254 lines
32 KiB
XML
254 lines
32 KiB
XML
<document id="4950A94BAEAA931714384C641B926B07" ID-CLB-Dataset="182450" ID-DOI="10.11646/zootaxa.5165.3.8" ID-GBIF-Dataset="762b9b99-738b-48e5-a7b6-cba72e455c71" ID-ISSN="1175-5326" ID-Zenodo-Dep="6838529" ID-ZooBank="63D31FC2-0954-40A5-8537-3D9AAA77D17A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1657877487759" checkinUser="plazi" docAuthor="Alekseev, Vitalii I. & Bukejs, Andris" docDate="2022" docId="5D0DC56DFFD4FF8CFF01F987613DFBD9" docLanguage="en" docName="zootaxa.5165.3.8.pdf" docOrigin="Zootaxa 5165 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Eustrophus praecursor Alekseev & Bukejs 2022, sp. nov." docType="treatment" docUuid="568D8333-4E82-4A41-9AF9-AD23B2F45E8B" docUuidSource="ZooBank" docVersion="9" lastPageNumber="440" masterDocId="A134BD15FFD5FF89FF96FFDC6371FFA4" masterDocTitle="Eustrophus praecursor sp. nov.: the first Eocene species of Eustrophinae (Coleoptera: Tetratomidae) from Baltic amber" masterLastPageNumber="442" masterPageNumber="435" pageNumber="436" updateTime="1699359133968" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="FC556CD468BAB77E33F655C498FAA84B" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="E0460EBDDB11B0290CFD4728BC8B5070">
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<mods:title id="E114DEEBE0584E1F0122B5F77BA54160">Eustrophus praecursor sp. nov.: the first Eocene species of Eustrophinae (Coleoptera: Tetratomidae) from Baltic amber</mods:title>
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<mods:name id="6FEBA0B82094724733EF24CEFC4CDA48" type="personal">
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<mods:role id="0F3D3993EC982D7BD024C7D63D963862">
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<mods:roleTerm id="333F35238961117E24CA6649EF38794A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F5A563A27CE5DA0B02B90D6C50EE32B8">Alekseev, Vitalii I.</mods:namePart>
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<mods:affiliation id="71A110304E3290593DB9D66C96ACBE9E">Kaliningrad Regional Amber Museum, Marshal Vasilevskii square 1, Kaliningrad 236016, Russia. & Immanuel Kant Baltic Federal University, Nevskogo street 14, Kaliningrad 236016, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="7B606E4CD2DD6EC8B69EF5E03ACD11FD" type="personal">
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<mods:roleTerm id="AD6BDD880E66150031495912BE63D1D9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="993B63ED8D666B83CED67DBCD497FC5D">Bukejs, Andris</mods:namePart>
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<mods:nameIdentifier id="58EF0A40310F5C1EDE7A56482F89D111" type="ORCID">0000-0001-7165-3023</mods:nameIdentifier>
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<mods:affiliation id="97F9AF403568E945382196761EDEF2CF">Institute of Life Sciences and Technologies, Daugavpils University, Vienîbas 13, Daugavpils LV- 5401, Latvia. carabidae @ inbox. lv; https: // orcid. org / 0000 - 0001 - 7165 - 3023</mods:affiliation>
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<mods:nameIdentifier id="0329BB5DF1AB566A62BD82FAD5D3BACA" type="email">carabidae@inbox.lv</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="98E8AFCF222E4C4CC0F3C9C155C13B65">text</mods:typeOfResource>
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<mods:title id="28212A616212B30DE0A400D0F1283923">Zootaxa</mods:title>
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<mods:part id="5B801479A6B5C1338D9258BA06E03370">
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<mods:date id="AFFB00F89248343CD01015DC0F3D7C12">2022</mods:date>
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<mods:number id="60168F32CA61126DC24FC04EFE5C0A11">2022-07-15</mods:number>
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<mods:number id="1041A66F014AA8A60B36E2DF0F679C1E">5165</mods:number>
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<mods:detail id="D98423E93C84F651C5ED43D27CF5AB27" type="issue">
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<mods:number id="065173A6E68F4139FFF8D8BCAC6CEB65">3</mods:number>
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<mods:classification id="3BA369A1369C80B5BEE28167F945779C">journal article</mods:classification>
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<mods:identifier id="D99273ECCA9D942A11FC71D67F50BAA4" type="CLB-Dataset">182450</mods:identifier>
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<mods:identifier id="587918E57B4715480E2CB026C70FD1FF" type="DOI">10.11646/zootaxa.5165.3.8</mods:identifier>
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<mods:identifier id="BD6162F2426013225678CD1CA0BDCEE8" type="ISSN">1175-5326</mods:identifier>
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</mods:mods>
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<treatment id="5D0DC56DFFD4FF8CFF01F987613DFBD9" ID-DOI="http://doi.org/10.5281/zenodo.6938936" ID-GBIF-Taxon="204252300" ID-Zenodo-Dep="6938936" LSID="urn:lsid:zoobank.org:act:568D8333-4E82-4A41-9AF9-AD23B2F45E8B" httpUri="http://treatment.plazi.org/id/5D0DC56DFFD4FF8CFF01F987613DFBD9" lastPageId="5" lastPageNumber="440" pageId="1" pageNumber="436">
