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<mods:title id="05ED7E561D5B36F6A320CA6136F647C2">Revision of the giant geckos of New Caledonia (Reptilia: Diplodactylidae: Rhacodactylus)</mods:title>
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<mods:namePart id="B1440AFE6C04C9E8DD6E152551277F8A">Whitaker, Anthony H.</mods:namePart>
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<treatment id="5C4F87E1FFA8FFB1088FF931D293B1A9" ID-DOI="http://doi.org/10.5281/zenodo.6166578" ID-GBIF-Taxon="119521666" ID-Zenodo-Dep="6166578" LSID="urn:lsid:plazi:treatment:5C4F87E1FFA8FFB1088FF931D293B1A9" httpUri="http://treatment.plazi.org/id/5C4F87E1FFA8FFB1088FF931D293B1A9" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<taxonomicName id="13E64D74FFA8FFB0088FF931D631B0B6" box="[151,308,1712,1738]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="genus">
<emphasis id="E692EAE5FFA8FFB0088FF931D631B0B6" bold="true" box="[151,308,1712,1738]" pageId="22" pageNumber="23">Correlophus</emphasis>
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<paragraph id="D45936F7FFA8FFB1088FF976D213B760" blockId="22.[151,1436,1783,2025]" lastBlockId="23.[151,1436,151,284]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<taxonomicName id="13E64D74FFA8FFB0088FF976D6BDB173" box="[151,440,1783,1807]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="22" pageNumber="23" phylum="Chordata" rank="species" species="sarasinorum">
<emphasis id="E692EAE5FFA8FFB0088FF976D6BDB173" box="[151,440,1783,1807]" italics="true" pageId="22" pageNumber="23">Correlophus sarasinorum</emphasis>
</taxonomicName>
exhibits almost no intraspecific variation in the genetic markers we studied, even in the rapidly evolving mitochondrial genes (
<figureCitation id="4CDD2A72FFA8FFB00A71F89AD5D8B14F" box="[617,733,1819,1843]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="10.[151,250,1813,1836]" captionStartId-1="11.[151,250,1554,1577]" captionTargetBox-0="[185,1384,161,1812]" captionTargetBox-1="[185,1392,253,1495]" captionTargetId-0="figure@10.[185,1409,137,1826]" captionTargetId-1="figure@11.[167,1419,235,1518]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 1. Maximum likelihood tree based on the mitochondrial ND 2 gene and flanking tRNAs showing relationships among species of New Caledonian diplodactylids and their immediate sister-group, the Australian genus Pseudothecadactylus. Values subtending branches are maximum likelihood / Bayesian posterior probabilities above the line and maximum parsimony bootstrap values below the line. Dashes for posterior probabilities indicate no support for the maximum likelihood topology whereas dashes for maximum parsimony bootstraps indicates values &lt;50 %. Support values are not shown for conspecific relationships where samples differ by three or fewer bases. In the case of Rhacodactylus auriculatus southern ultramafic block samples are cumulatively represented as a triangle." captionText-1="FIGURE 2. Maximum likelihood tree based on the combined mitochondrial (ND 2 and flanking tRNAs) and nuclear genes (RAG 1) showing relationships among species of New Caledonian diplodactylids and their immediate sister-group, the Australian genus Pseudothecadactylus. The tree has been pruned to show only one exemplar for each taxon. Both intra- and interspecific patterns of relationship are nearly identical to those supported by ND 2 only (Fig. 1). Only within Dierogekko and Rhacodactylus sensu