,496,973,995]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCE1D69AB8403CE086CFC29" box="[151,292,973,995]" italics="true" pageId="9" pageNumber="466">Drilus rufipes</emphasis>
:
<treatmentCitation id="541010C9FFCE1D69AA2103CE08B8FC29" author="Zurcher" box="[306,496,973,995]" page="243" pageId="9" pageNumber="466" year="1911">
<bibRefCitation id="B1204B29FFCE1D69AA2103CE08F5FC29" author="Zurcher" box="[306,445,973,995]" pageId="9" pageNumber="466" refString="Zurcher, C. (1911) Note sur deux Drilus de l`ile de Chypre (Col. Drilini). Bulletin de la Societe Entomologique de France, 1911, 243 - 244." type="journal article" year="1911">Zürcher 1911</bibRefCitation>
: 243
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D50E36D8FFCE1D69AB8403EF0B9BFBC8" blockId="9.[151,723,942,1026]" box="[151,723,1004,1026]" pageId="9" pageNumber="466">
<treatmentCitationGroup id="F5A111F6FFCE1D69AB8403EF0B9BFBC8" box="[151,723,1004,1026]" pageId="9" pageNumber="466">
<taxonomicName id="12B14D5BFFCE1D69AB8403EF0B4FFBC8" authority="Bourgeois, 1908: 241" authorityName="Bourgeois" authorityPageNumber="241" authorityYear="1908" box="[151,519,1004,1026]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCE1D69AB8403EF086BFBC8" box="[151,291,1004,1026]" italics="true" pageId="9" pageNumber="466">Drilus reitteri</emphasis>
<treatmentCitation id="541010C9FFCE1D69AA3903EF0B4FFBC8" author="Bourgeois" box="[298,519,1004,1026]" page="241" pageId="9" pageNumber="466" year="1908">
<bibRefCitation id="B1204B29FFCE1D69AA3903EF089CFBC8" author="Bourgeois" box="[298,468,1004,1026]" pageId="9" pageNumber="466" refString="Bourgeois, J. (1908) Description d`une nouvelle espece de Drilus [Col.]. Bulletin de la Societe Entomologique de France, 1908, 241 - 242." type="journal article" year="1908">Bourgeois, 1908</bibRefCitation>
: 241
</treatmentCitation>
</taxonomicName>
;
<treatmentCitation id="541010C9FFCE1D69A90203EF0B87FBC8" author="Zurcher" box="[529,719,1004,1026]" page="243" pageId="9" pageNumber="466" year="1911">
<bibRefCitation id="B1204B29FFCE1D69A90203EF0BD5FBC8" author="Zurcher" box="[529,669,1004,1026]" pageId="9" pageNumber="466" refString="Zurcher, C. (1911) Note sur deux Drilus de l`ile de Chypre (Col. Drilini). Bulletin de la Societe Entomologique de France, 1911, 243 - 244." type="journal article" year="1911">Zürcher 1911</bibRefCitation>
: 243
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9DAB6553FFCE1D69AB84042D0812FBA1" pageId="9" pageNumber="466" type="materials_examined">
<paragraph id="D50E36D8FFCE1D69AB84042D0812FBA1" blockId="9.[151,1437,1070,2031]" pageId="9" pageNumber="466">
<emphasis id="E7C5EACAFFCE1D69AB84042D08CAFB8D" bold="true" box="[151,386,1070,1095]" pageId="9" pageNumber="466">Material examined.</emphasis>
<specimenCount id="C3B7FD51FFCE1D69AA99042C0892FB8D" box="[394,474,1070,1095]" pageId="9" pageNumber="466" type="male">1 male</specimenCount>
,
<collectingCountry id="ADA67648FFCE1D69AAF4042D0B6DFB8D" box="[487,549,1070,1095]" name="Syria" pageId="9" pageNumber="466">Syria</collectingCountry>
, / Haifa //
<collectingCountry id="ADA67648FFCE1D69A9B2042D0B94FB8D" box="[673,732,1070,1095]" name="Syria" pageId="9" pageNumber="466">Syrie</collectingCountry>
//
<typeStatus id="0A0A887AFFCE1D69A9EE042D0A3AFB8D" box="[765,882,1070,1095]" pageId="9" pageNumber="466" type="holotype">Holotypus</typeStatus>
1885 /
<taxonomicName id="12B14D5BFFCE1D69A8DF042D0D66FB8D" box="[972,1070,1070,1095]" class="Insecta" family="Rhagophthalmidae" genus="Cydistus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="466" phylum="Arthropoda" rank="genus">Cydistus</taxonomicName>
♂ /