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<subSubSection id="9DBE27F0FFD4FF88FF01F9876165F9D1" box="[151,532,1627,1654]" pageId="1" pageNumber="436" type="nomenclature">
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<paragraph id="D51B747BFFD4FF88FF01F9876165F9D1" blockId="1.[151,532,1627,1690]" box="[151,532,1627,1654]" pageId="1" pageNumber="436">
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<heading id="8E53C317FFD4FF88FF01F9876165F9D1" bold="true" box="[151,532,1627,1654]" fontSize="11" level="1" pageId="1" pageNumber="436" reason="1">
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<emphasis id="E7D0A869FFD4FF88FF01F9876165F9D1" bold="true" box="[151,532,1627,1654]" pageId="1" pageNumber="436">
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<taxonomicName id="12A40FF8FFD4FF88FF01F98762DEF9D1" authority="Alekseev & Bukejs, 2022" authorityName="Alekseev & Bukejs" authorityYear="2022" box="[151,431,1627,1654]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="436" phylum="Arthropoda" rank="species" species="praecursor" status="sp. nov.">
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<emphasis id="E7D0A869FFD4FF88FF01F98762DEF9D1" bold="true" box="[151,431,1627,1654]" italics="true" pageId="1" pageNumber="436">Eustrophus praecursor</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FCE31512FFD4FF88FE20F9876165F9D1" box="[438,532,1627,1653]" pageId="1" pageNumber="436" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="9DBE27F0FFD4FF88FF01F9A3627AF93E" box="[151,267,1663,1690]" pageId="1" pageNumber="436" type="description">
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<paragraph id="D51B747BFFD4FF88FF01F9A3627AF93E" blockId="1.[151,532,1627,1690]" box="[151,267,1663,1690]" pageId="1" pageNumber="436">
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||
(
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<figureCitation id="4D9F68FEFFD4FF88FF09F9A36273F93E" box="[159,258,1663,1690]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–5" captionStartId-0="2.[151,264,1947,1971]" captionStartId-1="3.[151,264,1933,1957]" captionTargetBox-0="[151,1435,594,1921]" captionTargetBox-1="[151,1435,181,1909]" captionTargetId-0="figure-199@2.[151,1435,594,1921]" captionTargetId-1="figure-17@3.[151,1435,181,1909]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1–2. Eustrophus praecursor sp. nov., holotype, No 6819 [MAIG]: 1—habitus, dorsal view; 2—habitus, ventral view. Scale bar = 1.0 mm." captionText-1="FIGURES 3–5. Eustrophus praecursor sp. nov., holotype, No 6819 [MAIG]: 3—details of head and prothorax, ventral view; 4—habitus, right lateral view; 5—right metatibia and metatarsus. Not to scale." figureDoi-0="http://doi.org/10.5281/zenodo.6838531" figureDoi-1="http://doi.org/10.5281/zenodo.6838537" httpUri-0="https://zenodo.org/record/6838531/files/figure.png" httpUri-1="https://zenodo.org/record/6838537/files/figure.png" pageId="1" pageNumber="436">Figs 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="9DBE27F0FFD4FF88FF01F91B67D0F81D" pageId="1" pageNumber="436" type="materials_examined">
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<paragraph id="D51B747BFFD4FF88FF01F91B61F7F8E9" blockId="1.[151,1437,1735,2014]" pageId="1" pageNumber="436">
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<emphasis id="E7D0A869FFD4FF88FF01F91B6234F945" bold="true" box="[151,325,1735,1761]" pageId="1" pageNumber="436">Type material.</emphasis>
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<typeStatus id="0A1FCAD9FFD4FF88FEDDF91B62C9F945" box="[331,440,1735,1761]" pageId="1" pageNumber="436" type="holotype">Holotype</typeStatus>
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: collection number No 6819 [
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<collectionCode id="B3B5ECBEFFD4FF88FC94F91B6023F945" box="[770,850,1735,1761]" pageId="1" pageNumber="436">MAIG</collectionCode>
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] (ex coll. Jonas Damzen 9230); adult, sex unknown. A complete beetle with a deformed right elytron is included in a transparent, yellow amber piece with dimensions of 6×
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<quantity id="125CD99EFFD4FF88FF20F8D36387F88E" box="[182,246,1807,1834]" metricMagnitude="-3" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="436" unit="mm" value="8.0">8 mm</quantity>
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and a maximum thickness of
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||
<quantity id="125CD99EFFD4FF88FDD5F8CC61F7F88E" box="[579,646,1808,1834]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="1" pageNumber="436" unit="mm" value="7.0">7 mm</quantity>
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; preserved without supplementary fixation. Syninclusions: few stellate trichomes of
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<taxonomicName id="12A40FF8FFD4FF88FEBDF8E862E2F8EA" authorityName="Dumortier" authorityYear="1829" box="[299,403,1844,1870]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="1" pageNumber="436" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
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and detrital particles.