stricto are alternative patterns hypothesized (see text). Values subtending branches are maximum likelihood / Bayesian posterior probabilities above the line and maximum parsimony bootstrap values below the line. Dashes for posterior probabilities indicate no support for the maximum likelihood topology whereas dashes for maximum parsimony bootstraps indicates values &lt;50 %." httpUri-0="https://zenodo.org/record/211735/files/figure.png" httpUri-1="https://zenodo.org/record/211736/files/figure.png" pageId="22" pageNumber="23">Figs. 1–2</figureCitation>
). This is perhaps not surprising given the very restricted distribution of this species, which is limited to the southern ultramafic block of the Grande Terre (Bauer 1990; 
<bibRefCitation id="B0774B06FFA8FFB0088FF8E5D685B107" author="Bauer" box="[151,384,1891,1915]" pageId="22" pageNumber="23" refString="Bauer, A. M. &amp; Henle, K. (1994) Liste der rezenten Amphibien und Reptilien: Gekkonidae. Part 1, Australia and Oceania. Das Tierreich 109. Walter De Gruyter, Berlin, xiii + 306 pp." type="book" year="1994">Bauer &amp; Henle 1994</bibRefCitation>
; Bauer &amp; Sadlier 2000; 
<figureCitation id="4CDD2A72FFA8FFB00A94F8E2D5CAB107" box="[652,719,1891,1915]" captionStart="FIGURE 9" captionStartId="23.[151,250,1276,1299]" captionTargetBox="[161,1421,343,1246]" captionTargetId="figure@23.[151,1436,337,1255]" captionTargetPageId="23" captionText="FIGURE 9. Distribution map of Correlophus sarasinorum in southern New Caledonia. Type locality indicated by a green star, other vouchered localities by green circles, orange circle indicates unvouchered record. See Appendix for a list of localities mapped. Named subdivisions are communes." httpUri="https://zenodo.org/record/211743/files/figure.png" pageId="22" pageNumber="23">Fig. 9</figureCitation>
). Within this limited variation specimens from Fôrét Nord in the far south of the Plaine des Lacs differ only minimally from those further north at Mt. Koghis and Bois du Sud. Variation in color pattern has been previously mentioned (
<bibRefCitation id="B0774B06FFA8FFB00B05F82AD320B1BF" author="Bohme" box="[797,1061,1963,1987]" pageId="22" pageNumber="23" refString="Bohme, W. &amp; Henkel, F. - W. (1985) Zur Kenntnis der Herpetofauna Neukaledoniens, speziell der Gattung Rhacodactylus (Sauria: Gekkonidae). herpetofauna, 7 (34), 23 - 29." type="journal article" year="1985">Böhme &amp; Henkel 1985</bibRefCitation>
; 
<bibRefCitation id="B0774B06FFA8FFB00C28F82AD3C7B1BF" author="Henkel" box="[1072,1218,1963,1987]" pageId="22" pageNumber="23" refString="Henkel, F. - W. (1987) Haltung und Zucht von Rhacodactylus sarasinorum. Herpetofauna, 9 (50), 25 - 26." type="journal article" year="1987">Henkel 1987</bibRefCitation>
, 
<bibRefCitation id="B0774B06FFA8FFB00CD5F82AD20DB1BF" author="Henkel" box="[1229,1288,1963,1987]" pageId="22" pageNumber="23" refString="Henkel, F. - W. (1988) Rhacodactylus sarasinorum Roux [Amphib. - Reptil. Kartei: 125 - 128]. Sauria, 10 (4), 125 - 128." type="journal article" year="1988">1988</bibRefCitation>
; Bauer 1990; 
<bibRefCitation id="B0774B06FFA8FFB0088FF850D61BB19B" author="Myers" box="[151,286,1999,2025]" pageId="22" pageNumber="23" refString="Myers, A. (1997) The maintenance and breeding of Sarasin's gecko, Rhacodactylus sarasinorum Roux. Reptilian, 5 (3), 34 - 36, 38 - 39." type="journal article" year="1997">Myers 1997</bibRefCitation>
) but this appears to have no obvious phylogenetic basis. 