<taxonomicName id="12B14D5BFFCE1D69AF77042D0DF0FB8D" box="[1124,1208,1070,1095]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">Reitteri</taxonomicName>
/ Bourgeois // Coll. Reitter (
<collectionCode id="B3A0AE1DFFCE1D69ABE104500806FBA1" LSID="urn:lsid:biocol.org:col:34991" box="[242,334,1107,1131]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="9" pageNumber="466">HNHM</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="9DAB6553FFCE1D69ABD404750BA4FB79" pageId="9" pageNumber="466" type="diagnosis">
<paragraph id="D50E36D8FFCE1D69ABD404750BA4FB79" blockId="9.[151,1437,1070,2031]" pageId="9" pageNumber="466">
<emphasis id="E7C5EACAFFCE1D69ABD404750809FB45" bold="true" box="[199,321,1142,1167]" pageId="9" pageNumber="466">Diagnosis.</emphasis>
<taxonomicName id="12B14D5BFFCE1D69AA59047508A1FB45" box="[330,489,1142,1167]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="466" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCE1D69AA59047508A1FB45" box="[330,489,1142,1167]" italics="true" pageId="9" pageNumber="466">Drilus rufipes</emphasis>
</taxonomicName>
is easily recognizable by its minute body size (3.0 mm), pronotum dark brown to black (
<figureCitation id="4D8A2A5DFFCE1D69ABF00499086FFB79" box="[227,295,1178,1203]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[170,1419,200,1853]" captionTargetId="figure@3.[156,1430,195,1858]" captionTargetPageId="3" captionText="FIGURES 1 – 6. General appearance. 1 — Drilus adustus; 2 — D. akbesianus; 3 — D. bicolor; 4 — D. nemethi sp. nov.; 5 — D. rectus; 6 — D. rufipes. Scale 2.0 mm." httpUri="https://zenodo.org/record/226484/files/figure.png" pageId="9" pageNumber="466">Fig. 6</figureCitation>
), and slightly serrate antennae (
<figureCitation id="4D8A2A5DFFCE1D69A99804990B97FB79" box="[651,735,1178,1203]" captionStart="FIGURES 7 – 18" captionStartId="4.[151,264,1576,1599]" captionTargetBox="[154,1423,199,1547]" captionTargetId="figure@4.[151,1436,193,1555]" captionTargetPageId="4" captionText="FIGURES 7 – 18. Antenna. 7 — Drilus adustus; 8 — D. akbesianus; 9 — D. bicolor; 10 — D. nemethi sp. nov.; 11 — D. rectus; 12 — D. rufipes. Pronotum. 13 — Drilus adustus; 14 — D. akbesianus; 15 — D. bicolor; 16 — D. nemethi sp. nov.; 17 — D. rectus; 18 — D. rufipes. Scale 1.0 mm (Figs 7 – 12), 0.5 mm (Figs 13 – 18)." httpUri="https://zenodo.org/record/226485/files/figure.png" pageId="9" pageNumber="466">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9DAB6553FFCE1D6AABD404BD0CD4FF65" lastPageId="10" lastPageNumber="467" pageId="9" pageNumber="466" type="description">
<paragraph id="D50E36D8FFCE1D69ABD404BD0D1BFAD5" blockId="9.[151,1437,1070,2031]" pageId="9" pageNumber="466">
<emphasis id="E7C5EACAFFCE1D69ABD404BD083CFB1D" bold="true" box="[199,372,1214,1239]" pageId="9" pageNumber="466">Redescription.</emphasis>
Male. Body small, elongate, 3.5 times longer than width at humeri, moderately convex dorsally (
<figureCitation id="4D8A2A5DFFCE1D69AA1704E10803FB31" box="[260,331,1250,1275]" captionStart="FIGURES 1 – 6" captionStartId="3.[151,264,1879,1902]" captionTargetBox="[170,1419,200,1853]" captionTargetId="figure@3.[156,1430,195,1858]" captionTargetPageId="3" captionText="FIGURES 1 – 6. General appearance. 1 — Drilus adustus; 2 — D. akbesianus; 3 — D. bicolor; 4 — D. nemethi sp. nov.; 5 — D. rectus; 6 — D. rufipes. Scale 2.0 mm." httpUri="https://zenodo.org/record/226484/files/figure.png" pageId="9" pageNumber="466">Fig. 6</figureCitation>
). All parts of body dark brown to black, legs (except tarsi) and apical two abdominal ventrites yellowish to light brown; entire body covered with light brown to brown pubescence.