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</paragraph>
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<paragraph id="D51B747BFFD4FF88FF51F88B6091F831" blockId="1.[151,1437,1735,2014]" pageId="1" pageNumber="436">
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<emphasis id="E7D0A869FFD4FF88FF51F88B621FF8D5" bold="true" box="[199,366,1879,1905]" pageId="1" pageNumber="436">
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<typeStatus id="0A1FCAD9FFD4FF88FF51F88B6273F8D5" box="[199,258,1879,1905]" pageId="1" pageNumber="436">Type</typeStatus>
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stratum.
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</emphasis>
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Baltic amber from amber-bearing blue Earth layers; Middle-Late Eocene, 48–34 Ma (
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<bibRefCitation id="B135098AFFD4FF88FAB8F88B6260F831" author="Sadowski, E. - M. & Seyfullah, L. J. & Schmidt, A. R. & Kunzmann, L." pageId="1" pageNumber="436" pagination="1 - 73" refId="ref3991" refString="Sadowski, E. - M., Seyfullah, L. J., Schmidt, A. R. & Kunzmann, L. (2017) Conifers of the ' Baltic amber forest' and their palaeoecological significance. Stapfia, 106, 1 - 73." type="journal article" year="2017">
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||
Sadowski
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<emphasis id="E7D0A869FFD4FF88FF01F8A063B9F831" box="[151,200,1915,1941]" italics="true" pageId="1" pageNumber="436">et al</emphasis>
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||
. 2017
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||
</bibRefCitation>
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||
;
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||
<bibRefCitation id="B135098AFFD4FF88FE8BF8A76178F831" author="Seyfullah, L. J. & Beimforde, C. & Dal Corso, J. & Perrichot, V. & Rikkinen, J. & Schmidt, A. R." box="[285,521,1915,1942]" pageId="1" pageNumber="436" pagination="1684 - 1714" refId="ref4040" refString="Seyfullah, L. J., Beimforde, C., Dal Corso, J., Perrichot, V., Rikkinen, J. & Schmidt, A. R. (2018) Production and preservation of resins - past and present. Biological Reviews, 93, 1684 - 1714. https: // doi. org / 10.1111 / brv. 12414" type="journal article" year="2018">
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||
Seyfullah
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<emphasis id="E7D0A869FFD4FF88FE19F8A062B1F831" box="[399,448,1915,1941]" italics="true" pageId="1" pageNumber="436">et al</emphasis>
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||
. 2018
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||
</bibRefCitation>
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||
;
|
||
<bibRefCitation id="B135098AFFD4FF88FD83F8A76195F831" author="Bukejs, A. & Alekseev, V. I. & Pollock, D. A." box="[533,740,1915,1942]" pageId="1" pageNumber="436" pagination="261 - 273" refId="ref3243" refString="Bukejs, A., Alekseev, V. I. & Pollock, D. A. (2019) Waidelotinae, a new subfamily of Pyrochroidae (Coleoptera: Tenebrionoidea) from Baltic amber of the Sambian peninsula and the interpretation of Sambian amber stratigraphy, age and location. Zootaxa, 4664 (2), 261 - 273. https: // doi. org / 10.11646 / zootaxa. 4664.2.8" type="journal article" year="2019">
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Bukejs
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<emphasis id="E7D0A869FFD4FF88FDFCF8A061EAF831" box="[618,667,1915,1941]" italics="true" pageId="1" pageNumber="436">et al</emphasis>
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||
. 2019
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||
</bibRefCitation>
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||
;
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||
<bibRefCitation id="B135098AFFD4FF88FD66F8A760A4F831" author="Kasinski, J. R. & Kramarska, R. & Slodkowska, B. & Sivkov, V. & Piwocki, M." box="[752,981,1915,1942]" pageId="1" pageNumber="436" pagination="29 - 53" refId="ref3511" refString="Kasinski, J. R., Kramarska, R., Slodkowska, B., Sivkov, V. & Piwocki, M. (2020) Paleocene and Eocene deposits on the eastern margin of the Gulf of Gdansk (Yantarny P- 1 bore hole, Kaliningrad region, Russia). Geological Quarterly, 64, 29 - 53. https: // doi. org / 10.7306 / gq. 1513" type="journal article" year="2020">
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||
Kasiński
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<emphasis id="E7D0A869FFD4FF88FCCFF8A060FBF831" box="[857,906,1915,1941]" italics="true" pageId="1" pageNumber="436">et al</emphasis>
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||
. 2020
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||
</bibRefCitation>
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||
).