<bibRefCitation id="B0774B06FFA8FFB00B87F84ED36AB19B" box="[927,1135,1999,2024]" pageId="22" pageNumber="23" refString="Good, D. A., Bauer, A. M. &amp; Sadlier, R. A. (1997) Allozyme evidence for the phylogeny of giant New Caledonain geckos (Squamata: Diplodactylidae: Rhacodactylus), with comments on the status of R. leachianus henkeli. Australian Journal of Zoology, 45, 317 - 330." type="journal article">
Good 
<emphasis id="E692EAE5FFA8FFB00BFDF850D318B194" box="[997,1053,2000,2024]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
(1997)
</bibRefCitation>
found four fixed allozyme differences between single individuals of 
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<emphasis id="E692EAE5FFA9FFB10A6CFF19D421B6CC" box="[628,804,152,176]" italics="true" pageId="23" pageNumber="24">C. sarasinorum</emphasis>
</taxonomicName>
from Touaourou and Rivière Bleue, leading Bauer and Sadlier (2001) to hypothesize that two species might be represented. The information from the DNA sequence data presented here clearly contradicts this assumption and highlights the pitfalls of limited sampling. Several “morphs” are recognized by hobbyists (de 
<bibRefCitation id="B0774B06FFA9FFB10A1AFE85D5D6B760" author="Vosjoli" box="[514,723,260,284]" pageId="23" pageNumber="24" refString="de Vosjoli, P., Fast, F. &amp; Repashy, A. (2003) Rhacodactylus, the Complete Guide to their Selelction and Care. Advanced Visions, Inc., Vista, California. 296 pp." type="book" year="2003">
Vosjoli 
<emphasis id="E692EAE5FFA9FFB10A40FE84D595B760" box="[600,656,260,284]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2003
</bibRefCitation>
), but these are also of no phylogenetic significance.
</paragraph>
<caption id="8099667FFFA9FFB1088FFB7DD567B32C" httpUri="https://zenodo.org/record/211743/files/figure.png" pageId="23" pageNumber="24" targetBox="[161,1421,343,1246]" targetPageId="23">
<paragraph id="D45936F7FFA9FFB1088FFB7DD567B32C" blockId="23.[151,1436,1276,1360]" pageId="23" pageNumber="24">
<emphasis id="E692EAE5FFA9FFB1088FFB7DD617B36E" bold="true" box="[151,274,1276,1299]" pageId="23" pageNumber="24">FIGURE 9.</emphasis>
Distribution map of 
<taxonomicName id="13E64D74FFA9FFB109FFFB7CD5E8B36F" box="[487,749,1277,1299]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="sarasinorum">
<emphasis id="E692EAE5FFA9FFB109FFFB7CD5E8B36F" box="[487,749,1277,1299]" italics="true" pageId="23" pageNumber="24">Correlophus sarasinorum</emphasis>
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in southern New Caledonia. Type locality indicated by a green star, other vouchered localities by green circles, orange circle indicates unvouchered record. See Appendix for a list of localities mapped. Named subdivisions are communes.