</paragraph>
<paragraph id="D50E36D8FFCE1D69ABD405290CD4F9F5" blockId="9.[151,1437,1070,2031]" pageId="9" pageNumber="466">
Head slightly hypognathous, including eyes as wide as anterior margin of pronotum. Surface of cranium basally coarsely wrinkled; moderately deeply punctured, with wide, shallow, glabrous depression between antennal sockets; semierect pubescence sparse basally and laterally, almost absent dorsally. Eyes medium-sized, hemispherically prominent, their frontal distance 2.3 times eye diameter. Mandibles moderately long, considerably curved, incisor margin with tooth in middle part, shiny. Maxillary palpi with apical palpomere narrow, with slightly compressed, obliquely cut apex; labial palpi tiny, apical palpomere pointed. Antennae 11-segmented, slightly serrate, robust, reaching third of elytral length, scapus long, robust, pedicel short, small, third antennomere 2.8 times longer than antennomere 2, antennomeres 3–10 slightly serrate, apical antennomere simple, longest (
<figureCitation id="4D8A2A5DFFCE1D69AE2F06250CC7F9F5" box="[1340,1423,1574,1599]" captionStart="FIGURES 7 – 18" captionStartId="4.[151,264,1576,1599]" captionTargetBox="[154,1423,199,1547]" captionTargetId="figure@4.[151,1436,193,1555]" captionTargetPageId="4" captionText="FIGURES 7 – 18. Antenna. 7 — Drilus adustus; 8 — D. akbesianus; 9 — D. bicolor; 10 — D. nemethi sp. nov.; 11 — D. rectus; 12 — D. rufipes. Pronotum. 13 — Drilus adustus; 14 — D. akbesianus; 15 — D. bicolor; 16 — D. nemethi sp. nov.; 17 — D. rectus; 18 — D. rufipes. Scale 1.0 mm (Figs 7 – 12), 0.5 mm (Figs 13 – 18)." httpUri="https://zenodo.org/record/226485/files/figure.png" pageId="9" pageNumber="466">Fig. 12</figureCitation>
).
</paragraph>
<paragraph id="D50E36D8FFCE1D69ABD406480C76F86D" blockId="9.[151,1437,1070,2031]" pageId="9" pageNumber="466">
Pronotum flat, widest in middle, 1.5 times wider posteriorly than length at midline. Anterior margin almost straight, lateral margins convex, posterior margin sinuate. Anterior angles obtuse, posterior angles inconspicuous, obtuse (
<figureCitation id="4D8A2A5DFFCE1D69ABE006910802F961" box="[243,330,1682,1707]" captionStart="FIGURES 7 – 18" captionStartId="4.[151,264,1576,1599]" captionTargetBox="[154,1423,199,1547]" captionTargetId="figure@4.[151,1436,193,1555]" captionTargetPageId="4" captionText="FIGURES 7 – 18. Antenna. 7 — Drilus adustus; 8 — D. akbesianus; 9 — D. bicolor; 10 — D. nemethi sp. nov.; 11 — D. rectus; 12 — D. rufipes. Pronotum. 13 — Drilus adustus; 14 — D. akbesianus; 15 — D. bicolor; 16 — D. nemethi sp. nov.; 17 — D. rectus; 18 — D. rufipes. Scale 1.0 mm (Figs 7 – 12), 0.5 mm (Figs 13 – 18)." httpUri="https://zenodo.org/record/226485/files/figure.png" pageId="9" pageNumber="466">Fig. 18</figureCitation>
); surface of disc coarsely wrinkled, with sparse, semierect, long setae. Scutellum flat, triangleshaped, with apex rounded. Prosternum transverse, prosternal process short, slightly elevated medially. Mesoventrite narrow, v-shaped, with depression in middle part of frontal margin, hind margin rounded. Metaventrite large, trapezoidal, sparsely punctured. Elytra subparallel-sided, widest at humeri, 2.5 times longer than width at humeri, tapered apically, wrinkled, covered with semierect pubescence. Abdomen short, slender, ventrites covered with sparse, moderately deep punctures and sparse, long hairs. Legs slender, slightly compressed, with sparse, long, semierect setae, coxae long, robust, trochanters slender, obliquely attached to femora, tarsomeres 1–3 subequal in length, tarsomere 4 shortest, apical tarsomere long, claws simple, slender, slightly curved.