|
||
</paragraph>
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||
<paragraph id="D51B747BFFD4FF88FF51F84367D0F81D" blockId="1.[151,1437,1735,2014]" box="[199,1185,1951,1978]" pageId="1" pageNumber="436">
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||
<materialsCitation id="65CC7E26FFD4FF88FF51F84367ECF81D" ID-GBIF-Occurrence="3962318301" box="[199,1181,1951,1978]" country="Russia" location="Yantarny village" pageId="1" pageNumber="436" specimenCount="1" stateProvince="Kaliningrad" typeStatus="holotype">
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||
<emphasis id="E7D0A869FFD4FF88FF51F8436217F81D" bold="true" box="[199,358,1951,1977]" pageId="1" pageNumber="436">
|
||
<typeStatus id="0A1FCAD9FFD4FF88FF51F8436273F81D" box="[199,258,1951,1977]" pageId="1" pageNumber="436">Type</typeStatus>
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||
locality.
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||
</emphasis>
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||
<location id="D07B22A0FFD4FF88FEFAF87C6154F81D" LSID="urn:lsid:plazi:treatment:5D0DC56DFFD4FF8CFF01F987613DFBD9:D07B22A0FFD4FF88FEFAF87C6154F81D" box="[364,549,1951,1978]" country="Russia" name="Yantarny village" pageId="1" pageNumber="436" stateProvince="Kaliningrad">Yantarny village</location>
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||
(formerly Palmnicken), the
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||
<collectingRegion id="1760BA99FFD4FF88FCF5F8436735F81D" box="[867,1092,1951,1977]" country="Russia" name="Kaliningrad" pageId="1" pageNumber="436">Kaliningrad Region</collectingRegion>
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||
,
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||
<collectingCountry id="ADB334EBFFD4FF88FBD9F84367ECF81D" box="[1103,1181,1951,1977]" name="Russia" pageId="1" pageNumber="436">Russia</collectingCountry>
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DBE27F0FFD4FF8CFF51F81F639AFD55" lastPageId="5" lastPageNumber="440" pageId="1" pageNumber="436" type="description">
|
||
<paragraph id="D51B747BFFD4FF8BFF51F81F6219FEB9" blockId="1.[151,1437,1735,2014]" lastBlockId="2.[151,1436,150,573]" lastPageId="2" lastPageNumber="437" pageId="1" pageNumber="436">
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||
<emphasis id="E7D0A869FFD4FF88FF51F81F6228F879" bold="true" box="[199,345,1987,2013]" pageId="1" pageNumber="436">Description.</emphasis>
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||
Measurements: body length (from anterior margin of pronotum to elytral apex along the midline) about
|
||
<quantity id="125CD99EFFD7FF8BFF4AFF4A6244FF15" box="[220,309,150,177]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="2" pageNumber="437" unit="mm" value="5.1">5.1 mm</quantity>
|
||
; maximum width across both elytra
|
||
<quantity id="125CD99EFFD7FF8BFD5AFF4B6053FF15" box="[716,802,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="2" pageNumber="437" unit="mm" value="2.2">2.2 mm</quantity>
|
||
(measured in basal one-third); pronotum length
|
||
<quantity id="125CD99EFFD7FF8BFAD6FF4B66E9FF15" box="[1344,1432,151,177]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="2" pageNumber="437" unit="mm" value="1.3">1.3 mm</quantity>
|
||
, pronotum maximum width
|
||
<quantity id="125CD99EFFD7FF8BFE45FF67615EFF71" box="[467,559,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="2" pageNumber="437" unit="mm" value="2.1">2.1 mm</quantity>
|
||
; elytra length
|
||
<quantity id="125CD99EFFD7FF8BFD41FF676040FF71" box="[727,817,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="2" pageNumber="437" unit="mm" value="3.8">3.8 mm</quantity>
|
||
, elytron maximum width
|
||
<quantity id="125CD99EFFD7FF8BFBC8FF6767CBFF71" box="[1118,1210,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="2" pageNumber="437" unit="mm" value="1.1">1.1 mm</quantity>
|
||
. Body oblong-oval, moderately convex dorsally; integument uniformly dark brown (as preserved); densely covered with fine, short, semierect setation.