</paragraph>
</caption>
<paragraph id="D45936F7FFA9FFB108DFFA20D3A5B115" blockId="23.[151,1436,1441,2005]" pageId="23" pageNumber="24">
<taxonomicName id="13E64D74FFA9FFB108DFFA20D6A9B3C5" box="[199,428,1441,1465]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA9FFB108DFFA20D6A9B3C5" box="[199,428,1441,1465]" italics="true" pageId="23" pageNumber="24">Correlophus ciliatus</emphasis>
</taxonomicName>
—The greatest intraspecific genetic divergence within any giant 
<taxonomicName id="13E64D74FFA9FFB10C67FA20D3C7B3C5" box="[1151,1218,1441,1465]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
was seen within 
<taxonomicName id="13E64D74FFA9FFB10D9AFA23D7EFB3A1" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA9FFB10D9AFA23D7EFB3A1" italics="true" pageId="23" pageNumber="24">C. ciliatus</emphasis>
</taxonomicName>
(
<figureCitation id="4CDD2A72FFA9FFB108E3FA44D66CB3A1" box="[251,361,1477,1501]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="10.[151,250,1813,1836]" captionStartId-1="11.[151,250,1554,1577]" captionTargetBox-0="[185,1384,161,1812]" captionTargetBox-1="[185,1392,253,1495]" captionTargetId-0="figure@10.[185,1409,137,1826]" captionTargetId-1="figure@11.[167,1419,235,1518]" captionTargetPageId-0="10" captionTargetPageId-1="11" captionText-0="FIGURE 1. Maximum likelihood tree based on the mitochondrial ND 2 gene and flanking tRNAs showing relationships among species of New Caledonian diplodactylids and their immediate sister-group, the Australian genus Pseudothecadactylus. Values subtending branches are maximum likelihood / Bayesian posterior probabilities above the line and maximum parsimony bootstrap values below the line. Dashes for posterior probabilities indicate no support for the maximum likelihood topology whereas dashes for maximum parsimony bootstraps indicates values &lt;50 %. Support values are not shown for conspecific relationships where samples differ by three or fewer bases. In the case of Rhacodactylus auriculatus southern ultramafic block samples are cumulatively represented as a triangle." captionText-1="FIGURE 2. Maximum likelihood tree based on the combined mitochondrial (ND 2 and flanking tRNAs) and nuclear genes (RAG 1) showing relationships among species of New Caledonian diplodactylids and their immediate sister-group, the Australian genus Pseudothecadactylus. The tree has been pruned to show only one exemplar for each taxon. Both intra- and interspecific patterns of relationship are nearly identical to those supported by ND 2 only (Fig. 1). Only within Dierogekko and Rhacodactylus sensu stricto are alternative patterns hypothesized (see text). Values subtending branches are maximum likelihood / Bayesian posterior probabilities above the line and maximum parsimony bootstrap values below the line. Dashes for posterior probabilities indicate no support for the maximum likelihood topology whereas dashes for maximum parsimony bootstraps indicates values &lt;50 %." httpUri-0="https://zenodo.org/record/211735/files/figure.png" httpUri-1="https://zenodo.org/record/211736/files/figure.png" pageId="23" pageNumber="24">Figs. 1–2</figureCitation>
). This species was described in 1866 (
<figureCitation id="4CDD2A72FFA9FFB10B30FA44D47EB3A1" box="[808,891,1477,1501]" captionStart="FIGURE 10" captionStartId="24.[151,250,1052,1075]" captionTargetBox="[212,1374,416,1016]" captionTargetId="figure@24.[205,1382,407,1030]" captionTargetPageId="24" captionText="FIGURE 10. Lectotype of Correlophus ciliatus (MNHN 701 A) showing the absence of lumbosacral tubercles and presence of heterogeneous dorsolateral scalation. Photo courtesy of Muséum National d’Histoire Naturelle, Paris." httpUri="https://zenodo.org/record/211744/files/figure.png" pageId="23" pageNumber="24">Fig. 10</figureCitation>