</paragraph>
<paragraph id="D50E36D8FFCE1D69ABD407B10DD9F825" blockId="9.[151,1437,1070,2031]" pageId="9" pageNumber="466">
Male genitalia with phallus longer than paramerae, robust, considerably curved, parallel-sided in dorsoventral view; paramerae short, sparsely setose; phallobase robust, longer than paramerae (
<figureCitation id="4D8A2A5DFFCE1D69AF2307D50DCCF825" box="[1072,1156,2006,2031]" captionStart="FIGURES 19 – 24" captionStartId="5.[151,264,1605,1628]" captionTargetBox="[162,1421,342,1577]" captionTargetId="figure@5.[151,1436,337,1584]" captionTargetPageId="5" captionText="FIGURES 19 – 24. Male genitalia. 19 — Drilus adustus; 20 — D. akbesianus; 21 — D. bicolor; 22 — D. nemethi sp. nov.; 23 — D. rectus; 24 — D. rufipes. Scale 0.25 mm." httpUri="https://zenodo.org/record/226486/files/figure.png" pageId="9" pageNumber="466">Fig. 24</figureCitation>
).
</paragraph>
<paragraph id="D50E36D8FFCD1D6AABD4009B0CD4FF65" blockId="10.[151,1436,151,717]" box="[199,1436,151,176]" pageId="10" pageNumber="467">
Measurements. BL 3.0 mm, EL
<quantity id="12499B3DFFCD1D6AA93C009B0BCFFF65" box="[559,647,152,176]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="10" pageNumber="467" unit="mm" value="2.2">2.2 mm</quantity>
, WH
<quantity id="12499B3DFFCD1D6AA9DB009B0A68FF65" box="[712,800,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="10" pageNumber="467" unit="mm" value="0.9">0.9 mm</quantity>
, PL
<quantity id="12499B3DFFCD1D6AA84700940AE3FF65" box="[852,939,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="10" pageNumber="467" unit="mm" value="0.6">0.6 mm</quantity>
, PW
<quantity id="12499B3DFFCD1D6AA8FB009B0D08FF65" box="[1000,1088,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="10" pageNumber="467" unit="mm" value="0.8">0.8 mm</quantity>
, Edist
<quantity id="12499B3DFFCD1D6AAF9F00940DACFF65" box="[1164,1252,151,176]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="467" unit="mm" value="0.5">0.5 mm</quantity>
, Ediam
<quantity id="12499B3DFFCD1D6AAE2C009B0CD0FF65" box="[1343,1432,152,176]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="467" unit="mm" value="0.2">0.2 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="9DAB6553FFCD1D6AABD400BF0B44FF1F" box="[199,524,188,213]" pageId="10" pageNumber="467" type="distribution">
<paragraph id="D50E36D8FFCD1D6AABD400BF0B44FF1F" blockId="10.[151,1436,151,717]" box="[199,524,188,213]" pageId="10" pageNumber="467">
<emphasis id="E7C5EACAFFCD1D6AABD400BF0828FF1F" bold="true" box="[199,352,188,213]" pageId="10" pageNumber="467">Distribution.</emphasis>
<collectingCountry id="ADA67648FFCD1D6AAA7400BF08F3FF1E" box="[359,443,188,212]" name="Cyprus" pageId="10" pageNumber="467">Cyprus</collectingCountry>
,
<collectingCountry id="ADA67648FFCD1D6AAAD500BF0B40FF1F" box="[454,520,188,213]" name="Israel" pageId="10" pageNumber="467">Israel</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9DAB6553FFCD1D6AABD400DC0A32FD06" pageId="10" pageNumber="467" type="discussion">
<paragraph id="D50E36D8FFCD1D6AABD400DC089CFE66" blockId="10.[151,1436,151,717]" pageId="10" pageNumber="467">
<emphasis id="E7C5EACAFFCD1D6AABD400DC0873FF32" bold="true" box="[199,315,223,248]" pageId="10" pageNumber="467">Remarks.</emphasis>