|
||
</paragraph>
|
||
<paragraph id="D51B747BFFD7FF8BFF51FEFB673FFD99" blockId="2.[151,1436,150,573]" pageId="2" pageNumber="437">
|
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<emphasis id="E7D0A869FFD7FF8BFF51FEFB6273FEE5" box="[199,258,295,321]" pageId="2" pageNumber="437">Head</emphasis>
|
||
deflexed, uniformly and coarsely punctate. Compound eyes slightly flattened, large, with vertical diameter about 1.8× transverse diameter, deeply and narrowly emarginated around antennal insertions, without interfacetal setae; widely separated dorsally, distance between eyes about 1.5× as wide as transverse eye diameter. Antennae 11-segmented, not clubbed, with gradually widened distal antennomeres, moderately long, extending about to the elytral base. Scape subcylindrical, about 2.0× as long as pedicel; pedicel cylindrical, as long as antennomere 4; antennomeres 3–4 conical, antennomere 4 about 0.7× as wide as antennomere 5; antennomeres 5–10 trapezoidal, distinctly widened apically, as wide as long, flattened; antennomere 11 longer than antennomere 10, ovoid, pointed apically. Relative length ratios of antennomeres 1–11 equal to 10:5:6:5:7:7:7:7:7:7:9.
|
||
</paragraph>
|
||
<caption id="81DB24F3FFD7FF8BFF01F84762D0F873" ID-DOI="http://doi.org/10.5281/zenodo.6838531" ID-Zenodo-Dep="6838531" httpUri="https://zenodo.org/record/6838531/files/figure.png" pageId="2" pageNumber="437" startId="2.[151,264,1947,1971]" targetBox="[151,1435,594,1921]" targetPageId="2">
|
||
<paragraph id="D51B747BFFD7FF8BFF01F84762D0F873" blockId="2.[151,1436,1947,2007]" pageId="2" pageNumber="437">
|
||
<emphasis id="E7D0A869FFD7FF8BFF01F847624CF817" bold="true" box="[151,317,1947,1971]" pageId="2" pageNumber="437">FIGURES 1–2.</emphasis>
|
||
<taxonomicName id="12A40FF8FFD7FF8BFED0F8476142F817" authority="Alekseev & Bukejs, 2022" authorityName="Alekseev & Bukejs" authorityYear="2022" box="[326,563,1947,1971]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="437" phylum="Arthropoda" rank="species" species="praecursor" status="sp. nov.">
|
||
<emphasis id="E7D0A869FFD7FF8BFED0F8476142F817" box="[326,563,1947,1971]" italics="true" pageId="2" pageNumber="437">Eustrophus praecursor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7D0A869FFD7FF8BFDAAF84761FEF817" bold="true" box="[572,655,1947,1971]" pageId="2" pageNumber="437">
|
||
<taxonomicNameLabel id="FCE31512FFD7FF8BFDAAF84761FEF817" box="[572,655,1947,1971]" pageId="2" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No 6819 [MAIG]: 1—habitus, dorsal view; 2—habitus, ventral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="81DB24F3FFD6FF8AFF01F85160C3F86D" ID-DOI="http://doi.org/10.5281/zenodo.6838537" ID-Zenodo-Dep="6838537" httpUri="https://zenodo.org/record/6838537/files/figure.png" pageId="3" pageNumber="438" startId="3.[151,264,1933,1957]" targetBox="[151,1435,181,1909]" targetPageId="3">
|
||
<paragraph id="D51B747BFFD6FF8AFF01F85160C3F86D" blockId="3.[151,1436,1933,1993]" pageId="3" pageNumber="438">
|
||
<emphasis id="E7D0A869FFD6FF8AFF01F851624AF801" bold="true" box="[151,315,1933,1957]" pageId="3" pageNumber="438">FIGURES 3–5.</emphasis>
|
||
<taxonomicName id="12A40FF8FFD6FF8AFED4F851615DF801" authority="Alekseev & Bukejs, 2022" authorityName="Alekseev & Bukejs" authorityYear="2022" box="[322,556,1933,1957]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="438" phylum="Arthropoda" rank="species" species="praecursor" status="sp. nov.">
|
||
<emphasis id="E7D0A869FFD6FF8AFED4F851615DF801" box="[322,556,1933,1957]" italics="true" pageId="3" pageNumber="438">Eustrophus praecursor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7D0A869FFD6FF8AFDA5F85161F5F801" bold="true" box="[563,644,1933,1957]" pageId="3" pageNumber="438">
|
||