) and was apparently not uncommon in that era (
<bibRefCitation id="B0774B06FFA9FFB10886FA68D62EB07D" author="Bavay" box="[158,299,1513,1537]" pageId="23" pageNumber="24" refString="Bavay, A. (1869) Catalogue des reptiles de la Nouvelle-Caledonie et description d'especes nouvelles. Memoires de la Societe Linneenne de Normandie, 15, 1 - 37. Bocage, J. V. Barboza du (1873 a) Note sur quelques geckotiens nouveaux ou peu connus de la Nouvelle Caledonie. Jornal de Sciencias Mathematicas, Physicas, e Naturaes, Academia Real das Sciencas de Lisboa, 4, 201 - 207." type="journal article" year="1869">Bavay 1869</bibRefCitation>
). It was then “lost” to science for over 100 years and considered likely to be extinct (
<bibRefCitation id="B0774B06FFA9FFB10CFCFA68D7D5B059" author="Bauer" pageId="23" pageNumber="24" refString="Bauer, A. M. &amp; Sadlier, R. A. (1993) Systematics, biogeography and conservation of the lizards of New Caledonia. Biodiversity Letters, 1, 107 - 122." type="journal article" year="1993">Bauer &amp; Sadlier 1993</bibRefCitation>
) until rediscovered on the Île des Pins in the 1990s (
<bibRefCitation id="B0774B06FFA9FFB10B06F98CD4B0B059" author="Storelli" box="[798,949,1549,1573]" pageId="23" pageNumber="24" refString="Storelli, A. (1994) Gecko nouveau! On a retrouve le ciliatus! Les Nouvelles Hebdo n ° 320 (21 avril 1994), 16 - 17." type="journal article" year="1994">Storelli 1994</bibRefCitation>
; 
<bibRefCitation id="B0774B06FFA9FFB10BD9F98CD3D5B059" author="Seipp" box="[961,1232,1549,1573]" pageId="23" pageNumber="24" refString="Seipp, R. &amp; Klemmer, K. (1994) Wiederentdeckung von Rhacodactylus ciliatus Guichenot 1866 im Suden Neukaledoniens (Reptilia: Sauria: Gekkonidae). Senckenbergiana Biologica, 24, 199 - 204." type="journal article" year="1994">Seipp &amp; Klemmer 1994</bibRefCitation>
; 
<bibRefCitation id="B0774B06FFA9FFB10CC4F98CD295B059" author="Kullmann" box="[1244,1424,1549,1573]" pageId="23" pageNumber="24" refString="Kullimann, B. (1995) Uber die Wiederentdeckung des Kronengeckos (Rhacodactylus ciliatus) in Neu Kaledonien. Elaphe, 3, 68 - 71." type="journal article" year="1995">Kullmann 1995</bibRefCitation>
). It was subsequently found on several smaller satellite islands around Île des Pins (de Vosjoli 1995) and in the southern Grande Terre (
<bibRefCitation id="B0774B06FFA9FFB109BEF9D4D5B0B011" author="Girard" box="[422,693,1621,1645]" pageId="23" pageNumber="24" refString="Girard, F. &amp; Heuclin, D. (1998) Premiere mention de gecko Rhacodactylus ciliatus sur la Grande Terre (Nouvelle-Caledonie) depuis sa description en 1866. Bulletin de la Societe Herpetologique de France, 85 - 86, 60 - 61." type="journal article" year="1998">Girard &amp; Heuclin 1998</bibRefCitation>
; Bauer &amp; Sadlier 2000, 2001; 
<figureCitation id="4CDD2A72FFA9FFB10C0DF9D4D362B011" box="[1045,1127,1621,1645]" captionStart="FIGURE 11" captionStartId="25.[151,250,1357,1380]" captionTargetBox="[160,1426,418,1328]" captionTargetId="figure@25.[151,1436,409,1336]" captionTargetPageId="25" captionText="FIGURE 11. Distribution map of Correlophus ciliatus (circles) and C. belepensis sp. nov. (red star). For C. ciliatus green circles symbols represent vouchered records, orange circles represent unvouchered sight or literature records. Vouchered records from the Île des Pins lack precise localities. See Appendix for a list of localities mapped." httpUri="https://zenodo.org/record/211745/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
). Since then it has become one of the most popular of all lizard pets and is bred in at least the tens of thousands around the world (