<taxonomicName id="12B14D5BFFCD1D6AAA5500E308AEFF32" box="[326,486,224,248]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCD1D6AAA5500E308AEFF32" box="[326,486,224,248]" italics="true" pageId="10" pageNumber="467">Drilus rufipes</emphasis>
</taxonomicName>
belongs to the fauna of
<collectingCountry id="ADA67648FFCD1D6AA81E00E30A16FF32" box="[781,862,224,248]" name="Cyprus" pageId="10" pageNumber="467">Cyprus</collectingCountry>
and northern
<collectingCountry id="ADA67648FFCD1D6AAF1500DC0D0FFF32" box="[1030,1095,223,248]" name="Israel" pageId="10" pageNumber="467">Israel</collectingCountry>
, however, we include it here because its occurrence in the Northern Levant is highly probable. The border between the Northern and Southern Levant (i.e., between
<collectingCountry id="ADA67648FFCD1D6AAA980124088EFE8A" box="[395,454,295,320]" name="Syria" pageId="10" pageNumber="467">Syria</collectingCountry>
and
<collectingCountry id="ADA67648FFCD1D6AAAED01240B76FE8A" box="[510,574,295,320]" name="Israel" pageId="10" pageNumber="467">Israel</collectingCountry>
) is artificial and there is no barrier for animal migration (
<bibRefCitation id="B1204B29FFCD1D6AAFD6012B0C0EFE8A" author="Genz" box="[1221,1350,296,320]" pageId="10" pageNumber="467" refString="Genz, H. (2012) The Northern Levant. In: Potts, D. T. (Ed.), A Companion to the Archaeology of the Ancient Near East. Volume 1. Blackwell Publishing Ltd., Chichester, UK, pp. 607 - 628." type="book" year="2012">Genz 2012</bibRefCitation>
). Baudi di
<bibRefCitation id="B1204B29FFCD1D6AABA7014F080DFEAE" author="Selve" box="[180,325,332,357]" pageId="10" pageNumber="467" refString="Baudi di Selve, F. (1871) Coleopterorum messis in insula Cypro et Asia minore ab Eugenio Truqui congregatae recensitio: de Europaeis notis quibusdam additis. Pars quarta. Berliner Entomologische Zeitschrift, 15, 49 - 71. http: // dx. doi. org / 10.1002 / mmnd. 18710150113" type="journal article" year="1871">Selve (1871)</bibRefCitation>
described
<taxonomicName id="12B14D5BFFCD1D6AAAD3014F0BFAFEAE" box="[448,690,332,356]" class="Insecta" family="Drilidae" genus="Malacogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCD1D6AAAD3014F0BFAFEAE" box="[448,690,332,356]" italics="true" pageId="10" pageNumber="467">Malacogaster rufipes</emphasis>
</taxonomicName>
from
<collectingCountry id="ADA67648FFCD1D6AA9EB014F0A01FEAE" box="[760,841,332,356]" name="Cyprus" pageId="10" pageNumber="467">Cyprus</collectingCountry>
and
<bibRefCitation id="B1204B29FFCD1D6AA892014F0D0EFEAE" box="[897,1094,332,357]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1908) Description d`une nouvelle espece de Drilus [Col.]. Bulletin de la Societe Entomologique de France, 1908, 241 - 242." type="journal article">Bourgeois (1908)</bibRefCitation>
described
<taxonomicName id="12B14D5BFFCD1D6AAFD2014F0C16FEAE" box="[1217,1374,332,356]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AAFD2014F0C16FEAE" box="[1217,1374,332,356]" italics="true" pageId="10" pageNumber="467">Drilus reitteri</emphasis>
</taxonomicName>
from mainland Levant (
<typeStatus id="0A0A887AFFCD1D6AAA77017308DEFE42" box="[356,406,368,392]" pageId="10" pageNumber="467">type</typeStatus>
locality: &quot;
<collectingCountry id="ADA67648FFCD1D6AA918016C0B06FE42" box="[523,590,367,392]" name="Syria" pageId="10" pageNumber="467">Syrie</collectingCountry>
&quot;).