<taxonomicNameLabel id="FCE31512FFD6FF8AFDA5F85161F5F801" box="[563,644,1933,1957]" pageId="3" pageNumber="438" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype, No 6819 [MAIG]: 3—details of head and prothorax, ventral view; 4—habitus, right lateral view; 5—right metatibia and metatarsus. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="81DB24F3FFD1FF8DFF01FA2A6775F9F2" ID-DOI="http://doi.org/10.5281/zenodo.6838539" ID-Zenodo-Dep="6838539" httpUri="https://zenodo.org/record/6838539/files/figure.png" pageId="4" pageNumber="439" startId="4.[151,264,1526,1550]" targetBox="[151,1435,181,1501]" targetPageId="4">
|
||
<paragraph id="D51B747BFFD1FF8DFF01FA2A6775F9F2" blockId="4.[151,1437,1526,1622]" pageId="4" pageNumber="439">
|
||
<emphasis id="E7D0A869FFD1FF8DFF01FA2A624BF9AA" bold="true" box="[151,314,1526,1550]" pageId="4" pageNumber="439">FIGURES 6–8.</emphasis>
|
||
Cenozoic representatives of
|
||
<taxonomicName id="12A40FF8FFD1FF8DFDC8FA2A61A1F9AA" authorityName="Illiger" authorityYear="1802" box="[606,720,1526,1550]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="439" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E7D0A869FFD1FF8DFDC8FA2A61A1F9AA" box="[606,720,1526,1550]" italics="true" pageId="4" pageNumber="439">Eustrophus</emphasis>
|
||
</taxonomicName>
|
||
: 6—distribution of extant species (green area) and Eocene record of
|
||
<taxonomicName id="12A40FF8FFD1FF8DFA10FA2B6279F996" authority="Alekseev & Bukejs, 2022" authorityName="Alekseev & Bukejs" authorityYear="2022" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="439" phylum="Arthropoda" rank="species" species="praecursor" status="sp. nov.">
|
||
<emphasis id="E7D0A869FFD1FF8DFA10FA2B6279F996" italics="true" pageId="4" pageNumber="439">E. praecursor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7D0A869FFD1FF8DFE98F9C6622FF996" bold="true" box="[270,350,1562,1586]" pageId="4" pageNumber="439">
|
||
<taxonomicNameLabel id="FCE31512FFD1FF8DFE98F9C6622FF996" box="[270,350,1562,1586]" pageId="4" pageNumber="439" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(red dot); 7—extant European
|
||
<taxonomicName id="12A40FF8FFD1FF8DFD0AF9C7604FF996" authorityName="Fabricius" authorityYear="1792" box="[668,830,1562,1586]" class="Insecta" family="Mycetophagidae" genus="Mycetophagus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="439" phylum="Arthropoda" rank="species" species="dermestoides">
|
||
<emphasis id="E7D0A869FFD1FF8DFD0AF9C7604FF996" box="[668,830,1562,1586]" italics="true" pageId="4" pageNumber="439">E. dermestoides</emphasis>
|
||
</taxonomicName>
|
||
in the natural habitat from the Sambian Peninsula (Kaliningrad region, Russia), 26 May 2021, dorso-frontal view; 8—ibidem, dorso-lateral view.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D51B747BFFD1FF8DFF51F959617CF81B" blockId="4.[151,1437,1668,2019]" pageId="4" pageNumber="439">Pronotum semicircular, transverse, about 1.6× as wide as long, with maximum width in posterior one-fourth, as wide as elytra at base; pronotal disc slightly convex, with two paired very shallow, rounded subbasal impression. Lateral pronotal sides smooth, rounded. Pronotum weakly bisinuate at the base, with weak middle lobe, not margined posteriorly; evenly arcuate anteriorly. Anterior angles are weakly developed, obtusely rounded; posterior angles distinct, orthogonal. Pronotal surface uniformly and shallowly punctate; shortly pubescent. Prothoracic episternal suture absent. Prosternal punctation coarse and dense, the distance between punctures smaller than the diameter of one puncture, and interspaces convex. Hypomera rugosely punctate, without transverse suture. Procoxae are separated by the triangular prostenal process. Prosternal process elongate, with rounded apex, not extending to the posterior margin of procoxae.</paragraph>
|
||