<bibRefCitation id="B0774B06FFA9FFB10D07F9F8D639B0C9" author="Baldwin" pageId="23" pageNumber="24" refString="Baldwin, R. &amp; Repashey, A. (1998) Back with a vengeance: the New Caledonian crested gecko (Rhacodactylus ciliatus). Reptiles, 6 (4), 32 - 42." type="journal article" year="1998">Baldwin &amp; Repashy 1998</bibRefCitation>
; 
<bibRefCitation id="B0774B06FFA9FFB10953F91CD6C3B0C9" author="Both" box="[331,454,1693,1717]" pageId="23" pageNumber="24" refString="Both, A. R. (1999) The New Caledonian crested gecko (Rhacodactylus ciliatus). Reptile Hobbyist, 4 (5), 36 - 41." type="journal article" year="1999">Both 1999</bibRefCitation>
; 
<bibRefCitation id="B0774B06FFA9FFB109CDF91CD550B0C9" author="Bach" box="[469,597,1693,1717]" pageId="23" pageNumber="24" refString="Bach, S. (2006) Der Kronengecko, Rhacodactylus ciliatus. Natur und Tier-Verlag, Munster. 64 pp." type="book" year="2006">Bach 2006</bibRefCitation>
). All or most of these animals appear to originate from the Île des Pins, rather than Grande Terre. We found very little divergence between Île des Pins specimens and those from Rivière Bleue on the mainland, but quite deep divergences, comparable to the deepest within 
<taxonomicName id="13E64D74FFA9FFB10C4AF967D3F5B081" box="[1106,1264,1765,1789]" class="Reptilia" family="Diplodactylidae" genus="Rhacodactylus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="auriculatus">
<emphasis id="E692EAE5FFA9FFB10C4AF967D3F5B081" box="[1106,1264,1765,1789]" italics="true" pageId="23" pageNumber="24">R. auriculatus</emphasis>
</taxonomicName>
, between these and a single specimen from Mt. Dzumac, only about 
<quantity id="131E9B12FFA9FFB10AF5F888D430B15D" box="[749,821,1801,1825]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="23" pageNumber="24" unit="km" value="20.0">20 km</quantity>
distant from Rivière Bleue. However, specimens from the recently discovered population from the Îles Belep (
<bibRefCitation id="B0774B06FFA9FFB10B3DF8ACD319B139" author="Whitaker" box="[805,1052,1837,1861]" pageId="23" pageNumber="24" refString="Whitaker, A. H., Sadlier, R. A., Bauer, A. M. &amp; Whitaker, V. A. (2004) Biodiversity and conservation status of lizards in threatened and restricted habitats of north-western New Caledonia. Unpublished report by Whitaker Consultants Limited to Direction du Developpement Economique et de l'Environnement, Province Nord, Kone, New Caledonia, viii + 106 pp." type="book" year="2004">
Whitaker 
<emphasis id="E692EAE5FFA9FFB10B82F8AFD4D3B139" box="[922,982,1837,1861]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2004
</bibRefCitation>
; Wirth &amp; Peukert 2009) were as divergent as the most deeply-divergent splits between species in the 
<taxonomicName id="13E64D74FFA9FFB10B8CF8D3D353B115" box="[916,1110,1874,1897]" class="Reptilia" family="Diplodactylidae" genus="Bavayia" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="sauvagii">
<emphasis id="E692EAE5FFA9FFB10B8CF8D3D353B115" box="[916,1110,1874,1897]" italics="true" pageId="23" pageNumber="24">Bavayia sauvagii</emphasis>
</taxonomicName>
clade.
</paragraph>
<paragraph id="D45936F7FFA9FFB108DFF8F4D293B1A9" blockId="23.[151,1436,1441,2005]" pageId="23" pageNumber="24">
Further, there are morphological differences between the southern and northern populations concordant with the genetic differences retrieved, lending futher support to the recognition of the Belep Island populations as an independent evolutionary lineage, and on these criteria we here recognize this northern “ 
<taxonomicName id="13E64D74FFA9FFB10C6AF83CD3C1B1A9" box="[1138,1220,1981,2005]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="24" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA9FFB10C6AF83CD3C1B1A9" box="[1138,1220,1981,2005]" italics="true" pageId="23" pageNumber="24">ciliatus</emphasis>
</taxonomicName>
” as a new species:
</paragraph>
</subSubSection>
</treatment>
</document>