<bibRefCitation id="B1204B29FFCD1D6AA978016C0A5DFE42" author="Zurcher" box="[619,789,367,392]" pageId="10" pageNumber="467" refString="Zurcher, C. (1911) Note sur deux Drilus de l`ile de Chypre (Col. Drilini). Bulletin de la Societe Entomologique de France, 1911, 243 - 244." type="journal article" year="1911">Zürcher (1911)</bibRefCitation>
transferred
<taxonomicName id="12B14D5BFFCD1D6AA88D01720D5AFE42" box="[926,1042,368,392]" class="Insecta" family="Drilidae" genus="Malacogaster" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCD1D6AA88D01720AFDFE42" box="[926,949,369,392]" italics="true" pageId="10" pageNumber="467">M</emphasis>
.
<emphasis id="E7C5EACAFFCD1D6AA8D701730D5AFE42" box="[964,1042,368,392]" italics="true" pageId="10" pageNumber="467">rufipes</emphasis>
</taxonomicName>
to
<taxonomicName id="12B14D5BFFCD1D6AAF2B01730D37FE42" box="[1080,1151,368,392]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="E7C5EACAFFCD1D6AAF2B01730D37FE42" box="[1080,1151,368,392]" italics="true" pageId="10" pageNumber="467">Drilus</emphasis>
</taxonomicName>
and synonymized
<taxonomicName id="12B14D5BFFCD1D6AAE47017309ACFE66" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AAE47017309ACFE66" italics="true" pageId="10" pageNumber="467">Drilus reitteri</emphasis>
</taxonomicName>
under
<taxonomicName id="12B14D5BFFCD1D6AAA2201970886FE66" box="[305,462,404,428]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="rufipes">
<emphasis id="E7C5EACAFFCD1D6AAA2201970886FE66" box="[305,462,404,428]" italics="true" pageId="10" pageNumber="467">Drilus rufipes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D50E36D8FFCD1D6AABD401B40A32FD06" blockId="10.[151,1436,151,717]" pageId="10" pageNumber="467">
The
<typeStatus id="0A0A887AFFCD1D6AAA1301BB0878FE1A" box="[256,304,440,464]" pageId="10" pageNumber="467">type</typeStatus>
series of
<taxonomicName id="12B14D5BFFCD1D6AAABC01BB0A54FE1A" authority="Bourgeois, 1908" authorityName="Bourgeois" authorityYear="1908" box="[431,796,439,464]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AAABC01BB0B19FE05" box="[431,593,440,464]" italics="true" pageId="10" pageNumber="467">Drilus reitteri</emphasis>
<bibRefCitation id="B1204B29FFCD1D6AA94E01B40A54FE1A" author="Bourgeois" box="[605,796,439,464]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1908) Description d`une nouvelle espece de Drilus [Col.]. Bulletin de la Societe Entomologique de France, 1908, 241 - 242." type="journal article" year="1908">Bourgeois, 1908</bibRefCitation>
</taxonomicName>
should contain two
<typeStatus id="0A0A887AFFCD1D6AAF0B01BB0D36FE1A" box="[1048,1150,440,464]" pageId="10" pageNumber="467" type="syntype">syntypes</typeStatus>
; one
<typeStatus id="0A0A887AFFCD1D6AAFD601BB0C54FE1A" box="[1221,1308,440,464]" pageId="10" pageNumber="467" type="syntype">syntype</typeStatus>
should be deposited in the collection of Bourgeois in MHNP (no
<typeStatus id="0A0A887AFFCD1D6AA83301DE0A3FFE3F" box="[800,887,477,501]" pageId="10" pageNumber="467" type="syntype">syntype</typeStatus>
found) and the second
<typeStatus id="0A0A887AFFCD1D6AAF8301DE0DAFFE3F" box="[1168,1255,477,501]" pageId="10" pageNumber="467" type="syntype">syntype</typeStatus>
in the Reitter's collection in HNHM (