<paragraph id="D51B747BFFD1FF8CFF51F8146018FEB9" blockId="4.[151,1437,1668,2019]" lastBlockId="5.[151,1437,151,1149]" lastPageId="5" lastPageNumber="440" pageId="4" pageNumber="439">
|
||
Elytra elongate, slightly convex, striate-punctate. Punctation small, deep and dense, arranged in longitudinal striae, each elytron with 10 complete striae. Interstriae are densely covered with finer punctures bearing short, semierect setae. Epipleura narrowed posteriorly, extending to elytral apex. Metanepistenum densely and finely punctate; distinctly subdivided in two parts: anterior triangular area and posterior
|
||
<emphasis id="E7D0A869FFD0FF8CFC5CFF036725FF5D" box="[970,1108,223,249]" pageId="5" pageNumber="440">longitudinal</emphasis>
|
||
, rectangle area. Metaventrite with dense, fine punctuation; disc convex, with sharp discrimen.
|
||
</paragraph>
|
||
<paragraph id="D51B747BFFD0FF8CFF51FEFB61C0FDBD" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="440">Legs slender, long, relatively similar in shape, finely and densely punctate. All coxae separated; procoxae large, nearly hemispherical, strongly convex; metacoxae simple. Femora slender, suboval, flattened; metafemora widest, about 3× as long as wide. Femora and tibiae are subequal in length. Tibiae almost straight; with two rather short (about 0.8× as long as metatibia width at apex), simple, paired apical spurs of equal length; outer surfaces of meso- and metatibiae with numerous oblique, comb-like ridges (with about 15 ridges on metatibia). Tarsal formula 5-5-4. All tarsomeres narrow, simple (non-lobed). Relative length ratios of metatarsomeres 1–4 equal to 14:7:5:5. Tarsal claws slender, without basal tooth or expansion.</paragraph>
|
||
<paragraph id="D51B747BFFD0FF8CFF51FDFF6663FDC5" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="440">Abdomen with five visible, freely articulated ventrites; densely covered with uniform, fine punctation and decumbent setation. Relative length ratios of ventrites 1–5 equal to 40:35:32:25:23 (measured medially).</paragraph>
|
||
<paragraph id="D51B747BFFD0FF8CFF51FDB7639AFD55" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="440">
|
||
<emphasis id="E7D0A869FFD0FF8CFF51FDB76277FD21" bold="true" box="[199,262,619,645]" pageId="5" pageNumber="440">Note.</emphasis>
|
||
The presence or absence of ovate, setiferous pit on the ventral surface of profemur is unclear due to legs position in the studied specimen. The genitalia are not exposed, making definite sexual determination impossible. The shape of scutellar shield is unclear (probably trapezoidal and slightly transverse) because it is covered by air bubble.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DBE27F0FFD0FF8CFF51FD276298FCF9" pageId="5" pageNumber="440" type="etymology">
|
||
<paragraph id="D51B747BFFD0FF8CFF51FD276298FCF9" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="440">
|
||
<emphasis id="E7D0A869FFD0FF8CFF51FD27623FFCB1" bold="true" box="[199,334,763,789]" pageId="5" pageNumber="440">Etymology.</emphasis>
|
||
The specific epithet “
|
||
<taxonomicName id="12A40FF8FFD0FF8CFDD3FD2761B0FCB0" authorityName="Alekseev & Bukejs" authorityYear="2022" box="[581,705,763,788]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="praecursor">
|
||
<emphasis id="E7D0A869FFD0FF8CFDD3FD2761B0FCB0" box="[581,705,763,788]" italics="true" pageId="5" pageNumber="440">praecursor</emphasis>
|
||
</taxonomicName>
|
||
” is used as noun in apposition and stems from the Latin substantive, meaning “forerunner, vanguard, predecessor”. The name refers to the pioneer status of this species, pre-dating the other species of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9DBE27F0FFD0FF8CFF51FCBA613DFBD9" pageId="5" pageNumber="440" type="diagnosis">
|
||