<bibRefCitation id="B1204B29FFCD1D6AAA9D01FC0B0DFDD2" author="Bourgeois" box="[398,581,511,536]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1908) Description d`une nouvelle espece de Drilus [Col.]. Bulletin de la Societe Entomologique de France, 1908, 241 - 242." type="journal article" year="1908">Bourgeois 1908</bibRefCitation>
). We found a specimen of
<taxonomicName id="12B14D5BFFCD1D6AA86302030AFFFDD2" box="[880,951,512,536]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="E7C5EACAFFCD1D6AA86302030AFFFDD2" box="[880,951,512,536]" italics="true" pageId="10" pageNumber="467">Drilus</emphasis>
</taxonomicName>
in the Reitter's collection in HNHM which bears the
<typeStatus id="0A0A887AFFCD1D6AAA1902260872FDF7" box="[266,314,549,573]" pageId="10" pageNumber="467">type</typeStatus>
label &quot;
<typeStatus id="0A0A887AFFCD1D6AAA8302270B42FDF7" box="[400,522,548,573]" pageId="10" pageNumber="467" type="holotype">Holotypus</typeStatus>
,
<taxonomicName id="12B14D5BFFCD1D6AA90A02270AD4FDF7" authority="Bourgeois, 1885" authorityName="Bourgeois" authorityYear="1885" box="[537,924,548,573]" class="Insecta" family="Rhagophthalmidae" genus="Cydistus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AA90A02270B9AFDF6" box="[537,722,548,572]" italics="true" pageId="10" pageNumber="467">Cydistus reitteri</emphasis>
<bibRefCitation id="B1204B29FFCD1D6AA9CF02270AD4FDF7" author="Bourgeois" box="[732,924,548,573]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1885) Cydistus, nov. gen. Malacodermatorum. Bulletin des Seances et Bulletin Bibliographique de la Societe Entomologique de France, 1885, 37 - 38." type="journal article" year="1885">Bourgeois, 1885</bibRefCitation>
</taxonomicName>
, ♂&quot;. The label was apparently written by a curator after the collection of Edmund Reitter was deposited in the museum.
<taxonomicName id="12B14D5BFFCD1D6AAF12024B0C6BFDAA" authority="Bourgeois, 1885" authorityName="Bourgeois" authorityYear="1885" box="[1025,1315,583,608]" class="Insecta" family="Rhagophthalmidae" genus="Cydistus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="E7C5EACAFFCD1D6AAF12024B0D2AFDAA" box="[1025,1122,584,608]" italics="true" pageId="10" pageNumber="467">Cydistus</emphasis>
<bibRefCitation id="B1204B29FFCD1D6AAF7902440C6BFDAA" author="Bourgeois" box="[1130,1315,583,608]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1885) Cydistus, nov. gen. Malacodermatorum. Bulletin des Seances et Bulletin Bibliographique de la Societe Entomologique de France, 1885, 37 - 38." type="journal article" year="1885">Bourgeois, 1885</bibRefCitation>
</taxonomicName>
is a genus of uncertain position in Elateriformia (
<bibRefCitation id="B1204B29FFCD1D6AA947026F0A04FD4E" author="Lawrence" box="[596,844,620,644]" pageId="10" pageNumber="467" refString="Lawrence, J. F., Kawashima, I. &amp; Branham, M. A. (2010) Elateriformia Incertae Sedis. In: Leschen, R. A. B., Beutel, R. G. &amp; Lawrence, J. F. (Eds.), Coleoptera, Beetles. Vol. 2. Morphology and Systematics (Elateroidea, Bostrichiformia, Cucujiformia partim). In: Kristensen, N. P. &amp; Beutel, R. G. (Eds), Handbook of Zoology, Arthropoda: Insecta. Berlin / New York, Walter de Gruyter GmbH &amp; Co. KG, pp. 162 - 177." type="book" year="2010">
Lawrence
<emphasis id="E7C5EACAFFCD1D6AA9DD026E0A49FD4E" box="[718,769,620,644]" italics="true" pageId="10" pageNumber="467">et al</emphasis>
. 2010
</bibRefCitation>
) with
<taxonomicName id="12B14D5BFFCD1D6AA88A026F0C5DFD4F" authority="Bourgeois, 1885" authorityName="Bourgeois" authorityYear="1885" box="[921,1301,620,645]" class="Insecta" family="Rhagophthalmidae" genus="Cydistus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AA88A026F0D18FD4E" box="[921,1104,620,644]" italics="true" pageId="10" pageNumber="467">Cydistus reitteri</emphasis>
<bibRefCitation id="B1204B29FFCD1D6AAF49026F0C5DFD4F" author="Bourgeois" box="[1114,1301,620,645]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1885) Cydistus, nov. gen. Malacodermatorum. Bulletin des Seances et Bulletin Bibliographique de la Societe Entomologique de France, 1885, 37 - 38." type="journal article" year="1885">Bourgeois, 1885</bibRefCitation>
</taxonomicName>
as the
<typeStatus id="0A0A887AFFCD1D6AAE7F026E0CD4FD4F" box="[1388,1436,621,645]" pageId="10" pageNumber="467">type</typeStatus>
species. The
<typeStatus id="0A0A887AFFCD1D6AAA3F02930814FD62" box="[300,348,656,680]" pageId="10" pageNumber="467">type</typeStatus>
specimen bearing the subsequently attached label
<taxonomicName id="12B14D5BFFCD1D6AA8B202930C52FD62" authority="Bourgeois, 1885" authorityName="Bourgeois" authorityYear="1885" box="[929,1306,655,680]" class="Insecta" family="Rhagophthalmidae" genus="Cydistus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AA8B202930D1FFD6D" box="[929,1111,656,680]" italics="true" pageId="10" pageNumber="467">Cydistus reitteri</emphasis>
<bibRefCitation id="B1204B29FFCD1D6AAF4C028C0C52FD62" author="Bourgeois" box="[1119,1306,655,680]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1885) Cydistus, nov. gen. Malacodermatorum. Bulletin des Seances et Bulletin Bibliographique de la Societe Entomologique de France, 1885, 37 - 38." type="journal article" year="1885">Bourgeois, 1885</bibRefCitation>
</taxonomicName>
is a
<taxonomicName id="12B14D5BFFCD1D6AAE4702930CD4FD62" box="[1364,1436,656,680]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="genus">
<emphasis id="E7C5EACAFFCD1D6AAE4702930CD4FD62" box="[1364,1436,656,680]" italics="true" pageId="10" pageNumber="467">Drilus</emphasis>
</taxonomicName>
and we consider it as a
<typeStatus id="0A0A887AFFCD1D6AAA8F02B608BBFD07" box="[412,499,693,717]" pageId="10" pageNumber="467" type="syntype">syntype</typeStatus>
of
<taxonomicName id="12B14D5BFFCD1D6AA90B02B70A3EFD06" authority="Bourgeois, 1908" authorityName="Bourgeois" authorityYear="1908" box="[536,886,692,717]" class="Insecta" family="Drilidae" genus="Drilus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="467" phylum="Arthropoda" rank="species" species="reitteri">
<emphasis id="E7C5EACAFFCD1D6AA90B02B70BFCFD06" box="[536,692,692,716]" italics="true" pageId="10" pageNumber="467">Drilus reitteri</emphasis>
<bibRefCitation id="B1204B29FFCD1D6AA9A802B70A3EFD06" author="Bourgeois" box="[699,886,692,717]" pageId="10" pageNumber="467" refString="Bourgeois, J. (1908) Description d`une nouvelle espece de Drilus [Col.]. Bulletin de la Societe Entomologique de France, 1908, 241 - 242." type="journal article" year="1908">Bourgeois, 1908</bibRefCitation>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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