<paragraph id="D51B747BFFD0FF8CFF51FCBA613DFBD9" blockId="5.[151,1437,151,1149]" pageId="5" pageNumber="440">
|
||
<emphasis id="E7D0A869FFD0FF8CFF51FCBA62A0FC24" bold="true" box="[199,465,870,896]" pageId="5" pageNumber="440">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="12A40FF8FFD0FF8CFE4CFCBB61AEFC24" authority="Alekseev & Bukejs, 2022" authorityName="Alekseev & Bukejs" authorityYear="2022" box="[474,735,871,897]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="praecursor" status="sp. nov.">
|
||
<emphasis id="E7D0A869FFD0FF8CFE4CFCBB61AEFC24" box="[474,735,871,897]" italics="true" pageId="5" pageNumber="440">Eustrophus praecursor</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="E7D0A869FFD0FF8CFD7FFCBB6037FC25" bold="true" box="[745,838,871,897]" pageId="5" pageNumber="440">
|
||
<taxonomicNameLabel id="FCE31512FFD0FF8CFD7FFCBB6037FC25" box="[745,838,871,897]" pageId="5" pageNumber="440" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished from extant congeners by following characters: basal pronotal margination absent (in contrast to finely margined pronotum in
|
||
<taxonomicName id="12A40FF8FFD0FF8CFB4FFC576607FC00" authorityName="Say" authorityYear="1826" box="[1241,1398,907,932]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="tomentosus">
|
||
<emphasis id="E7D0A869FFD0FF8CFB4FFC576607FC00" box="[1241,1398,907,932]" italics="true" pageId="5" pageNumber="440">E. tomentosus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="12A40FF8FFD0FF8CFA12FC57625AFC6D" authorityName="Fabricius" authorityYear="1792" class="Insecta" family="Mycetophagidae" genus="Mycetophagus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="dermestoides">
|
||
<emphasis id="E7D0A869FFD0FF8CFA12FC57625AFC6D" italics="true" pageId="5" pageNumber="440">E. dermestoides</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="12A40FF8FFD0FF8CFEF1FC736282FC6C" authorityName="Lewis" authorityYear="1895" box="[359,499,943,968]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="niponicus">
|
||
<emphasis id="E7D0A869FFD0FF8CFEF1FC736282FC6C" box="[359,499,943,968]" italics="true" pageId="5" pageNumber="440">E. niponicus</emphasis>
|
||
</taxonomicName>
|
||
), and compound eyes rather widely separated by a distance about 1.5× as transverse eye diameter (in contrast to the narrowly separated eyes with the distance between them about 0.4× as transverse eye diameter in
|
||
<taxonomicName id="12A40FF8FFD0FF8CFED1FC2B629EFBB4" authorityName="Nikitsky" authorityYear="1998" box="[327,495,1015,1040]" class="Insecta" family="Tetratomidae" genus="Eustrophus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="species" species="yunnanensis">
|
||
<emphasis id="E7D0A869FFD0FF8CFED1FC2B629EFBB4" box="[327,495,1015,1040]" italics="true" pageId="5" pageNumber="440">E. yunnanensis</emphasis>
|
||
</taxonomicName>
|
||
). The new extinct species from Baltic amber can be easily distinguished from extinct
|
||
<taxonomicName id="12A40FF8FFD0FF8CFF01FBC7625AFB91" authorityName="Gistel" authorityYear="1856" box="[151,299,1051,1077]" class="Insecta" family="Tetratomidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="subFamily" subFamily="Eustrophinae">Eustrophinae</taxonomicName>
|
||
from Cretaceous ambers by the combination of tribal and generic characters, most obviously in the presence of oblique, comb-like ridges on outer surfaces of meso- and metatibiae in contrast to the simple tibiae in all other described fossil
|
||
<taxonomicName id="12A40FF8FFD0FF8CFE27FBBF6138FBD9" authorityName="Gistel" authorityYear="1856" box="[433,585,1123,1149]" class="Insecta" family="Tetratomidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="440" phylum="Arthropoda" rank="subFamily" subFamily="Eustrophinae">Eustrophinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |