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<document id="B9E40C95E90B9B94B7FCECEA89EE63EA" ID-DOI="10.5281/zenodo.211734" ID-GBIF-Dataset="1837d7e6-3418-4081-9525-50f0b78d2125" ID-ISSN="1175-5326" ID-Zenodo-Dep="211734" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460502342988" checkinUser="plazi" docAuthor="Bauer, Aaron M., Jackman, Todd R., Sadlier, Ross A. &amp; Whitaker, Anthony H." docDate="2012" docId="5C4F87E1FFA6FFB9088FFF16D643B788" docLanguage="en" docName="zt03404p052.pdf" docOrigin="Zootaxa 3404" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Correlophus belepensis Bauer, Whitaker, Sadlier &amp; Jackman, sp. nov." docType="treatment" docVersion="10" lastPageNumber="32" masterDocId="A076FF99FFBEFFA60818FF81D705B67C" masterDocTitle="Revision of the giant geckos of New Caledonia (Reptilia: Diplodactylidae: Rhacodactylus)" masterLastPageNumber="52" masterPageNumber="1" pageNumber="25" updateTime="1698312116073" updateUser="plazi">
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<mods:title id="08B2FBE7F21E1415F3897B89BEBC80AB">Revision of the giant geckos of New Caledonia (Reptilia: Diplodactylidae: Rhacodactylus)</mods:title>
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<mods:namePart id="230F3732F0F238C3E7C0CEFB52669C49">Bauer, Aaron M.</mods:namePart>
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<mods:namePart id="946D3A3A677E4D5C98C6E707697299C1">Jackman, Todd R.</mods:namePart>
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<mods:namePart id="E8BBFD6CBC085CC7BEFFB230B2D01165">Sadlier, Ross A.</mods:namePart>
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<mods:namePart id="A5062776536313FB83F69EFC4AA39E4F">Whitaker, Anthony H.</mods:namePart>
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<treatment id="5C4F87E1FFA6FFB9088FFF16D643B788" ID-DOI="http://doi.org/10.5281/zenodo.6166580" ID-GBIF-Taxon="119521669" ID-Zenodo-Dep="6166580" LSID="urn:lsid:plazi:treatment:5C4F87E1FFA6FFB9088FFF16D643B788" httpUri="http://treatment.plazi.org/id/5C4F87E1FFA6FFB9088FFF16D643B788" lastPageId="31" lastPageNumber="32" pageId="24" pageNumber="25">
<subSubSection id="9CFC657CFFA6FFBE088FFF16D62BB6AE" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="D45936F7FFA6FFBE088FFF16D4E9B6CD" blockId="24.[151,1004,151,210]" box="[151,1004,151,177]" pageId="24" pageNumber="25">
<heading id="8F11819BFFA6FFBE088FFF16D4E9B6CD" bold="true" box="[151,1004,151,177]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="E692EAE5FFA6FFBE088FFF16D4E9B6CD" bold="true" box="[151,1004,151,177]" pageId="24" pageNumber="25">
<taxonomicName id="13E64D74FFA6FFBE088FFF16D48DB6CD" authority="Bauer, Whitaker, Sadlier &amp; Jackman" authorityName="Bauer, Whitaker, Sadlier &amp; Jackman" box="[151,904,151,177]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA6FFBE088FFF16D6B4B6CD" bold="true" box="[151,433,151,177]" italics="true" pageId="24" pageNumber="25">Correlophus belepensis</emphasis>
Bauer, Whitaker, Sadlier &amp; Jackman
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA6FFBE0B97FF16D4E9B6CD" box="[911,1004,151,177]" pageId="24" pageNumber="25" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D45936F7FFA6FFBE088FFF3BD62BB6AE" blockId="24.[151,1004,151,210]" box="[151,302,186,210]" pageId="24" pageNumber="25">
(
<figureCitation id="4CDD2A72FFA6FFBE0887FF3BD623B6AE" box="[159,294,186,210]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStartId-0="26.[151,250,1177,1200]" captionStartId-1="26.[151,250,1817,1840]" captionStartId-2="27.[151,250,1318,1341]" captionStartId-3="29.[151,250,1469,1492]" captionTargetBox-0="[151,1435,445,1156]" captionTargetBox-1="[151,1435,1280,1794]" captionTargetBox-2="[151,1436,841,1296]" captionTargetBox-3="[217,1369,733,1447]" captionTargetId-0="figure@26.[151,1436,445,1156]" captionTargetId-1="figure@26.[151,1436,1280,1796]" captionTargetId-2="figure@27.[151,1436,841,1297]" captionTargetId-3="figure@29.[217,1369,733,1448]" captionTargetPageId-0="26" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="29" captionText-0="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." captionText-1="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." captionText-2="FIGURE 14. Life photographs of Correlophus belepensis sp. nov. (A) Male holotype (MNHN 2011.1100) and (B) smaller female paratype (CAS 250865). Photos by A. H. Whitaker." captionText-3="FIGURE 15. Type series of Correlophus belepensis sp. nov. showing variation in color and in lumbosacral tubercles. Left to right: holotype (MNHN 2011.1100), paratypes (AMS R 161282, AMS R 161283, CAS 250865). Photo by A. H. Whitaker." httpUri-0="https://zenodo.org/record/211746/files/figure.png" httpUri-1="https://zenodo.org/record/211747/files/figure.png" httpUri-2="https://zenodo.org/record/211748/files/figure.png" httpUri-3="https://zenodo.org/record/211749/files/figure.png" pageId="24" pageNumber="25">Figs. 1215</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9CFC657CFFA6FFBE088FFE82D25BB2EA" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="D45936F7FFA6FFBE088FFE82D51EB71E" blockId="24.[151,1436,258,355]" pageId="24" pageNumber="25">
<emphasis id="E692EAE5FFA6FFBE088FFE82D60EB767" bold="true" box="[151,267,259,283]" pageId="24" pageNumber="25">
<typeStatus id="0B5D8855FFA6FFBE088FFE82D603B767" box="[151,262,259,283]" pageId="24" pageNumber="25" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="B2F7AE32FFA6FFBE090AFE82D66DB766" LSID="urn:lsid:biocol.org:col:34988" box="[274,360,259,282]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="24" pageNumber="25">MNHN</collectionCode>
2011.1100 (formerly
<collectionCode id="B2F7AE32FFA6FFBE0A47FE83D599B766" box="[607,668,258,282]" pageId="24" pageNumber="25">AMS</collectionCode>
<date id="A0581037FFA6FFBE0ABCFE83D5F9B766" box="[676,764,258,282]" pageId="24" pageNumber="25" value="81-12-16">161281</date>
), adult male (
<figureCitation id="4CDD2A72FFA6FFBE0B8DFE83D325B766" box="[917,1056,258,282]" captionStart-0="FIGURE 12" captionStart-1="FIGURE 13" captionStart-2="FIGURE 14" captionStart-3="FIGURE 15" captionStartId-0="26.[151,250,1177,1200]" captionStartId-1="26.[151,250,1817,1840]" captionStartId-2="27.[151,250,1318,1341]" captionStartId-3="29.[151,250,1469,1492]" captionTargetBox-0="[151,1435,445,1156]" captionTargetBox-1="[151,1435,1280,1794]" captionTargetBox-2="[151,1436,841,1296]" captionTargetBox-3="[217,1369,733,1447]" captionTargetId-0="figure@26.[151,1436,445,1156]" captionTargetId-1="figure@26.[151,1436,1280,1796]" captionTargetId-2="figure@27.[151,1436,841,1297]" captionTargetId-3="figure@29.[217,1369,733,1448]" captionTargetPageId-0="26" captionTargetPageId-1="26" captionTargetPageId-2="27" captionTargetPageId-3="29" captionText-0="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." captionText-1="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." captionText-2="FIGURE 14. Life photographs of Correlophus belepensis sp. nov. (A) Male holotype (MNHN 2011.1100) and (B) smaller female paratype (CAS 250865). Photos by A. H. Whitaker." captionText-3="FIGURE 15. Type series of Correlophus belepensis sp. nov. showing variation in color and in lumbosacral tubercles. Left to right: holotype (MNHN 2011.1100), paratypes (AMS R 161282, AMS R 161283, CAS 250865). Photo by A. H. Whitaker." httpUri-0="https://zenodo.org/record/211746/files/figure.png" httpUri-1="https://zenodo.org/record/211747/files/figure.png" httpUri-2="https://zenodo.org/record/211748/files/figure.png" httpUri-3="https://zenodo.org/record/211749/files/figure.png" pageId="24" pageNumber="25">Figs. 1215</figureCitation>
).
<collectingCountry id="ACF17667FFA6FFBE0C2CFE82D3E1B766" box="[1076,1252,258,282]" name="New Caledonia" pageId="24" pageNumber="25">New Caledonia</collectingCountry>
, Province Nord, Îles Belep, Île Art,
<quantity id="131E9B12FFA6FFBE0968FEA7D6ACB742" box="[368,425,294,318]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="24" pageNumber="25" unit="km" value="2.0">2 km</quantity>
E Waala, Wênè Côgat,
<geoCoordinate id="B1D25030FFA6FFBE0AAFFEA7D456B742" box="[695,851,294,318]" direction="south" orientation="latitude" pageId="24" pageNumber="25" precision="1" value="-19.713028">19°42'46.9&quot; S</geoCoordinate>
,
<geoCoordinate id="B1D25030FFA6FFBE0B78FEA7D308B742" box="[864,1037,294,318]" direction="east" orientation="longitude" pageId="24" pageNumber="25" precision="1" value="163.66046">163°39'37.7&quot; E</geoCoordinate>
,
<specimenCount id="C2E0FD7EFFA6FFBE0C02FEA7D36DB742" box="[1050,1128,294,318]" pageId="24" pageNumber="25" type="generic">
<quantity id="131E9B12FFA6FFBE0C02FEA7D361B742" box="[1050,1124,294,318]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="25" unit="m" value="230.0">230 m</quantity>
.
</specimenCount>
Collected
<date id="A0581037FFA6FFBE0CFAFEA7D27CB742" box="[1250,1401,294,319]" pageId="24" pageNumber="25" value="2002-05-23">23 May 2002</date>
by A.H. Whitaker and V.A. Whitaker.
</paragraph>
<caption id="8099667FFFA6FFBE088FFB9DD39EB22D" httpUri="https://zenodo.org/record/211744/files/figure.png" pageId="24" pageNumber="25" targetBox="[212,1374,416,1016]" targetPageId="24">
<paragraph id="D45936F7FFA6FFBE088FFB9DD39EB22D" blockId="24.[151,1436,1052,1105]" pageId="24" pageNumber="25">
<emphasis id="E692EAE5FFA6FFBE088FFB9DD61AB24D" bold="true" box="[151,287,1052,1075]" pageId="24" pageNumber="25">FIGURE 10.</emphasis>
Lectotype of
<taxonomicName id="13E64D74FFA6FFBE09B4FB9DD578B24E" box="[428,637,1052,1074]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA6FFBE09B4FB9DD578B24E" box="[428,637,1052,1074]" italics="true" pageId="24" pageNumber="25">Correlophus ciliatus</emphasis>
</taxonomicName>
(MNHN 701A) showing the absence of lumbosacral tubercles and presence of heterogeneous dorsolateral scalation. Photo courtesy of Muséum National dHistoire Naturelle, Paris.
</paragraph>
</caption>
<paragraph id="D45936F7FFA6FFBE08DFFBFFD25BB2EA" blockId="24.[151,1437,1150,2002]" box="[199,1374,1150,1174]" pageId="24" pageNumber="25">
<emphasis id="E692EAE5FFA6FFBE08DFFBFFD643B2EA" bold="true" box="[199,326,1150,1174]" pageId="24" pageNumber="25">
<typeStatus id="0B5D8855FFA6FFBE08DFFBFFD644B2EA" box="[199,321,1150,1174]" pageId="24" pageNumber="25" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="B2F7AE32FFA6FFBE0954FBFFD68CB2EA" box="[332,393,1150,1174]" pageId="24" pageNumber="25">AMS</collectionCode>
R
<date id="A0581037FFA6FFBE09BBFBFFD6FDB2EA" box="[419,504,1150,1174]" pageId="24" pageNumber="25" value="82-12-16">161282</date>
283,
<collectionCode id="B2F7AE32FFA6FFBE0A25FBFFD576B2EA" LSID="urn:lsid:biocol.org:col:34700" box="[573,627,1150,1174]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" name="California Academy of Sciences" pageId="24" pageNumber="25">CAS</collectionCode>
<date id="A0581037FFA6FFBE0A62FBFFD5CBB2EA" box="[634,718,1150,1174]" pageId="24" pageNumber="25" value="65-08-25">250865</date>
(formerly
<collectionCode id="B2F7AE32FFA6FFBE0B5FFBFFD486B2EA" box="[839,899,1150,1174]" pageId="24" pageNumber="25">AMS</collectionCode>
R
<date id="A0581037FFA6FFBE0B86FBFFD4F6B2EA" box="[926,1011,1150,1174]" pageId="24" pageNumber="25" value="84-12-16">161284</date>
) (
<figureCitation id="4CDD2A72FFA6FFBE0C12FBFFD358B2EA" box="[1034,1117,1150,1174]" captionStart="FIGURE 15" captionStartId="29.[151,250,1469,1492]" captionTargetBox="[217,1369,733,1447]" captionTargetId="figure@29.[217,1369,733,1448]" captionTargetPageId="29" captionText="FIGURE 15. Type series of Correlophus belepensis sp. nov. showing variation in color and in lumbosacral tubercles. Left to right: holotype (MNHN 2011.1100), paratypes (AMS R 161282, AMS R 161283, CAS 250865). Photo by A. H. Whitaker." httpUri="https://zenodo.org/record/211749/files/figure.png" pageId="24" pageNumber="25">Fig. 15</figureCitation>
). Data as for
<typeStatus id="0B5D8855FFA6FFBE0CECFBFFD25FB2EA" box="[1268,1370,1150,1174]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="9CFC657CFFA6FFBE08DFFB20D47BB312" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="D45936F7FFA6FFBE08DFFB20D47BB312" blockId="24.[151,1437,1150,2002]" pageId="24" pageNumber="25">
<emphasis id="E692EAE5FFA6FFBE08DFFB20D644B2C6" bold="true" box="[199,321,1185,1210]" pageId="24" pageNumber="25">Diagnosis.</emphasis>
<taxonomicName id="13E64D74FFA6FFBE0955FB23D55FB2C6" box="[333,602,1186,1210]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA6FFBE0955FB23D55FB2C6" box="[333,602,1186,1210]" italics="true" pageId="24" pageNumber="25">Correlophus belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA6FFBE0A7EFB23D5C0B2C6" box="[614,709,1186,1210]" pageId="24" pageNumber="25" rank="species">
<emphasis id="E692EAE5FFA6FFBE0A7EFB23D5C0B2C6" bold="true" box="[614,709,1186,1210]" pageId="24" pageNumber="25">sp. nov.</emphasis>
</taxonomicNameLabel>
is a large (to
<quantity id="131E9B12FFA6FFBE0B6EFB23D4DDB2C6" box="[886,984,1186,1210]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="24" pageNumber="25" unit="mm" value="100.0">100 mm</quantity>
SVL) diplodactylid. It differs from
<taxonomicName id="13E64D74FFA6FFBE0D9BFB22D624B2A2" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="sarasinorum">
<emphasis id="E692EAE5FFA6FFBE0D9BFB22D624B2A2" italics="true" pageId="24" pageNumber="25">C. sarasinorum</emphasis>
</taxonomicName>
in possessing a prominent crest of spinose scales extending from behind the orbit to the shoulder region. It is most similar to its sister taxon
<taxonomicName id="13E64D74FFA6FFBE0A0CFB6AD581B37E" box="[532,644,1258,1282]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA6FFBE0A0CFB6AD581B37E" box="[532,644,1258,1282]" italics="true" pageId="24" pageNumber="25">C. ciliatus</emphasis>
</taxonomicName>
, but may be distinguished from it by its homogeneous dorsolateral trunk scalation (
<figureCitation id="4CDD2A72FFA6FFBE0912FA8FD66DB35A" box="[266,360,1294,1318]" captionStart="FIGURE 12" captionStartId="26.[151,250,1177,1200]" captionTargetBox="[151,1435,445,1156]" captionTargetId="figure@26.[151,1436,445,1156]" captionTargetPageId="26" captionText="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." httpUri="https://zenodo.org/record/211746/files/figure.png" pageId="24" pageNumber="25">Figs. 12</figureCitation>
,
<figureCitation id="4CDD2A72FFA6FFBE0960FA8FD697B35A" box="[376,402,1294,1318]" captionStart="FIGURE 13" captionStartId="26.[151,250,1817,1840]" captionTargetBox="[151,1435,1280,1794]" captionTargetId="figure@26.[151,1436,1280,1796]" captionTargetPageId="26" captionText="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." httpUri="https://zenodo.org/record/211747/files/figure.png" pageId="24" pageNumber="25">13</figureCitation>
A) (
<emphasis id="E692EAE5FFA6FFBE09DCFA8ED50CB35A" box="[452,521,1295,1318]" italics="true" pageId="24" pageNumber="25">versus</emphasis>
a raised and enlarged series of scales extending posteriorly from the end of the spinose crest,
<figureCitation id="4CDD2A72FFA6FFBE0928FAB3D67BB336" box="[304,382,1330,1354]" captionStart="FIGURE 10" captionStartId="24.[151,250,1052,1075]" captionTargetBox="[212,1374,416,1016]" captionTargetId="figure@24.[205,1382,407,1030]" captionTargetPageId="24" captionText="FIGURE 10. Lectotype of Correlophus ciliatus (MNHN 701 A) showing the absence of lumbosacral tubercles and presence of heterogeneous dorsolateral scalation. Photo courtesy of Muséum National dHistoire Naturelle, Paris." httpUri="https://zenodo.org/record/211744/files/figure.png" pageId="24" pageNumber="25">Fig. 10</figureCitation>
), and (in three of the four
<typeStatus id="0B5D8855FFA6FFBE0A85FAB3D5DDB336" box="[669,728,1330,1354]" pageId="24" pageNumber="25">types</typeStatus>
) its possession of a series of small whitish tubercles on the lower back and/or tail base (
<figureCitation id="4CDD2A72FFA6FFBE099DFAD7D6DAB312" box="[389,479,1366,1390]" captionStart="FIGURE 12" captionStartId="26.[151,250,1177,1200]" captionTargetBox="[151,1435,445,1156]" captionTargetId="figure@26.[151,1436,445,1156]" captionTargetPageId="26" captionText="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." httpUri="https://zenodo.org/record/211746/files/figure.png" pageId="24" pageNumber="25">Figs. 12</figureCitation>
,
<figureCitation id="4CDD2A72FFA6FFBE09F3FAD7D503B312" box="[491,518,1366,1390]" captionStart="FIGURE 13" captionStartId="26.[151,250,1817,1840]" captionTargetBox="[151,1435,1280,1794]" captionTargetId="figure@26.[151,1436,1280,1796]" captionTargetPageId="26" captionText="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." httpUri="https://zenodo.org/record/211747/files/figure.png" pageId="24" pageNumber="25">13</figureCitation>
A, 14A, 15) (absent in
<taxonomicName id="13E64D74FFA6FFBE0AE7FAD6D46BB312" box="[767,878,1366,1390]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="25" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA6FFBE0AE7FAD6D46BB312" box="[767,878,1366,1390]" italics="true" pageId="24" pageNumber="25">C. ciliatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="9CFC657CFFA6FFBA08DFFAF8D698B051" lastPageId="28" lastPageNumber="29" pageId="24" pageNumber="25" type="description">
<paragraph id="D45936F7FFA6FFBF08DFFAF8D6FCB718" blockId="24.[151,1437,1150,2002]" lastBlockId="25.[151,1436,151,356]" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25">
<emphasis id="E692EAE5FFA6FFBE08DFFAF8D65CB3EE" bold="true" box="[199,345,1401,1426]" pageId="24" pageNumber="25">Description.</emphasis>
(data from adult male
<typeStatus id="0B5D8855FFA6FFBE0A73FAFBD5D5B3EE" box="[619,720,1402,1426]" pageId="24" pageNumber="25" type="holotype">holotype</typeStatus>
, MNHN 2011.1100). Specimen fixed with mouth open wide; abdominal incision for removal of liver sample for DNA. SVL
<quantity id="131E9B12FFA6FFBE0B8FFA1FD300B3CA" box="[919,1029,1438,1462]" metricMagnitude="-2" metricUnit="m" metricValue="9.559999999999999" pageId="24" pageNumber="25" unit="mm" value="95.6">95.6 mm</quantity>
; TailL 11.6 (broken with minimal regeneration); TrunkL
<quantity id="131E9B12FFA6FFBE0981FA43D504B3A6" box="[409,513,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="3.94" pageId="24" pageNumber="25" unit="mm" value="39.4">39.4 mm</quantity>
; HeadL
<quantity id="131E9B12FFA6FFBE0A7AFA43D5CEB3A6" box="[610,715,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="2.7399999999999998" pageId="24" pageNumber="25" unit="mm" value="27.4">27.4 mm</quantity>
; HeadW
<quantity id="131E9B12FFA6FFBE0B2DFA43D498B3A6" box="[821,925,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="2.16" pageId="24" pageNumber="25" unit="mm" value="21.6">21.6 mm</quantity>
; SnEye
<quantity id="131E9B12FFA6FFBE0BE3FA43D366B3A6" box="[1019,1123,1474,1498]" metricMagnitude="-2" metricUnit="m" metricValue="1.1199999999999999" pageId="24" pageNumber="25" unit="mm" value="11.2">11.2 mm</quantity>
; OrbD
<quantity id="131E9B12FFA6FFBE0CAFFA43D217B3A6" box="[1207,1298,1474,1498]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" pageId="24" pageNumber="25" unit="mm" value="7.1">7.1 mm</quantity>
; EyeEar
<quantity id="131E9B12FFA6FFBE0D61FA43D7C3B382" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="24" pageNumber="25" unit="mm" value="7.8">7.8 mm</quantity>
. Body moderately long (TrunkL = 41% SVL), slender, slightly depressed. Head triangular, large (HeadL = 29% SVL), very wide (HeadW = 79% HeadL), very well demarcated from neck (
<figureCitation id="4CDD2A72FFA6FFBE0C43F98BD3B6B05E" box="[1115,1203,1546,1570]" captionStart="FIGURE 12" captionStartId="26.[151,250,1177,1200]" captionTargetBox="[151,1435,445,1156]" captionTargetId="figure@26.[151,1436,445,1156]" captionTargetPageId="26" captionText="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." httpUri="https://zenodo.org/record/211746/files/figure.png" pageId="24" pageNumber="25">Fig. 12</figureCitation>
); nasofrontal region depressed, dorsal orbital rims raised; canthus distinct; snout relatively long (SnEye = 41% HeadL), much longer than eye diameter (OrbD = 64% SnEye). Scales on dorsum of snout approximately two to five times the diameter of those on occipital region, largest along canthus and immediately posterior to supranasals. Eye large (OrbD = 26% HeadL); pupil oval, margins faintly crenellated. Ear opening approximately two to three times longer than high, canted posterodorsally to anteroventrally at &lt;45° to the horizontal; eye to ear distance longer than diameter of eyes (EyeEar = 110% OrbD). Rostral rectangular, much broader (
<quantity id="131E9B12FFA6FFBE0BB9F963D4FFB086" box="[929,1018,1762,1786]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="24" pageNumber="25" unit="mm" value="4.1">4.1 mm</quantity>
) than high (
<quantity id="131E9B12FFA6FFBE0C9DF963D3E7B086" box="[1157,1250,1762,1786]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="24" pageNumber="25" unit="mm" value="2.2">2.2 mm</quantity>
), without rostral crease, contacted posteriorly by three internasals — median smaller and pentagonal, lateral ones larger and hexagonal — and two large supranasals, each somewhat larger than the lateral internasals; contacted posteroventrally by first supralabial. Nostrils oval, laterally oriented, surrounded by eight circumnarial scales, including enlarged supranasal, and broadly contacted by first supralabial. Mental subtriangular, deeper (
<quantity id="131E9B12FFA6FFBE0D22F8F3D291B1F6" box="[1338,1428,1906,1930]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="24" pageNumber="25" unit="mm" value="2.3">2.3 mm</quantity>
) than broad (
<quantity id="131E9B12FFA6FFBE0938F817D679B1D2" box="[288,380,1942,1966]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="24" pageNumber="25" unit="mm" value="1.7">1.7 mm</quantity>
). Two somewhat enlarged (68 times size of throat granules) postmental scales separate first infralabials from one another. Postmentals bordered posterolaterally and posteriorly by series of five smaller (~50% size of postmentals) scales. Scales in two to five rows medial to infralabials somewhat enlarged and elongate. 13(left)15(right) enlarged supralabial scales, posteriormost only about 3 times size of rictal scales, 10 (left) or 12 (right) supralabials to midpoint of orbit; 12 (left)13 (right) enlarged infralabial scales (right infralabials 811 fragmented into two rows of scales); 38 scale rows between supraciliaries, 18 scale rows across frontal bones at midpoint of orbit. Supraciliary scales forming 17 (L) and 18 (R) large pointed spines, smaller anteriorly, largest posterodorsally and posteriorly.
</paragraph>
<caption id="8099667FFFA7FFBF088FFACCD36EB3DC" httpUri="https://zenodo.org/record/211745/files/figure.png" pageId="25" pageNumber="26" targetBox="[160,1426,418,1328]" targetPageId="25">
<paragraph id="D45936F7FFA7FFBF088FFACCD36EB3DC" blockId="25.[151,1437,1357,1440]" pageId="25" pageNumber="26">
<emphasis id="E692EAE5FFA7FFBF088FFACCD624B31E" bold="true" box="[151,289,1357,1380]" pageId="25" pageNumber="26">FIGURE 11.</emphasis>
Distribution map of
<taxonomicName id="13E64D74FFA7FFBF0A18FACCD5D6B31F" box="[512,723,1357,1379]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA7FFBF0A18FACCD5D6B31F" box="[512,723,1357,1379]" italics="true" pageId="25" pageNumber="26">Correlophus ciliatus</emphasis>
</taxonomicName>
(circles) and
<taxonomicName id="13E64D74FFA7FFBF0B7DFACFD4F5B31F" box="[869,1008,1357,1379]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA7FFBF0B7DFACFD4F5B31F" box="[869,1008,1357,1379]" italics="true" pageId="25" pageNumber="26">C. belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA7FFBF0BE0FACFD349B31F" box="[1016,1100,1358,1379]" pageId="25" pageNumber="26" rank="species">
<emphasis id="E692EAE5FFA7FFBF0BE0FACFD349B31F" bold="true" box="[1016,1100,1358,1379]" pageId="25" pageNumber="26">sp. nov.</emphasis>
</taxonomicNameLabel>
(red star). For
<taxonomicName id="13E64D74FFA7FFBF0CE8FACFD25EB31F" box="[1264,1371,1357,1379]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="26" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA7FFBF0CE8FACFD25EB31F" box="[1264,1371,1357,1379]" italics="true" pageId="25" pageNumber="26">C. ciliatus</emphasis>
</taxonomicName>
green circles symbols represent vouchered records, orange circles represent unvouchered sight or literature records. Vouchered records from the Île des Pins lack precise localities. See Appendix for a list of localities mapped.
</paragraph>
</caption>
<paragraph id="D45936F7FFA7FFBF08DFFA4CD3C4B179" blockId="25.[151,1437,1485,2013]" pageId="25" pageNumber="26">
Dorsal scales small, mostly homogeneous (
<figureCitation id="4CDD2A72FFA7FFBF0AA9FA4CD40AB399" box="[689,783,1485,1509]" captionStart="FIGURE 12" captionStartId="26.[151,250,1177,1200]" captionTargetBox="[151,1435,445,1156]" captionTargetId="figure@26.[151,1436,445,1156]" captionTargetPageId="26" captionText="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." httpUri="https://zenodo.org/record/211746/files/figure.png" pageId="25" pageNumber="26">Figs. 12</figureCitation>
,
<figureCitation id="4CDD2A72FFA7FFBF0B04FA4CD433B399" box="[796,822,1485,1509]" captionStart="FIGURE 13" captionStartId="26.[151,250,1817,1840]" captionTargetBox="[151,1435,1280,1794]" captionTargetId="figure@26.[151,1436,1280,1796]" captionTargetPageId="26" captionText="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." httpUri="https://zenodo.org/record/211747/files/figure.png" pageId="25" pageNumber="26">13</figureCitation>
A), conical; ventral scales 1.32 times size of dorsals, smooth, flattened, oval, juxtaposed to subimbricate, enlarged under sternum, midventrally on posterior abdomen, and in precloacal region. Sternal and posterior abdominal scales rounded. Approximately 154 scale rows around mid-body. A dorsolateral fold of skin from anterior to ear to approximately level of adpressed elbow, strongly projecting above ear and on neck, continuing posteriorly as a slightly raised crest. Entire fold/crest bearing elongate spinose scales, becoming smaller above shoulder; a few small, scattered spinose scales between anterior margin of fold and posteriormost spinose supraciliaries. Lumbosacral region with six scattered, enlarged, semi-erect conical white tubercles, with their apices directed posteriorly or posterolaterally. Another two such tubercles at tail base, followed posteriorly by several slightly enlarged and irregularly-shaped non-tuberculate scales.
</paragraph>
<paragraph id="D45936F7FFA7FFBF08DFF890D298B1A1" blockId="25.[151,1437,1485,2013]" pageId="25" pageNumber="26">
Skin folds small but distinct on ventrolateral trunk between limb insertions, on ventrolateral border of neck, and on posterior margins of forelimbs. Well developed folds along postaxial border of hindlimbs, forming deep popliteal pockets behind the knees (
<figureCitation id="4CDD2A72FFA7FFBF0A2AF8D8D583B10D" box="[562,646,1881,1905]" captionStart="FIGURE 13" captionStartId="26.[151,250,1817,1840]" captionTargetBox="[151,1435,1280,1794]" captionTargetId="figure@26.[151,1436,1280,1796]" captionTargetPageId="26" captionText="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." httpUri="https://zenodo.org/record/211747/files/figure.png" pageId="25" pageNumber="26">Fig. 13</figureCitation>
B). Scales on postaxial hindlimb fold as far as base of digit V of pes with scattered, white, conical to weakly spinose tubercles. Scales on limbs granular to low and conical, subimbricate to juxtaposed. Scales on palms and soles smooth, flattened. Fore- and hindlimbs, exclusive of skin folds, moderately long and slender (ForeaL = 14% SVL; CrusL = 18% SVL), axillary pocket weakly developed.
</paragraph>
<paragraph id="D45936F7FFA4FFBC088FFF16D3F3B7FB" blockId="26.[151,1437,151,391]" pageId="26" pageNumber="27">
Digits moderately long, all bearing claws, those on digit I of both manus and pes reduced and partially sheathed, remaining claws long and strongly recurved; relative length of digits of manus: IV&gt;III&gt;II&gt;V&gt;I, and of pes: IV&gt;III&gt;V&gt;II&gt;I; digits well webbed; digits III and IV of pes tightly bound along length of elongate metatarsals. Subdigital lamellae all unpaired, somewhat bowed, with lateral margins gently angled distally (except for proximalmost lamellae, which are straight). Claw of digit I of manus and pes, lies between a smaller lateral and a larger (twice size of lateral) medial apical scansor. Lamellar counts from right (and left) sides
<date id="A0581037FFA4FFBC0CADFECDD247B718" box="[1205,1346,332,356]" pageId="26" pageNumber="27" value="1918-10-13" valueMax="1918-10-17" valueMin="1918-10-13">10-13-17-18</date>
-15 (
<date id="A0581037FFA4FFBC0D6EFECDD7FBB7FB" pageId="26" pageNumber="27" value="1917-10-15" valueMax="1917-10-18" valueMin="1917-10-15">10- 15-18-17</date>
-15) manus and
<date id="A0581037FFA4FFBC09ABFEEED545B7FB" box="[435,576,367,391]" pageId="26" pageNumber="27" value="1920-12-15" valueMax="1920-12-17" valueMin="1920-12-15">12-15-17-20</date>
-18 (
<date id="A0581037FFA4FFBC0A6BFEEED404B7FB" box="[627,769,367,391]" pageId="26" pageNumber="27" value="1919-12-14" valueMax="1919-12-18" valueMin="1919-12-14">12-14-18-19</date>
-17) pes (excludes apical scansors of digit I).
</paragraph>
<caption id="8099667FFFA4FFBC088FFB18D610B2B2" httpUri="https://zenodo.org/record/211746/files/figure.png" pageId="26" pageNumber="27" targetBox="[151,1435,445,1156]" targetPageId="26">
<paragraph id="D45936F7FFA4FFBC088FFB18D610B2B2" blockId="26.[151,1436,1177,1230]" pageId="26" pageNumber="27">
<emphasis id="E692EAE5FFA4FFBC088FFB18D624B2D3" bold="true" box="[151,289,1177,1200]" pageId="26" pageNumber="27">FIGURE 12.</emphasis>
Preserved holotype (MNHN 2011.1100) of
<taxonomicName id="13E64D74FFA4FFBC0AF2FB18D4DEB2D3" box="[746,987,1177,1199]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA4FFBC0AF2FB18D4DEB2D3" box="[746,987,1177,1199]" italics="true" pageId="26" pageNumber="27">Correlophus belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA4FFBC0BFBFB1BD330B2D3" box="[995,1077,1178,1199]" pageId="26" pageNumber="27" rank="species">
<emphasis id="E692EAE5FFA4FFBC0BFBFB1BD330B2D3" bold="true" box="[995,1077,1178,1199]" pageId="26" pageNumber="27">sp. nov.</emphasis>
</taxonomicNameLabel>
from Île Art, Îles Belep. Photo by A.M. Bauer.
</paragraph>
</caption>
<caption id="8099667FFFA4FFBC088FF898D610B1F7" httpUri="https://zenodo.org/record/211747/files/figure.png" pageId="26" pageNumber="27" targetBox="[151,1435,1280,1794]" targetPageId="26">
<paragraph id="D45936F7FFA4FFBC088FF898D610B1F7" blockId="26.[151,1436,1817,1931]" pageId="26" pageNumber="27">
<emphasis id="E692EAE5FFA4FFBC088FF898D627B153" bold="true" box="[151,290,1817,1840]" pageId="26" pageNumber="27">FIGURE 13.</emphasis>
Holotype (MNHN 2011.1100) of
<taxonomicName id="13E64D74FFA4FFBC0A9FF898D472B153" box="[647,887,1817,1839]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="26" pageNumber="27" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA4FFBC0A9FF898D472B153" box="[647,887,1817,1839]" italics="true" pageId="26" pageNumber="27">Correlophus belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA4FFBC0B99F89BD4D6B153" box="[897,979,1818,1839]" pageId="26" pageNumber="27" rank="species">
<emphasis id="E692EAE5FFA4FFBC0B99F89BD4D6B153" bold="true" box="[897,979,1818,1839]" pageId="26" pageNumber="27">sp. nov.</emphasis>
</taxonomicNameLabel>
showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A.M. Bauer.
</paragraph>
</caption>
<paragraph id="D45936F7FFA4FFBD08DFF839D219B68B" blockId="26.[151,1436,1976,2036]" lastBlockId="27.[151,1437,151,788]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
Small precloacal pores in a patch of somewhat enlarged scales, arranged in three rows (anterior to posterior), each divided by two small poreless scales medially, of 15(L) + 15(R), 12(L) + 10(R), and 3 (L) + 4(R). Pores restricted to proximal 40% of thighs (
<figureCitation id="4CDD2A72FFA5FFBD0A58FF16D597B6D3" box="[576,658,151,175]" captionStart="FIGURE 13" captionStartId="26.[151,250,1817,1840]" captionTargetBox="[151,1435,1280,1794]" captionTargetId="figure@26.[151,1436,1280,1796]" captionTargetPageId="26" captionText="FIGURE 13. Holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. showing (A) homogeneous dorsal scalation and (B) rudimentary regenerated tail, skin folds on posterior margin of thighs, and precloacal pores extending onto thighs (the three pore-bearing rows of scales are near the posterior margin of the grayish patch of scales anterior to the vent). Photos by A. M. Bauer." httpUri="https://zenodo.org/record/211747/files/figure.png" pageId="27" pageNumber="28">Fig. 13</figureCitation>
B). Hemipenial bulge large; a single large, smooth, flattened, conical cloacal spur comprising a much larger dorsal scale and a small posterolateral scale subtending it on each side of tail base. Tail broken at base and supporting a rudimentary regenerate of narrow diameter (see Discussion).
</paragraph>
<paragraph id="D45936F7FFA5FFBD08DFFE85D218B46B" blockId="27.[151,1437,151,788]" pageId="27" pageNumber="28">
Color in preservative (based on
<typeStatus id="0B5D8855FFA5FFBD0A34FE85D591B760" box="[556,660,260,284]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
): Dorsum, lateral surfaces of trunk, side of neck and head posterior to orbit, and limbs more-or-less a uniform dark chocolate brown. Snout, dorsum of head and middorsal region between dorsolateral spinose crests a lighter brown with a tinge of brick red (
<figureCitation id="4CDD2A72FFA5FFBD0C1EFECDD35EB718" box="[1030,1115,332,356]" captionStart="FIGURE 12" captionStartId="26.[151,250,1177,1200]" captionTargetBox="[151,1435,445,1156]" captionTargetId="figure@26.[151,1436,445,1156]" captionTargetPageId="26" captionText="FIGURE 12. Preserved holotype (MNHN 2011.1100) of Correlophus belepensis sp. nov. from Île Art, Îles Belep. Photo by A. M. Bauer." httpUri="https://zenodo.org/record/211746/files/figure.png" pageId="27" pageNumber="28">Fig. 12</figureCitation>
). Posterior infralabial scales with whitish ventral borders. Midvertebral region between end of spinose crests and white tubercles of lumbar region mottled with some hint of alternating lighter and darker brown bands. Venter light brown with darker mottling, especially on anterior chest, mid- to posterior abdomen, in precloacal region, and under hemipenial bulge. Soles and palms pale grayish brown. A white line along the postaxial margin of the hindlimb. Tail base bearing a cream “Y”-shaped marking dorsally; vestigial regenerated tail beige with dark pigment around its base.
</paragraph>
<paragraph id="D45936F7FFA5FFBD08DFFDA4D522B568" blockId="27.[151,1437,151,788]" pageId="27" pageNumber="28">
In life the dorsal coloration of the
<typeStatus id="0B5D8855FFA5FFBD0A52FDA4D580B441" box="[586,645,549,573]" pageId="27" pageNumber="28">types</typeStatus>
was an orangey-brown with a slightly greenish suffusion, with darker diffuse irregular reticulations on the flanks. Crown of the head, area between denticulate crests, and anterior middorsal region more distinctly orangey to russet than adjacent flanks, fading either abruptly or gradually posterior to the shoulders. Anterior of head with yellowish highlights; orbital rim and ventrolateral surfaces of supraciliary scales yellow. Iris silvery to whitish (
<figureCitation id="4CDD2A72FFA5FFBD0A53FD35D5A7B4B0" box="[587,674,692,716]" captionStart="FIGURE 14" captionStartId="27.[151,250,1318,1341]" captionTargetBox="[151,1436,841,1296]" captionTargetId="figure@27.[151,1436,841,1297]" captionTargetPageId="27" captionText="FIGURE 14. Life photographs of Correlophus belepensis sp. nov. (A) Male holotype (MNHN 2011.1100) and (B) smaller female paratype (CAS 250865). Photos by A. H. Whitaker." httpUri="https://zenodo.org/record/211748/files/figure.png" pageId="27" pageNumber="28">Fig. 14</figureCitation>
). Venter reddish-brown, with diffuse, darker gray-brown irregular markings, particularly along the ventrolateral margins and with scattered isolated greenish-yellow scales. Tongue and interior of mouth unpigmented.
</paragraph>
<caption id="8099667FFFA5FFBD088FFAA7D5E2B327" httpUri="https://zenodo.org/record/211748/files/figure.png" pageId="27" pageNumber="28" targetBox="[151,1436,841,1296]" targetPageId="27">
<paragraph id="D45936F7FFA5FFBD088FFAA7D5E2B327" blockId="27.[151,1435,1318,1371]" pageId="27" pageNumber="28">
<emphasis id="E692EAE5FFA5FFBD088FFAA7D627B347" bold="true" box="[151,290,1318,1341]" pageId="27" pageNumber="28">FIGURE 14.</emphasis>
Life photographs of
<taxonomicName id="13E64D74FFA5FFBD0A19FAA7D5F7B340" box="[513,754,1318,1340]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="27" pageNumber="28" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA5FFBD0A19FAA7D5F7B340" box="[513,754,1318,1340]" italics="true" pageId="27" pageNumber="28">Correlophus belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA5FFBD0AE3FAA6D448B340" box="[763,845,1319,1340]" pageId="27" pageNumber="28" rank="species">
<emphasis id="E692EAE5FFA5FFBD0AE3FAA6D448B340" bold="true" box="[763,845,1319,1340]" pageId="27" pageNumber="28">sp. nov.</emphasis>
</taxonomicNameLabel>
(A) Male holotype (MNHN 2011.1100) and (B) smaller female paratype (CAS 250865). Photos by A.H. Whitaker.
</paragraph>
</caption>
<paragraph id="D45936F7FFA5FFBD08DFFA2AD6BCB0C3" blockId="27.[151,1436,1451,2015]" pageId="27" pageNumber="28">
<emphasis id="E692EAE5FFA5FFBD08DFFA2AD646B3B8" bold="true" box="[199,323,1451,1476]" pageId="27" pageNumber="28">Osteology.</emphasis>
Vertebral counts are typical for diplodactylid geckos, with 26 presacral and 2 sacral vertebrae. The first three cervical vertebrae are without ribs, as is the last presacral (lumbar) vertebra. Also typical for the family, the caudal skeleton includes 5 pygal vertebrae. The
<typeStatus id="0B5D8855FFA5FFBD0AF8FA72D447B077" box="[736,834,1523,1547]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
and all
<typeStatus id="0B5D8855FFA5FFBD0B85FA75D30CB070" box="[925,1033,1524,1548]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
have the tail broken within the first postpygal vertebra. The phalangeal formulae of the manus and pes are unreduced, 2-3-4-5-3 and 2-3-4-5-4, respectively. Total tooth loci in upper jaw of
<typeStatus id="0B5D8855FFA5FFBD0A88F9BAD5F7B02F" box="[656,754,1595,1619]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
85, of which
<quantity id="131E9B12FFA5FFBD0B96F9BAD4BCB02F" box="[910,953,1595,1619]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" pageId="27" pageNumber="28" unit="in" value="9.0">9 in</quantity>
premaxilla; total mandibular tooth loci 78. A single pair of crescentic cloacal bones is present in the
<typeStatus id="0B5D8855FFA5FFBD0B0DF9DED472B00B" box="[789,887,1631,1655]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
and in the male
<typeStatus id="0B5D8855FFA5FFBD0C36F9E1D38BB004" box="[1070,1166,1632,1656]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
AMS R
<date id="A0581037FFA5FFBD0CF4F9DED246B00B" box="[1260,1347,1631,1655]" pageId="27" pageNumber="28" value="82-12-16">161282</date>
. In both female
<typeStatus id="0B5D8855FFA5FFBD08F1F905D650B0E0" box="[233,341,1668,1692]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
the endolymphatic system is visible in x-rays by its radio-opaque calcium content and can be seen to be entirely intracranial.
</paragraph>
<paragraph id="D45936F7FFA5FFBD08DFF94AD407B1A3" blockId="27.[151,1436,1451,2015]" pageId="27" pageNumber="28">
<emphasis id="E692EAE5FFA5FFBD08DFF94AD63EB098" bold="true" box="[199,315,1739,1764]" pageId="27" pageNumber="28">Variation.</emphasis>
Comparative mensural and meristic data for the
<typeStatus id="0B5D8855FFA5FFBD0B92F94AD4E9B09F" box="[906,1004,1739,1763]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
and
<typeStatus id="0B5D8855FFA5FFBD0C34F94DD392B098" box="[1068,1175,1740,1764]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
are given in
<tableCitation id="9964034CFFA5FFBD0D25F94AD29DB09F" box="[1341,1432,1739,1763]" captionStart="TABLE 2" captionStartId="28.[151,239,151,175]" captionText="TABLE 2. Mensural data for the types of Correlophus belepensis sp. nov. Abbreviations as in Materials and methods. All measurements in mm. * All tails broken with rudimentary regenerated tips." pageId="27" pageNumber="28">Table 2</tableCitation>
. Meristic characters of
<typeStatus id="0B5D8855FFA5FFBD0981F971D501B174" box="[409,516,1776,1800]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
are mostly similar to those of the
<typeStatus id="0B5D8855FFA5FFBD0B8DF96ED4FFB17B" box="[917,1018,1775,1799]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
, and are mentioned hereafter if only they differ. All
<typeStatus id="0B5D8855FFA5FFBD0953F895D6B2B150" box="[331,439,1812,1836]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
had only a single postmental scale. AMS R
<date id="A0581037FFA5FFBD0BAFF892D30EB157" box="[951,1035,1811,1835]" pageId="27" pageNumber="28" value="83-12-16">161283</date>
had seven circumnarial scales and CAS
<date id="A0581037FFA5FFBD08CEF8B6D62FB133" box="[214,298,1847,1871]" pageId="27" pageNumber="28" value="65-08-25">250865</date>
had seven around the left naris and eight around the right. Precloacal pores are arranged in rows of 18(L) + 15(R), 10(L) + 10(R), and 3(L) + 2(R) in the male
<typeStatus id="0B5D8855FFA5FFBD0B2BF8DDD496B108" box="[819,915,1884,1908]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
(AMS R
<date id="A0581037FFA5FFBD0BE4F8DAD354B10F" box="[1020,1105,1883,1907]" pageId="27" pageNumber="28" value="82-12-16">161282</date>
) and are absent in the female
<typeStatus id="0B5D8855FFA5FFBD088FF801D603B1E4" box="[151,262,1920,1944]" pageId="27" pageNumber="28" type="paratype">paratypes</typeStatus>
. In the male
<typeStatus id="0B5D8855FFA5FFBD09B2F801D50FB1E4" box="[426,522,1920,1944]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
the posterolateral scale of the cloacal spur is separate from the larger scale, resulting in two discrete cloacal spurs on each side of the tail base. In the females the cloacal spurs are represented by one to three slightly enlarged smooth conical scales.
</paragraph>
<caption id="8099667FFFA2FFBA088FFF16D4E4B6B3" pageId="28" pageNumber="29">
<paragraph id="D45936F7FFA2FFBA088FFF16D4E4B6B3" blockId="28.[151,1436,151,207]" pageId="28" pageNumber="29">
<emphasis id="E692EAE5FFA2FFBA088FFF16D60EB6D3" bold="true" box="[151,267,151,175]" pageId="28" pageNumber="29">TABLE 2.</emphasis>
Mensural data for the types of
<taxonomicName id="13E64D74FFA2FFBA0A7AFF19D45AB6D3" box="[610,863,152,175]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA2FFBA0A7AFF19D45AB6D3" box="[610,863,152,175]" italics="true" pageId="28" pageNumber="29">Correlophus belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA2FFBA0B7FFF16D4BBB6D3" box="[871,958,151,175]" pageId="28" pageNumber="29" rank="species">
<emphasis id="E692EAE5FFA2FFBA0B7FFF16D4B2B6D3" bold="true" box="[871,951,151,175]" pageId="28" pageNumber="29">sp. nov</emphasis>
.
</taxonomicNameLabel>
Abbreviations as in Materials and methods. All measurements in mm. * All tails broken with rudimentary regenerated tips.
</paragraph>
</caption>
<paragraph id="D45936F7FFA2FFBA09B5FF70D249B74E" blockId="28.[164,1410,240,1240]" pageId="28" pageNumber="29">
MNHN 2011.1100 AMS R
<date id="A0581037FFA2FFBA0B08FF71D459B77A" box="[784,860,240,262]" pageId="28" pageNumber="29" value="82-12-16">161282</date>
AMS R
<date id="A0581037FFA2FFBA0C07FF71D36EB77A" box="[1055,1131,240,262]" pageId="28" pageNumber="29" value="83-12-16">161283</date>
CAS
<date id="A0581037FFA2FFBA0D0EFF71D267B77A" box="[1302,1378,240,262]" pageId="28" pageNumber="29" value="65-08-25">250865</date>
<typeStatus id="0B5D8855FFA2FFBA09F8FE9DD53CB74E" box="[480,569,284,306]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
<typeStatus id="0B5D8855FFA2FFBA0AFCFE9CD249B74E" box="[740,1356,285,306]" pageId="28" pageNumber="29" type="paratype">paratype paratype paratype</typeStatus>
</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFEC8D246B722" blockId="28.[164,1410,240,1240]" box="[164,1347,328,350]" pageId="28" pageNumber="29">Sex male male female female</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFEF5D233B7F5" blockId="28.[164,1410,240,1240]" box="[164,1334,372,394]" pageId="28" pageNumber="29">SVL 95.6 88.6 99.7 99.4</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFE20D233B7CA" blockId="28.[164,1410,240,1240]" box="[164,1334,417,438]" pageId="28" pageNumber="29">ForeaL 13.4 12.1 13.0 12.3</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFE4CD233B79E" blockId="28.[164,1410,240,1240]" box="[164,1334,460,482]" pageId="28" pageNumber="29">CrusL 17.2 15.3 16.7 16.4</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFE79D232B472" blockId="28.[164,1410,240,1240]" box="[164,1335,504,526]" pageId="28" pageNumber="29">TailL* 11.6 12.9 15.0 16.4</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFDA5D233B446" blockId="28.[164,1410,240,1240]" box="[164,1334,548,570]" pageId="28" pageNumber="29">TrunkL 39.4 34.5 40.6 41.0</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFDD1D233B41A" blockId="28.[164,1410,240,1240]" box="[164,1334,592,614]" pageId="28" pageNumber="29">HeadL 27.4 25.4 29.6 27.8</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFDFDD233B4EE" blockId="28.[164,1410,240,1240]" box="[164,1334,636,658]" pageId="28" pageNumber="29">HeadW 21.6 19.8 22.4 22.0</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFD29D235B4C2" blockId="28.[164,1410,240,1240]" box="[164,1328,680,702]" pageId="28" pageNumber="29">OrbD 7.1 6.8 7.6 7.0</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFD54D235B496" blockId="28.[164,1410,240,1240]" box="[164,1328,724,746]" pageId="28" pageNumber="29">EyeEar 7.8 7.5 8.8 8.0</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFC80D233B56A" blockId="28.[164,1410,240,1240]" box="[164,1334,769,790]" pageId="28" pageNumber="29">SnEye 11.2 11.1 11.8 11.7</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFCACD235B53E" blockId="28.[164,1410,240,1240]" box="[164,1328,812,834]" pageId="28" pageNumber="29">NarEye 8.0 8.2 8.5 8.5</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFCD9D233B512" blockId="28.[164,1410,240,1240]" box="[164,1334,856,878]" pageId="28" pageNumber="29">Interorb 11.3 10.3 13.2 13.9</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFC04D24BB5E6" blockId="28.[164,1410,240,1240]" box="[164,1358,900,922]" pageId="28" pageNumber="29">EarL 2.2 1.3 1.9 occluded</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFC30D235B5BA" blockId="28.[164,1410,240,1240]" box="[164,1328,945,966]" pageId="28" pageNumber="29">Internar 3.3 3.2 3.4 3.7</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFC5CD26FB58F" blockId="28.[164,1410,240,1240]" box="[164,1386,988,1011]" pageId="28" pageNumber="29">SupraL L/R 13(10)/15(12) 15(12)/ 16(13) 14(11)/ 14(11) 15(12)/ 14(11)</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFB89D238B262" blockId="28.[164,1410,240,1240]" box="[164,1341,1032,1054]" pageId="28" pageNumber="29">
InfraL L/R
<date id="A0581037FFA2FFBA09E8FB88D40FB262" box="[496,778,1033,1054]" pageId="28" pageNumber="29" value="1913-12-13">12/13 13</date>
/13 13/14 13/13
</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFBB4D265B207" blockId="28.[164,1410,240,1240]" pageId="28" pageNumber="29">
LamManus L/R
<date id="A0581037FFA2FFBA09B0FBB5D523B236" box="[424,550,1076,1098]" pageId="28" pageNumber="29" value="1917-10-15" valueMax="1917-10-18" valueMin="1917-10-15">10-15-18-17</date>
-15/
<date id="A0581037FFA2FFBA0A57FBB5D5F6B236" box="[591,755,1076,1098]" pageId="28" pageNumber="29" value="1916-10-14" valueMax="1916-10-15" valueMin="1916-10-14">10- 14-15-16</date>
-19-15/
<date id="A0581037FFA2FFBA0B5CFBB4D4F4B236" box="[836,1009,1076,1098]" pageId="28" pageNumber="29" value="1914-11-13" valueMax="1914-11-14" valueMin="1914-11-13">13-14- 11-14</date>
-17-19-15/
<date id="A0581037FFA2FFBA0C7CFBB4D209B236" box="[1124,1292,1077,1098]" pageId="28" pageNumber="29" value="1917-10-09" valueMax="1917-10-13" valueMin="1917-10-09">10- 9-13-17</date>
-20-14/
<date id="A0581037FFA2FFBA0D78FBB4D52EB207" pageId="28" pageNumber="29" value="1918-11-13" valueMax="1918-11-17" valueMin="1918-11-13">11- 13-17-18</date>
-15 16-19-15 14-18-18-15 13-16-17-15
</paragraph>
<paragraph id="D45936F7FFA2FFBA08BCFB13D698B051" blockId="28.[164,1410,240,1240]" lastBlockId="28.[151,1437,1305,2013]" pageId="28" pageNumber="29">
LamPes L/R
<date id="A0581037FFA2FFBA09B0FB13D523B2DB" box="[424,550,1170,1191]" pageId="28" pageNumber="29" value="1919-12-14" valueMax="1919-12-18" valueMin="1919-12-14">12-14-18-19</date>
-17/
<date id="A0581037FFA2FFBA0A57FB13D5F7B2DB" box="[591,754,1169,1191]" pageId="28" pageNumber="29" value="1918-12-13" valueMax="1918-12-16" valueMin="1918-12-13">12- 13-16-18</date>
-19-18/
<date id="A0581037FFA2FFBA0B5CFB10D4F4B2DB" box="[836,1009,1169,1191]" pageId="28" pageNumber="29" value="1913-10-13" valueMax="1913-10-15" valueMin="1913-10-13">13-15- 10-13</date>
-16-19-17/
<date id="A0581037FFA2FFBA0C7CFB13D20BB2DB" box="[1124,1294,1170,1191]" pageId="28" pageNumber="29" value="1919-10-08" valueMax="1919-10-14" valueMin="1919-10-08">10- 8-14-19</date>
-19-17/
<date id="A0581037FFA2FFBA0D78FB13D529B2A4" pageId="28" pageNumber="29" value="1920-10-15" valueMax="1920-10-17" valueMin="1920-10-15">10- 15-17-20</date>
-18 17-21-17 15-19-21-17 15-18-21-17 Color pattern is generally similar among all specimens examined, although in AMS R
<date id="A0581037FFA2FFBA0C9AFA98D3D3B34D" box="[1154,1238,1305,1329]" pageId="28" pageNumber="29" value="82-12-16">161282</date>
and CAS
<date id="A0581037FFA2FFBA0D50FA98D299B34D" box="[1352,1436,1305,1329]" pageId="28" pageNumber="29" value="65-08-25">250865</date>
the dark chocolate dorsal and lateral coloration is more mottled and less uniform than in the
<typeStatus id="0B5D8855FFA2FFBA0CBBFABCD20CB329" box="[1187,1289,1341,1365]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
. The number of white tubercles on the dorsum is also variable (
<figureCitation id="4CDD2A72FFA2FFBA0AD6FAE0D427B305" box="[718,802,1377,1401]" captionStart="FIGURE 15" captionStartId="29.[151,250,1469,1492]" captionTargetBox="[217,1369,733,1447]" captionTargetId="figure@29.[217,1369,733,1448]" captionTargetPageId="29" captionText="FIGURE 15. Type series of Correlophus belepensis sp. nov. showing variation in color and in lumbosacral tubercles. Left to right: holotype (MNHN 2011.1100), paratypes (AMS R 161282, AMS R 161283, CAS 250865). Photo by A. H. Whitaker." httpUri="https://zenodo.org/record/211749/files/figure.png" pageId="28" pageNumber="29">Fig. 15</figureCitation>
). AMS R
<date id="A0581037FFA2FFBA0B97FAE0D4E6B305" box="[911,995,1377,1401]" pageId="28" pageNumber="29" value="83-12-16">161283</date>
has three tubercles on the lumbosacral region, two to the left of the midline and one to the right. In AMS R
<date id="A0581037FFA2FFBA0B81FA04D4E8B3E1" box="[921,1005,1413,1437]" pageId="28" pageNumber="29" value="82-12-16">161282</date>
there is a single such scale anterior to the hindlimb insertion and left of the dorsal midline and several irregular scales, including one small white tubercle at the tail base, but neither tubercle is conical as in the
<typeStatus id="0B5D8855FFA2FFBA0B24FA4CD4A7B399" box="[828,930,1485,1509]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
. In CAS
<date id="A0581037FFA2FFBA0C00FA4CD369B399" box="[1048,1132,1485,1509]" pageId="28" pageNumber="29" value="65-08-25">250865</date>
there are no lumbosacral tubercles. Venter in life yellowish (CAS
<date id="A0581037FFA2FFBA0A78FA70D5B3B075" box="[608,694,1521,1545]" pageId="28" pageNumber="29" value="65-08-25">250865</date>
) to reddish-brown; females lacking the scattered greenish-yellow ventral scales of males.
</paragraph>
</subSubSection>
<subSubSection id="9CFC657CFFA2FFBA08DFF9BBD23FB02D" box="[199,1338,1593,1618]" pageId="28" pageNumber="29" type="etymology">
<paragraph id="D45936F7FFA2FFBA08DFF9BBD23FB02D" blockId="28.[151,1437,1305,2013]" box="[199,1338,1593,1618]" pageId="28" pageNumber="29">
<emphasis id="E692EAE5FFA2FFBA08DFF9BBD64BB02E" bold="true" box="[199,334,1594,1618]" pageId="28" pageNumber="29">Etymology.</emphasis>
The specific epithet refers to the Îles Belep, to which this species is apparently restricted.
</paragraph>
</subSubSection>
<subSubSection id="9CFC657CFFA2FFB808DFF9DCD50AB197" lastPageId="30" lastPageNumber="31" pageId="28" pageNumber="29" type="distribution">
<paragraph id="D45936F7FFA2FFBA08DFF9DCD3FEB1E9" blockId="28.[151,1437,1305,2013]" pageId="28" pageNumber="29">
<emphasis id="E692EAE5FFA2FFBA08DFF9DCD665B00A" bold="true" box="[199,352,1629,1654]" pageId="28" pageNumber="29">Distribution.</emphasis>
<taxonomicName id="13E64D74FFA2FFBA0970F9DFD577B00A" box="[360,626,1630,1654]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA2FFBA0970F9DFD577B00A" box="[360,626,1630,1654]" italics="true" pageId="28" pageNumber="29">Correlophus belepensis</emphasis>
</taxonomicName>
has been recorded only from Île Art, in the Îles Belep group more than
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north-west of Grande Terre (
<bibRefCitation id="B0774B06FFA2FFBA0A31F900D40AB0E5" author="Whitaker" box="[553,783,1665,1690]" pageId="28" pageNumber="29" refString="Whitaker, A. H., Sadlier, R. A., Bauer, A. M. &amp; Whitaker, V. A. (2004) Biodiversity and conservation status of lizards in threatened and restricted habitats of north-western New Caledonia. Unpublished report by Whitaker Consultants Limited to Direction du Developpement Economique et de l'Environnement, Province Nord, Kone, New Caledonia, viii + 106 pp." type="book" year="2004">
Whitaker
<emphasis id="E692EAE5FFA2FFBA0A83F902D5C8B0E6" box="[667,717,1666,1690]" italics="true" pageId="28" pageNumber="29">et al</emphasis>
2004
</bibRefCitation>
;
<figureCitation id="4CDD2A72FFA2FFBA0B04F900D468B0E5" box="[796,877,1665,1689]" captionStart="FIGURE 11" captionStartId="25.[151,250,1357,1380]" captionTargetBox="[160,1426,418,1328]" captionTargetId="figure@25.[151,1436,409,1336]" captionTargetPageId="25" captionText="FIGURE 11. Distribution map of Correlophus ciliatus (circles) and C. belepensis sp. nov. (red star). For C. ciliatus green circles symbols represent vouchered records, orange circles represent unvouchered sight or literature records. Vouchered records from the Île des Pins lack precise localities. See Appendix for a list of localities mapped." httpUri="https://zenodo.org/record/211745/files/figure.png" pageId="28" pageNumber="29">Fig. 11</figureCitation>
)). Two discrete sub-populations occur on Île Art,
<quantity id="131E9B12FFA2FFBA088FF924D7E0B0C1" box="[151,229,1701,1725]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="28" pageNumber="29" unit="km" value="4.5">4.5 km</quantity>
apart, one on the Tolé Munu plateau (c.700 ha) and the other on the Kalidan plateau (c. 220 ha). The
<typeStatus id="0B5D8855FFA2FFBA0D74F927D299B0C2" box="[1388,1436,1702,1726]" pageId="28" pageNumber="29">type</typeStatus>
series was collected on the former; specimens were observed, but not collected, at Puröbi (
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,
<geoCoordinate id="B1D25030FFA2FFBA088FF96CD63BB179" box="[151,318,1773,1797]" direction="east" orientation="longitude" pageId="28" pageNumber="29" precision="1" value="163.67473">163°40'29.0&quot;E</geoCoordinate>
,
<quantity id="131E9B12FFA2FFBA0949F96CD6A4B179" box="[337,417,1773,1797]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="28" pageNumber="29" unit="m" value="230.0">230 m</quantity>
) on the latter. These plateaux comprise an undulating ultramafic substrate between
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in elevation that is capped by a rocky cuirasse surface covered with closed forest or paraforestier. They are separated by a wide, low saddle (&lt;
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) with sparse fire-induced savannah woodland and grassland. A small area (c.90 ha) of similar habitat to that on the Tolé Munu and Kalidan plateaux occurs at lower elevation (
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) on Île Pott, the only other large island in the Belep group, but it has not been surveyed.
</paragraph>
<paragraph id="D45936F7FFA2FFBB08DFF823D30CB4DB" blockId="28.[151,1437,1305,2013]" lastBlockId="29.[151,1437,151,680]" lastPageId="29" lastPageNumber="30" pageId="28" pageNumber="29">
<emphasis id="E692EAE5FFA2FFBA08DFF823D694B1C6" bold="true" box="[199,401,1953,1978]" pageId="28" pageNumber="29">Natural History.</emphasis>
<taxonomicName id="13E64D74FFA2FFBA0985F823D5AFB1C6" box="[413,682,1954,1978]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="28" pageNumber="29" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA2FFBA0985F823D5AFB1C6" box="[413,682,1954,1978]" italics="true" pageId="28" pageNumber="29">Correlophus belepensis</emphasis>
</taxonomicName>
has only been found in old-growth, low-stature (&lt;
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) closed humidforest and paraforestier habitat (&lt;
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) on bouldery cuirasse surfaces (
<figureCitation id="4CDD2A72FFA2FFBA0BF5F844D344B1A1" box="[1005,1089,1989,2013]" captionStart="FIGURE 16" captionStartId="30.[151,250,861,884]" captionTargetBox="[337,1249,229,840]" captionTargetId="figure@30.[337,1249,229,840]" captionTargetPageId="30" captionText="FIGURE 16. Habitat of Correlophus belepensis sp. nov. and Mniarogekko jalu sp. nov. on the Tolé Munu plateau, Île Art, Îles Belep, Province Nord, New Caledonia. Photo by A. H. Whitaker." httpUri="https://zenodo.org/record/211750/files/figure.png" pageId="28" pageNumber="29">Fig. 16</figureCitation>
). The Tolé Munu plateau has a diverse flora (&gt;170 species) including several rare and threatened taxa, several of which are endemic to that site (
<bibRefCitation id="B0774B06FFA3FFBB0886FF3DD6A4B6A8" author="Munzinger" box="[158,417,188,212]" pageId="29" pageNumber="30" refString="Munzinger, J., Barrabe, L. &amp; Swenson, U. (2009) Compte-rendu de mission sur l'ile Art, 24 - 27 Aout 2009. Unpublished report, Laboratoire de Botanique et d'Ecologie Vegetale Appliquees, IRD, Noumea, 6 pp." type="book chapter" year="2009">
Munzinger
<emphasis id="E692EAE5FFA3FFBB093BFF3CD650B6A8" box="[291,341,188,212]" italics="true" pageId="29" pageNumber="30">et al</emphasis>
. 2009
</bibRefCitation>
). Dominant canopy trees in this forest, with large trunks with holes and crevices as retreat sites for geckos, are
<taxonomicName id="13E64D74FFA3FFBB0990FF60D588B68B" authority="G. Forst." authorityName="G. Forst." box="[392,653,223,248]" class="Gastropoda" family="Ficidae" genus="Ficus" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="prolixa">
<emphasis id="E692EAE5FFA3FFBB0990FF60D525B684" box="[392,544,224,248]" italics="true" pageId="29" pageNumber="30">Ficus prolixa</emphasis>
G. Forst.
</taxonomicName>
(Fam.
<taxonomicName id="13E64D74FFA3FFBB0AFBFF5ED453B68B" box="[739,854,223,247]" class="Magnoliopsida" family="Moraceae" kingdom="Plantae" order="Rosales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Moraceae</taxonomicName>
),
<taxonomicName id="13E64D74FFA3FFBB0B73FF61D38AB68B" authority="Baill." authorityName="Baill." box="[875,1167,223,248]" class="Magnoliopsida" family="Euphorbiaceae" genus="Alphandia" kingdom="Plantae" order="Malpighiales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="resinosa">
<emphasis id="E692EAE5FFA3FFBB0B73FF61D34DB684" box="[875,1096,224,248]" italics="true" pageId="29" pageNumber="30">Alphandia resinosa</emphasis>
Baill.
</taxonomicName>
(Fam.
<taxonomicName id="13E64D74FFA3FFBB0CFDFF5ED295B68B" box="[1253,1424,223,247]" class="Magnoliopsida" family="Euphorbiaceae" kingdom="Plantae" order="Malpighiales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Euphorbiaceae</taxonomicName>
),
<taxonomicName id="13E64D74FFA3FFBB088FFE85D7F1B760" box="[151,244,260,284]" class="Magnoliopsida" family="Sapotaceae" genus="Iteiluma" higherTaxonomySource="CoL" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="E692EAE5FFA3FFBB088FFE85D7F1B760" box="[151,244,260,284]" italics="true" pageId="29" pageNumber="30">Iteiluma</emphasis>
</taxonomicName>
sp. (Fam.
<taxonomicName id="13E64D74FFA3FFBB096FFE85D6FCB760" box="[375,505,260,284]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
),
<taxonomicName id="13E64D74FFA3FFBB0A08FE85D33DB760" authority="(Pancher &amp; Sebert) Pierre" authorityName="(Pancher &amp; Sebert) Pierre" box="[528,1080,260,284]" class="Magnoliopsida" family="Sapotaceae" genus="Planchonella" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="wakere">
<emphasis id="E692EAE5FFA3FFBB0A08FE85D405B760" box="[528,768,260,284]" italics="true" pageId="29" pageNumber="30">Planchonella wakere</emphasis>
(Pancher &amp; Sebert) Pierre
</taxonomicName>
(Fam.
<taxonomicName id="13E64D74FFA3FFBB0C88FE85D216B760" box="[1168,1299,260,284]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
),
<taxonomicName id="13E64D74FFA3FFBB0D31FE84D7D9B73C" class="Magnoliopsida" family="Sapotaceae" genus="Mimusops" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="elengi">
<emphasis id="E692EAE5FFA3FFBB0D31FE84D7D9B73C" italics="true" pageId="29" pageNumber="30">Mimusops elengi</emphasis>
</taxonomicName>
L. (Fam.
<taxonomicName id="13E64D74FFA3FFBB0941FEA6D6D9B743" box="[345,476,295,319]" class="Magnoliopsida" family="Sapotaceae" kingdom="Plantae" order="Ericales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Sapotaceae</taxonomicName>
), and
<taxonomicName id="13E64D74FFA3FFBB0A3FFEA9D4DBB743" authority="Brongn. &amp; Gris" authorityName="Brongn. &amp; Gris" box="[551,990,295,320]" class="Magnoliopsida" family="Myrtaceae" genus="Piliocalyx" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="species" species="laurifolius">
<emphasis id="E692EAE5FFA3FFBB0A3FFEA9D41FB73C" box="[551,794,296,320]" italics="true" pageId="29" pageNumber="30">Piliocalyx laurifolius</emphasis>
Brongn. &amp;
<collectingCountry id="ACF17667FFA3FFBB0BB6FEA6D4DBB743" box="[942,990,295,319]" name="Greece" pageId="29" pageNumber="30">Gris</collectingCountry>
</taxonomicName>
and
<taxonomicName id="13E64D74FFA3FFBB0C06FEA9D38AB73C" box="[1054,1167,296,320]" class="Magnoliopsida" family="Myrtaceae" genus="Piliocalyx" higherTaxonomySource="GBIF,CoL" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="genus">
<emphasis id="E692EAE5FFA3FFBB0C06FEA9D38AB73C" box="[1054,1167,296,320]" italics="true" pageId="29" pageNumber="30">Piliocalyx</emphasis>
</taxonomicName>
sp. (Fam.
<taxonomicName id="13E64D74FFA3FFBB0D0EFEA6D294B743" box="[1302,1425,295,319]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="29" pageNumber="30" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
). Fortuitously, browsing ungulates and pigs are absent from the Belep Islands so this forest is in reasonably good condition with thick undergrowth. All observations of
<taxonomicName id="13E64D74FFA3FFBB0AE3FEF1D494B7F4" box="[763,913,368,392]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA3FFBB0AE3FEF1D494B7F4" box="[763,913,368,392]" italics="true" pageId="29" pageNumber="30">C. belepensis</emphasis>
</taxonomicName>
have been at night while they are active and it appears to be an exclusively arboreal species. Unlike
<taxonomicName id="13E64D74FFA3FFBB0B0FFE15D4E6B7D0" box="[791,995,404,428]" class="Reptilia" family="Diplodactylidae" genus="Mniarogekko" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="jalu" status="sp. nov.">
<emphasis id="E692EAE5FFA3FFBB0B0FFE15D4E6B7D0" box="[791,995,404,428]" italics="true" pageId="29" pageNumber="30">Mniarogekko jalu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA3FFBB0BE8FE15D331B7D0" box="[1008,1076,404,428]" pageId="29" pageNumber="30" rank="species">
<emphasis id="E692EAE5FFA3FFBB0BE8FE15D328B7D0" bold="true" box="[1008,1069,404,428]" pageId="29" pageNumber="30">n. sp</emphasis>
.
</taxonomicNameLabel>
, with which it is syntopic,
<taxonomicName id="13E64D74FFA3FFBB0D9AFE14D609B7AC" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA3FFBB0D9AFE14D609B7AC" italics="true" pageId="29" pageNumber="30">C. belepensis</emphasis>
</taxonomicName>
was found at all levels within the forest from the upper surface of the canopy to low in sub-canopy and forest margin shrubs. Perches used were similarly varied and included foliage, twigs, branches and trunks. The new species is particularly agile when moving through the vegetation and frequently leaps between branches, as much as
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vertically upwards and&gt;
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horizontally. When threatened they jump from the canopy or other vegetation and fall, spread-eagled and horizontal, to lower plants or the forest floor. If pursued on the ground they move in a series of frog-like leaps, up to
<quantity id="131E9B12FFA3FFBB0A7EFDEDD5AEB4F8" box="[614,683,620,644]" metricMagnitude="-1" metricUnit="m" metricValue="1.2" pageId="29" pageNumber="30" unit="cm" value="12.0">12 cm</quantity>
high and covering
<quantity id="131E9B12FFA3FFBB0B9EFDEDD4F1B4F8" box="[902,1012,620,644]" metricMagnitude="-1" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="29" pageNumber="30" unit="cm" value="35.0" valueMax="40.0" valueMin="30.0">3040 cm</quantity>
in each bound. A similar tendency to jump has been documented in its sister species
<taxonomicName id="13E64D74FFA3FFBB0ABFFD11D301B4DB" authority="Vosjoli et al. 2003" authorityName="Vosjoli et al." authorityYear="2003" box="[679,1028,655,680]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA3FFBB0ABFFD11D41FB4D4" box="[679,794,656,680]" italics="true" pageId="29" pageNumber="30">C. ciliatus</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB0B31FD0ED4F9B4DB" author="Vosjoli" box="[809,1020,655,680]" pageId="29" pageNumber="30" refString="de Vosjoli, P., Fast, F. &amp; Repashy, A. (2003) Rhacodactylus, the Complete Guide to their Selelction and Care. Advanced Visions, Inc., Vista, California. 296 pp." type="book" year="2003">
Vosjoli
<emphasis id="E692EAE5FFA3FFBB0B99FD10D4BCB4D4" box="[897,953,656,680]" italics="true" pageId="29" pageNumber="30">et al.</emphasis>
2003
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
<caption id="8099667FFFA3FFBB088FFA3CD264B38E" httpUri="https://zenodo.org/record/211749/files/figure.png" pageId="29" pageNumber="30" targetBox="[217,1369,733,1447]" targetPageId="29">
<paragraph id="D45936F7FFA3FFBB088FFA3CD264B38E" blockId="29.[151,1436,1469,1522]" pageId="29" pageNumber="30">
<emphasis id="E692EAE5FFA3FFBB088FFA3CD625B3AE" bold="true" box="[151,288,1469,1492]" pageId="29" pageNumber="30">FIGURE 15.</emphasis>
Type series of
<taxonomicName id="13E64D74FFA3FFBB09A6FA3CD5AAB3AF" box="[446,687,1469,1491]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA3FFBB09A6FA3CD5AAB3AF" box="[446,687,1469,1491]" italics="true" pageId="29" pageNumber="30">Correlophus belepensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA3FFBB0AADFA3FD402B3AF" box="[693,775,1470,1491]" pageId="29" pageNumber="30" rank="species">
<emphasis id="E692EAE5FFA3FFBB0AADFA3FD402B3AF" bold="true" box="[693,775,1470,1491]" pageId="29" pageNumber="30">sp. nov.</emphasis>
</taxonomicNameLabel>
showing variation in color and in lumbosacral tubercles. Left to right: holotype (MNHN 2011.1100), paratypes (AMS R161282, AMS R161283, CAS 250865). Photo by A.H. Whitaker.
</paragraph>
</caption>
<paragraph id="D45936F7FFA3FFBB08DFF9C3D3F0B173" blockId="29.[151,1436,1602,2023]" pageId="29" pageNumber="30">
On Île Art
<taxonomicName id="13E64D74FFA3FFBB095CF9C2D6DEB027" box="[324,475,1603,1627]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA3FFBB095CF9C2D6DEB027" box="[324,475,1603,1627]" italics="true" pageId="29" pageNumber="30">C. belepensis</emphasis>
</taxonomicName>
is syntopic with
<taxonomicName id="13E64D74FFA3FFBB0AB8F9C2D46DB027" box="[672,872,1603,1627]" class="Reptilia" family="Diplodactylidae" genus="Mniarogekko" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="jalu" status="sp. nov.">
<emphasis id="E692EAE5FFA3FFBB0AB8F9C2D46DB027" box="[672,872,1603,1627]" italics="true" pageId="29" pageNumber="30">Mniarogekko jalu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA3FFBB0B68F9C2D4B5B027" box="[880,944,1603,1627]" pageId="29" pageNumber="30" rank="species">
<emphasis id="E692EAE5FFA3FFBB0B68F9C2D4AFB027" bold="true" box="[880,938,1603,1627]" pageId="29" pageNumber="30">n. sp</emphasis>
.
</taxonomicNameLabel>
,
<taxonomicName id="13E64D74FFA3FFBB0BD8F9C2D3E4B027" box="[960,1249,1603,1627]" class="Reptilia" family="Diplodactylidae" genus="Eurydactylodes" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="agricolae">
<emphasis id="E692EAE5FFA3FFBB0BD8F9C2D3E4B027" box="[960,1249,1603,1627]" italics="true" pageId="29" pageNumber="30">Eurydactylodes agricolae</emphasis>
</taxonomicName>
and
<taxonomicName id="13E64D74FFA3FFBB0D02F9C2D7FCB003" class="Reptilia" family="Diplodactylidae" genus="Dierogekko" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="insularis">
<emphasis id="E692EAE5FFA3FFBB0D02F9C2D7FCB003" italics="true" pageId="29" pageNumber="30">Dierogekko insularis</emphasis>
</taxonomicName>
. Species sympatric within its habitat include
<taxonomicName id="13E64D74FFA3FFBB0AE5F9E6D7D7B0DE" authority="Sadlier, Smith, Bauer &amp; Whitaker, 2004" authorityName="Sadlier, Smith, Bauer &amp; Whitaker" authorityYear="2004" class="Reptilia" family="Scincidae" genus="Kanakysaurus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="viviparus">
<emphasis id="E692EAE5FFA3FFBB0AE5F9E6D308B003" box="[765,1037,1639,1663]" italics="true" pageId="29" pageNumber="30">Kanakysaurus viviparus</emphasis>
<bibRefCitation id="B0774B06FFA3FFBB0C0EF9E6D7D7B0DE" author="Sadlier" pageId="29" pageNumber="30" refString="Whitaker, A. H., Sadlier, R. A., Bauer, A. M. &amp; Whitaker, V. A. (2004) Biodiversity and conservation status of lizards in threatened and restricted habitats of north-western New Caledonia. Unpublished report by Whitaker Consultants Limited to Direction du Developpement Economique et de l'Environnement, Province Nord, Kone, New Caledonia, viii + 106 pp." type="book" year="2004">Sadlier, Smith, Bauer &amp; Whitaker, 2004</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="13E64D74FFA3FFBB08E0F90DD421B0DE" authority="Peters, 1869" authorityName="Peters" authorityYear="1869" box="[248,804,1674,1699]" class="Reptilia" family="Scincidae" genus="Lioscincus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="nigrofasciolatus">
<emphasis id="E692EAE5FFA3FFBB08E0F90DD542B0DF" box="[248,583,1675,1699]" italics="true" pageId="29" pageNumber="30">Lioscincus nigrofasciolatus</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB0A69F90BD418B0DE" author="Peters" box="[625,797,1674,1698]" pageId="29" pageNumber="30" refString="Peters, W. (1869) Uber neue Gattungen und neue oder weniger bekannte Arten von Amphibien. Monatsberichte der Koniglichen Preussischen Akademie der Wissenschaften zu Berlin, 1869, 432 - 445, 1 pl." type="journal article" year="1869">Peters, 1869</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="13E64D74FFA3FFBB0B54F90AD292B0DE" authority="Roux, 1913" authorityName="Roux" authorityYear="1913" box="[844,1431,1674,1699]" class="Reptilia" family="Scincidae" genus="Caledoniscincus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="atropunctatus">
<emphasis id="E692EAE5FFA3FFBB0B54F90AD3C4B0DF" box="[844,1217,1675,1699]" italics="true" pageId="29" pageNumber="30">Caledoniscincus atropunctatus</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB0CF4F90BD28AB0DE" author="Roux" box="[1260,1423,1674,1698]" pageId="29" pageNumber="30" refString="Roux J. (1913) Les reptiles de la Nouvelle-Caledonie et des Iles Loyalty. In: Sarasin F. &amp; Roux J. (Eds.) Nova Caledonia, Zoologie. C. W. Kreidels Verlag, Wiesbaden, pp. 79 - 160, pls. 4 - 5." type="book chapter" year="1913">Roux, 1913</bibRefCitation>
)
</taxonomicName>
,
<taxonomicName id="13E64D74FFA3FFBB088FF931D528B0BB" authority="Bavay, 1869" authorityName="Bavay" authorityYear="1869" box="[151,557,1711,1735]" class="Reptilia" family="Scincidae" genus="Caledoniscincus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="austrocaledonicus">
<emphasis id="E692EAE5FFA3FFBB088FF931D686B0BB" box="[151,387,1711,1735]" italics="true" pageId="29" pageNumber="30">C. austrocaledonicus</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB098BF92ED520B0BB" author="Bavay" box="[403,549,1711,1735]" pageId="29" pageNumber="30" refString="Bavay, A. (1869) Catalogue des reptiles de la Nouvelle-Caledonie et description d'especes nouvelles. Memoires de la Societe Linneenne de Normandie, 15, 1 - 37. Bocage, J. V. Barboza du (1873 a) Note sur quelques geckotiens nouveaux ou peu connus de la Nouvelle Caledonie. Jornal de Sciencias Mathematicas, Physicas, e Naturaes, Academia Real das Sciencas de Lisboa, 4, 201 - 207." type="journal article" year="1869">Bavay, 1869</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="13E64D74FFA3FFBB0A73F931D4D5B0BB" authority="Gunther, 1872" authorityName="Gunther" authorityYear="1872" box="[619,976,1711,1735]" class="Reptilia" family="Scincidae" genus="Caledoniscincus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="haplorhinus">
<emphasis id="E692EAE5FFA3FFBB0A73F931D417B0BB" box="[619,786,1711,1735]" italics="true" pageId="29" pageNumber="30">C. haplorhinus</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB0B39F92ED4CCB0BB" author="Gunther" box="[801,969,1711,1735]" pageId="29" pageNumber="30" refString="Gunther, A. (1872) On some new species of reptiles and fishes collected by J. Brenchley, Esq. Annals and Magazine of Natural History, series 4, 10, 418 - 420." type="journal article" year="1872">Günther, 1872</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="13E64D74FFA3FFBB0C16F931D3F6B0BB" box="[1038,1267,1711,1735]" class="Reptilia" family="Diplodactylidae" genus="Bavayia" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="cyclura">
<emphasis id="E692EAE5FFA3FFBB0C16F931D36FB0BB" box="[1038,1130,1712,1735]" italics="true" pageId="29" pageNumber="30">Bavayia</emphasis>
aff.
<emphasis id="E692EAE5FFA3FFBB0C87F92ED3F6B0BB" box="[1183,1267,1711,1735]" italics="true" pageId="29" pageNumber="30">cyclura</emphasis>
</taxonomicName>
,
<taxonomicName id="13E64D74FFA3FFBB0D1AF92ED666B096" authority="Schlegel" authorityName="Schlegel" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="E692EAE5FFA3FFBB0D1AF92ED7F7B097" italics="true" pageId="29" pageNumber="30">Hemidactylus frenatus</emphasis>
Schlegel
</taxonomicName>
<emphasis id="E692EAE5FFA3FFBB096CF955D68FB097" box="[372,394,1748,1771]" italics="true" pageId="29" pageNumber="30">in</emphasis>
<bibRefCitation id="B0774B06FFA3FFBB0984F953D5D6B096" author="Dumeril" box="[412,723,1746,1770]" pageId="29" pageNumber="30" refString="Dumeril, A. - M. - C. &amp; Bibron, G. (1836) Erpetologie Generale ou Histoire Naturelle Complete des Reptiles, Tome 3; Librairie Encyclopedique de Roret, Paris, (2), (1), iv + 571 pp., errata, 2 folding tables., pls. 25 - 28, 33, 35, 43 - 48." type="book chapter" year="1836">Duméril &amp; Bibron, 1836</bibRefCitation>
,
<taxonomicName id="13E64D74FFA3FFBB0AF1F952D258B096" authority="Dumeril &amp; Bibron, 1836" authorityName="Dumeril &amp; Bibron" authorityYear="1836" box="[745,1373,1746,1771]" class="Reptilia" family="Gekkonidae" genus="Lepidodactylus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="lugubris">
<emphasis id="E692EAE5FFA3FFBB0AF1F952D307B097" box="[745,1026,1747,1771]" italics="true" pageId="29" pageNumber="30">Lepidodactylus lugubris</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB0C04F953D253B096" author="Dumeril" box="[1052,1366,1746,1770]" pageId="29" pageNumber="30" refString="Dumeril, A. - M. - C. &amp; Bibron, G. (1836) Erpetologie Generale ou Histoire Naturelle Complete des Reptiles, Tome 3; Librairie Encyclopedique de Roret, Paris, (2), (1), iv + 571 pp., errata, 2 folding tables., pls. 25 - 28, 33, 35, 43 - 48." type="book chapter" year="1836">Duméril &amp; Bibron, 1836</bibRefCitation>
)
</taxonomicName>
, and
<taxonomicName id="13E64D74FFA3FFBB088FF976D53EB173" authority="Bavay, 1869" authorityName="Bavay" authorityYear="1869" box="[151,571,1783,1807]" class="Reptilia" family="Scincidae" genus="Phoboscincus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="garnieri">
<emphasis id="E692EAE5FFA3FFBB088FF976D690B173" box="[151,405,1783,1807]" italics="true" pageId="29" pageNumber="30">Phoboscincus garnieri</emphasis>
(
<bibRefCitation id="B0774B06FFA3FFBB09BCF976D536B173" author="Bavay" box="[420,563,1783,1807]" pageId="29" pageNumber="30" refString="Bavay, A. (1869) Catalogue des reptiles de la Nouvelle-Caledonie et description d'especes nouvelles. Memoires de la Societe Linneenne de Normandie, 15, 1 - 37. Bocage, J. V. Barboza du (1873 a) Note sur quelques geckotiens nouveaux ou peu connus de la Nouvelle Caledonie. Jornal de Sciencias Mathematicas, Physicas, e Naturaes, Academia Real das Sciencas de Lisboa, 4, 201 - 207." type="journal article" year="1869">Bavay, 1869</bibRefCitation>
)
</taxonomicName>
were recorded in adjacent savannah and secondary shrubland.
</paragraph>
<paragraph id="D45936F7FFA3FFBB08DFF89BD682B12B" blockId="29.[151,1436,1602,2023]" pageId="29" pageNumber="30">Trombiculid mites are present between the subdigital lamellae and in one or both popliteal pockets in all of the specimens examined.</paragraph>
<paragraph id="D45936F7FFA3FFBB08DFF8E3D451B19B" blockId="29.[151,1436,1602,2023]" pageId="29" pageNumber="30">
<emphasis id="E692EAE5FFA3FFBB08DFF8E3D6BAB107" bold="true" box="[199,447,1890,1915]" pageId="29" pageNumber="30">Conservation status.</emphasis>
<taxonomicName id="13E64D74FFA3FFBB09D0F8E2D5D1B107" box="[456,724,1891,1915]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="29" pageNumber="30" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA3FFBB09D0F8E2D5D1B107" box="[456,724,1891,1915]" italics="true" pageId="29" pageNumber="30">Correlophus belepensis</emphasis>
</taxonomicName>
has been recorded only on Île Art, in the Belep archipelago, where it occurs as two sub-populations within a single locality. It is expected that the species may also occur in similar habitat on nearby Île Pott. Whether its entire distribution is confined to the Belep archipelago is unclear but at present there is no evidence of populations on the mainland.
</paragraph>
<caption id="8099667FFFA0FFB8088FFCDCD423B5EE" httpUri="https://zenodo.org/record/211750/files/figure.png" pageId="30" pageNumber="31" targetBox="[337,1249,229,840]" targetPageId="30">
<paragraph id="D45936F7FFA0FFB8088FFCDCD423B5EE" blockId="30.[151,1435,861,914]" pageId="30" pageNumber="31">
<emphasis id="E692EAE5FFA0FFB8088FFCDCD61AB50F" bold="true" box="[151,287,861,884]" pageId="30" pageNumber="31">FIGURE 16.</emphasis>
Habitat of
<taxonomicName id="13E64D74FFA0FFB80989FCDFD57AB508" box="[401,639,862,884]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="belepensis" status="sp. nov.">
<emphasis id="E692EAE5FFA0FFB80989FCDFD57AB508" box="[401,639,862,884]" italics="true" pageId="30" pageNumber="31">Correlophus belepensis</emphasis>
</taxonomicName>
<emphasis id="E692EAE5FFA0FFB80A9EFCDFD5D3B50F" bold="true" box="[646,726,862,883]" pageId="30" pageNumber="31">
<taxonomicNameLabel id="FDA1579EFFA0FFB80A9EFCDFD5D3B50F" box="[646,726,862,883]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="13E64D74FFA0FFB80B10FCDFD4B9B508" box="[776,956,862,884]" class="Reptilia" family="Diplodactylidae" genus="Mniarogekko" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="jalu" status="sp. nov.">
<emphasis id="E692EAE5FFA0FFB80B10FCDFD4B9B508" box="[776,956,862,884]" italics="true" pageId="30" pageNumber="31">Mniarogekko jalu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA0FFB80BDAFCDFD317B50F" box="[962,1042,862,883]" pageId="30" pageNumber="31" rank="species">
<emphasis id="E692EAE5FFA0FFB80BDAFCDFD317B50F" bold="true" box="[962,1042,862,883]" pageId="30" pageNumber="31">sp. nov.</emphasis>
</taxonomicNameLabel>
on the Tolé Munu plateau, Île Art, Îles Belep, Province Nord, New Caledonia. Photo by A.H. Whitaker.
</paragraph>
</caption>
<paragraph id="D45936F7FFA0FFB808DFFC3ED6CDB2D3" blockId="30.[151,1436,959,2027]" pageId="30" pageNumber="31">
Within the known extent of occurrence for
<taxonomicName id="13E64D74FFA0FFB80AD8FC41D45CB5AB" box="[704,857,959,983]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA0FFB80AD8FC41D45CB5AB" box="[704,857,959,983]" italics="true" pageId="30" pageNumber="31">C. belepensis</emphasis>
</taxonomicName>
, the closed forest and paraforestier formations on cuirasse surfaces are surprisingly intact given the high level of habitat modification that has occurred elsewhere on Île Art. Both plateaux are still largely covered by forest, but as a result of repeated burning and clearance for subsistence gardening since colonization by Melanesians, the surrounding slopes and remainder of the island have been modified and are characterized by depleted niaouli (
<taxonomicName id="13E64D74FFA0FFB80B03FBCED213B21B" authority="(Cav.) Blake, Fam." authorityName="(Cav.) Blake, Fam." box="[795,1302,1103,1127]" class="Magnoliopsida" family="Myrtaceae" genus="Melaleuca" higherTaxonomySource="CoL" kingdom="Fungi" order="Myrtales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="species" species="quinquenervia">
<emphasis id="E692EAE5FFA0FFB80B03FBCED33FB21B" box="[795,1082,1103,1127]" italics="true" pageId="30" pageNumber="31">Melaleuca quinquenervia</emphasis>
(Cav.) Blake, Fam.
</taxonomicName>
<taxonomicName id="13E64D74FFA0FFB80D04FBCED291B21B" box="[1308,1428,1103,1127]" class="Magnoliopsida" family="Myrtaceae" kingdom="Plantae" order="Myrtales" pageId="30" pageNumber="31" phylum="Tracheophyta" rank="family">Myrtaceae</taxonomicName>
) savannah, secondary shrublands and grassland. This is assumed to have resulted in a contraction of the area of occupation for this species.
</paragraph>
<paragraph id="D45936F7FFA0FFB808DFFB3AD4EEB023" blockId="30.[151,1436,959,2027]" pageId="30" pageNumber="31">
The sub-populations of
<taxonomicName id="13E64D74FFA0FFB809C4FB3DD573B2AF" box="[476,630,1211,1235]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA0FFB809C4FB3DD573B2AF" box="[476,630,1211,1235]" italics="true" pageId="30" pageNumber="31">C. belepensis</emphasis>
</taxonomicName>
on Île Art are at risk to several direct and indirect threats. Wildfires regularly affect large areas of the grassland, savannah and shrubland, and these inevitably encroach on the remaining forest, reducing it in extent. There is also sporadic timber cutting for local use but to date this has had only a very localized impact. Introduced rats (
<taxonomicName id="13E64D74FFA0FFB80ABEFAA9D5EBB343" box="[678,750,1320,1343]" class="Mammalia" family="Muridae" genus="Rattus" kingdom="Animalia" order="Rodentia" pageId="30" pageNumber="31" phylum="Chordata" rank="genus">
<emphasis id="E692EAE5FFA0FFB80ABEFAA9D5EBB343" box="[678,750,1320,1343]" italics="true" pageId="30" pageNumber="31">Rattus</emphasis>
</taxonomicName>
spp.) and feral cats are present throughout the habitat. Both are known to be predators of lizards but no direct evidence of predation on
<taxonomicName id="13E64D74FFA0FFB80BEAFACDD38DB31F" box="[1010,1160,1355,1379]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA0FFB80BEAFACDD38DB31F" box="[1010,1160,1355,1379]" italics="true" pageId="30" pageNumber="31">C. belepensis</emphasis>
</taxonomicName>
was obtained. Little red fire ants,
<taxonomicName id="13E64D74FFA0FFB8091FFAF1D5D1B3FB" authority="Roger, 1863" authorityName="Roger" authorityYear="1863" box="[263,724,1391,1415]" class="Insecta" family="Formicidae" genus="Wasmannia" kingdom="Animalia" order="Hymenoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="auropunctata">
<emphasis id="E692EAE5FFA0FFB8091FFAF1D52FB3FB" box="[263,554,1392,1415]" italics="true" pageId="30" pageNumber="31">Wasmannia auropunctata</emphasis>
(
<bibRefCitation id="B0774B06FFA0FFB80A24FAEED5C9B3FB" author="Roger" box="[572,716,1391,1415]" pageId="30" pageNumber="31" refString="Roger, J. (1863) Die neu aufgefuhrten Gattungen und Arten meines Formiciden Verzeichnisses. Berliner Entomologische Zeitschrift, 7, 131 - 214." type="journal article" year="1863">Roger, 1863</bibRefCitation>
)
</taxonomicName>
are also present in the forest on the plateaux. These highlyinvasive ants are known to have a severe detrimental impact on lizard populations (
<bibRefCitation id="B0774B06FFA0FFB80C79FA12D243B3D7" author="Jourdan" box="[1121,1350,1427,1451]" pageId="30" pageNumber="31" refString="Jourdan, H., Sadlier, R. &amp; Bauer, A. (2000) Premieres observations sur les consequences de l'invasion de Wasmannia auropunctata 1863 (Roger) sur les predateurs superieurs dans les ecosystemes Neo-caledoniens. Actes des Colloques Insectes Sociaux, 13, 121 - 126" type="journal article" year="2000">
Jourdan
<emphasis id="E692EAE5FFA0FFB80CDEFA15D204B3D7" box="[1222,1281,1427,1451]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2000
</bibRefCitation>
,
<bibRefCitation id="B0774B06FFA0FFB80D4CFA12D295B3D7" author="Jourdan" box="[1364,1424,1427,1451]" pageId="30" pageNumber="31" refString="Jourdan, H., Sadlier, R. A. &amp; Bauer, A. M. (2001) Little fire ant invasion (Wasmannia auropunctata) as a threat to New Caledonian lizards: evidences from the sclerophyll forest (Hymenoptera: Formicidae). Sociobiology, 38, 283 - 301." type="journal article" year="2001">2001</bibRefCitation>
).
<taxonomicName id="13E64D74FFA0FFB8088FFA36D696B3B3" box="[151,403,1463,1487]" class="Reptilia" family="Gekkonidae" genus="Hemidactylus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="frenatus">
<emphasis id="E692EAE5FFA0FFB8088FFA36D696B3B3" box="[151,403,1463,1487]" italics="true" pageId="30" pageNumber="31">Hemidactylus frenatus</emphasis>
</taxonomicName>
has colonized the Îles Belep relatively recently and was detected in the township of Waala, as well as in littoral vegetation, garden sites and savannah woodlands. If it spreads into forest habitat, as is expected, it is unclear what impact this invasive
<taxonomicName id="13E64D74FFA0FFB80ACCFA7ED412B06B" box="[724,791,1535,1559]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
would have on giant
<taxonomicName id="13E64D74FFA0FFB80C04FA7ED35AB06B" box="[1052,1119,1535,1559]" class="Reptilia" family="Gekkonidae" genus="Gekko" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="gecko">gecko</taxonomicName>
species but elsewhere it is known to result in competitive exclusion and local extirpation of smaller indigenous geckos (e.g.,
<bibRefCitation id="B0774B06FFA0FFB80CF8F9A2D292B047" author="Case" box="[1248,1431,1571,1595]" pageId="30" pageNumber="31" refString="Case, T. J., Bolger, D. T. &amp; Petren, K. (1994) Invasions and competitive displacement among house geckos in the tropical Pacific. Ecology, 75, 464 - 477" type="journal article" year="1994">
Case
<emphasis id="E692EAE5FFA0FFB80D05F9A5D250B047" box="[1309,1365,1571,1595]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
1994
</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA0FFB8088FF9C9D65AB023" author="Petren" box="[151,351,1607,1631]" pageId="30" pageNumber="31" refString="Petren, K., Bolger, D. T. &amp; Case, T. J. (1993) Mechanisms in the competitive success of an invading sexual gecko over an asexual native. Science, 259, 354 - 358." type="journal article" year="1993">
Petren
<emphasis id="E692EAE5FFA0FFB808FDF9C9D610B023" box="[229,277,1607,1631]" italics="true" pageId="30" pageNumber="31">et al</emphasis>
. 1993
</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA0FFB80973F9C9D54AB023" author="Petren" box="[363,591,1607,1631]" pageId="30" pageNumber="31" refString="Petren, K. &amp; Case, T. J. (1996) An experimental demonstration of exploitation competition in an ongoing invasion. Ecology, 77, 118 - 132" type="journal article" year="1996">Petren &amp; Case 1996</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA0FFB80A43F9C6D415B023" author="Cole" box="[603,784,1607,1631]" pageId="30" pageNumber="31" refString="Cole, N. C., Jones, C. G. &amp; Harris, S. (2005) The need for enemy-free species: the impact of an invasive gecko on island endemics. Biological Conservation, 125, 467 - 474" type="journal article" year="2005">
Cole
<emphasis id="E692EAE5FFA0FFB80A8EF9C9D5CAB023" box="[662,719,1607,1631]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2005
</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA0FFB80B04F9C6D4DAB023" author="Rivas" box="[796,991,1607,1631]" pageId="30" pageNumber="31" refString="Rivas Fuenmayor, G., Barros, T., Ugueto, G. N. &amp; Bauer, A. M. (2005) Expansion and natural habits of a successful colonizing gecko in Venezuela (Reptilia: Gekkonidae: Hemidactylus mabouia) and the discovery of H. frenatus in Venezuela. Herpetological Review, 36, 121 - 125." type="journal article" year="2005">
Rivas
<emphasis id="E692EAE5FFA0FFB80B7BF9C9D49EB023" box="[867,923,1607,1631]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2005
</bibRefCitation>
).
</paragraph>
<paragraph id="D45936F7FFA0FFB808DFF9EAD3AAB093" blockId="30.[151,1436,959,2027]" pageId="30" pageNumber="31">The ultramafic plateaux on both Île Art and Île Pott have been subjected to extensive prospecting in the past and are criss-crossed by old access tracks and drill sites. Although these areas are held under current mining licenses there appear to be no immediate plans to exploit the areas. However, should it occur, mining would almost certainly remove a significant proportion—if not all—of the available habitat for this species.</paragraph>
<paragraph id="D45936F7FFA0FFB808DFF97DD23EB127" blockId="30.[151,1436,959,2027]" pageId="30" pageNumber="31">
No quantitative data on population size and trends are available for
<taxonomicName id="13E64D74FFA0FFB80BC1F97DD374B16F" box="[985,1137,1787,1811]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA0FFB80BC1F97DD374B16F" box="[985,1137,1787,1811]" italics="true" pageId="30" pageNumber="31">C. belepensis</emphasis>
</taxonomicName>
.
<bibRefCitation id="B0774B06FFA0FFB80C9AF97DD3DFB16F" author="In" box="[1154,1242,1787,1811]" pageId="30" pageNumber="31" refString="IUCN (2001) IUCN Red List categories: Version 3.1. IUCN Species Survival Commission, Gland, Switzerland, and Cambridge, UK, 26 pp." type="book" year="2001">In 2001</bibRefCitation>
it was relatively abundant in the forest on the Tolé Munu plateau, with encounter rates of 0.570.83/person hour, implying a population density approximately half that of the syntopic
<taxonomicName id="13E64D74FFA0FFB80B3FF8C2D4EBB127" box="[807,1006,1859,1883]" class="Reptilia" family="Diplodactylidae" genus="Mniarogekko" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="jalu" status="sp. nov.">
<emphasis id="E692EAE5FFA0FFB80B3FF8C2D4EBB127" box="[807,1006,1859,1883]" italics="true" pageId="30" pageNumber="31">Mniarogekko jalu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FDA1579EFFA0FFB80BEDF8C2D331B127" box="[1013,1076,1859,1883]" pageId="30" pageNumber="31" rank="species">
<emphasis id="E692EAE5FFA0FFB80BEDF8C2D331B127" bold="true" box="[1013,1076,1859,1883]" pageId="30" pageNumber="31">n. sp.</emphasis>
</taxonomicNameLabel>
(
<bibRefCitation id="B0774B06FFA0FFB80C5AF8C2D22AB127" author="Whitaker" box="[1090,1327,1859,1883]" pageId="30" pageNumber="31" refString="Whitaker, A. H., Sadlier, R. A., Bauer, A. M. &amp; Whitaker, V. A. (2004) Biodiversity and conservation status of lizards in threatened and restricted habitats of north-western New Caledonia. Unpublished report by Whitaker Consultants Limited to Direction du Developpement Economique et de l'Environnement, Province Nord, Kone, New Caledonia, viii + 106 pp." type="book" year="2004">
Whitaker
<emphasis id="E692EAE5FFA0FFB80CABF8C5D3EEB127" box="[1203,1259,1859,1883]" italics="true" pageId="30" pageNumber="31">et al.</emphasis>
2004
</bibRefCitation>
).
</paragraph>
<paragraph id="D45936F7FFA0FFB808DFF8E9D50AB197" blockId="30.[151,1436,959,2027]" pageId="30" pageNumber="31">
Because of its extremely limited extent of occurrence (&lt;
<quantity id="131E9B12FFA0FFB80B23F8E6D486B103" box="[827,899,1895,1919]" metricMagnitude="4" metricUnit="m" metricValue="2.5" pageId="30" pageNumber="31" unit="km" value="25.0">25 km</quantity>
²) and area of occupation (&lt;
<quantity id="131E9B12FFA0FFB80CA0F8E6D207B103" box="[1208,1282,1895,1919]" metricMagnitude="4" metricUnit="m" metricValue="1.0" pageId="30" pageNumber="31" unit="km" value="10.0">10 km</quantity>
²), presence at a single locality, the threats to its habitat (wildfires, and perhaps mining), the presence of mammalian predators (rats, cats) and the impacts of fire ants,
<taxonomicName id="13E64D74FFA0FFB80A41F82ED466B1BB" box="[601,867,1967,1991]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="30" pageNumber="31" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA0FFB80A41F82ED466B1BB" box="[601,867,1967,1991]" italics="true" pageId="30" pageNumber="31">Correlophus belepensis</emphasis>
</taxonomicName>
is assessed as Critically Endangered (B1a, b[iiii, v]; B2a, b[iiii, v]) (
<bibRefCitation id="B0774B06FFA0FFB80961F852D506B197" author="IUCN" box="[377,515,2003,2027]" pageId="30" pageNumber="31" refString="IUCN (2001) IUCN Red List categories: Version 3.1. IUCN Species Survival Commission, Gland, Switzerland, and Cambridge, UK, 26 pp." type="book" year="2001">IUCN 2001</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9CFC657CFFA1FFB908DFFF19D643B788" pageId="31" pageNumber="32" type="discussion">
<paragraph id="D45936F7FFA1FFB908DFFF19D62FB743" blockId="31.[151,1436,151,500]" pageId="31" pageNumber="32">
<emphasis id="E692EAE5FFA1FFB908DFFF19D63EB6CC" bold="true" box="[199,315,152,176]" pageId="31" pageNumber="32">Remarks.</emphasis>
<taxonomicName id="13E64D74FFA1FFB90959FF19D522B6CC" box="[321,551,152,176]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA1FFB90959FF19D522B6CC" box="[321,551,152,176]" italics="true" pageId="31" pageNumber="32">Correlophus ciliatus</emphasis>
</taxonomicName>
occurs in southern Grande Terre and on the Île des Pins, with a northern known limit of range around
<quantity id="131E9B12FFA1FFB90988FF3DD6E0B6A8" box="[400,485,188,212]" metricMagnitude="5" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="km" value="300.0">300 km</quantity>
south-east of the Îles Belep (Bauer &amp; Sadlier 2000;
<figureCitation id="4CDD2A72FFA1FFB90C25FF3DD38BB6A8" box="[1085,1166,188,212]" captionStart="FIGURE 11" captionStartId="25.[151,250,1357,1380]" captionTargetBox="[160,1426,418,1328]" captionTargetId="figure@25.[151,1436,409,1336]" captionTargetPageId="25" captionText="FIGURE 11. Distribution map of Correlophus ciliatus (circles) and C. belepensis sp. nov. (red star). For C. ciliatus green circles symbols represent vouchered records, orange circles represent unvouchered sight or literature records. Vouchered records from the Île des Pins lack precise localities. See Appendix for a list of localities mapped." httpUri="https://zenodo.org/record/211745/files/figure.png" pageId="31" pageNumber="32">Fig. 11</figureCitation>
). The presence of these two taxa at either end of Grande Terre suggests the occurrence of
<taxonomicName id="13E64D74FFA1FFB90B86FF61D332B684" box="[926,1079,224,248]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="belepensis">
<emphasis id="E692EAE5FFA1FFB90B86FF61D332B684" box="[926,1079,224,248]" italics="true" pageId="31" pageNumber="32">C. belepensis</emphasis>
</taxonomicName>
is relictual and that it is not a primary endemic in the Belep group, but extensive fieldwork in the northern Grande Terre has not revealed any
<taxonomicName id="13E64D74FFA1FFB9088FFEA9D626B73C" box="[151,291,296,320]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="genus">
<emphasis id="E692EAE5FFA1FFB9088FFEA9D626B73C" box="[151,291,296,320]" italics="true" pageId="31" pageNumber="32">Correlophus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="D45936F7FFA1FFB908DFFECDD643B788" blockId="31.[151,1436,151,500]" pageId="31" pageNumber="32">
Although the diagnostic heterogeneous paravertebral trunk scalation of
<taxonomicName id="13E64D74FFA1FFB90BF5FECCD365B718" box="[1005,1120,332,356]" class="Reptilia" family="Diplodactylidae" genus="Correlophus" kingdom="Animalia" order="Squamata" pageId="31" pageNumber="32" phylum="Chordata" rank="species" species="ciliatus">
<emphasis id="E692EAE5FFA1FFB90BF5FECCD365B718" box="[1005,1120,332,356]" italics="true" pageId="31" pageNumber="32">C. ciliatus</emphasis>
</taxonomicName>
is not illustrated in the
<typeStatus id="0B5D8855FFA1FFB90D73FECCD29EB719" box="[1387,1435,333,357]" pageId="31" pageNumber="32">type</typeStatus>
illustration of that species (
<bibRefCitation id="B0774B06FFA1FFB909C4FEEED59EB7FB" author="Guichenot" box="[476,667,367,391]" pageId="31" pageNumber="32" refString="Guichenot, A. (1866) Notice sur un nouveau genre de sauriens de la famille geckotiens, du Museum de Paris. Memoires de la Societe des Sciences Naturelles de Cherbourg, 12, 248 - 252." type="journal article" year="1866">Guichenot 1866</bibRefCitation>
), it is clearly visible in the
<typeStatus id="0B5D8855FFA1FFB90BF7FEEED353B7FB" box="[1007,1110,367,391]" pageId="31" pageNumber="32" type="lectotype">lectotype</typeStatus>
specimens (
<figureCitation id="4CDD2A72FFA1FFB90CF4FEEED239B7FB" box="[1260,1340,367,391]" captionStart="FIGURE 10" captionStartId="24.[151,250,1052,1075]" captionTargetBox="[212,1374,416,1016]" captionTargetId="figure@24.[205,1382,407,1030]" captionTargetPageId="24" captionText="FIGURE 10. Lectotype of Correlophus ciliatus (MNHN 701 A) showing the absence of lumbosacral tubercles and presence of heterogeneous dorsolateral scalation. Photo courtesy of Muséum National dHistoire Naturelle, Paris." httpUri="https://zenodo.org/record/211744/files/figure.png" pageId="31" pageNumber="32">Fig 10</figureCitation>
) and in published photos of the species (e.g.,
<bibRefCitation id="B0774B06FFA1FFB90A25FE15D448B7D0" author="Seipp" box="[573,845,404,428]" pageId="31" pageNumber="32" refString="Seipp, R. &amp; Klemmer, K. (1994) Wiederentdeckung von Rhacodactylus ciliatus Guichenot 1866 im Suden Neukaledoniens (Reptilia: Sauria: Gekkonidae). Senckenbergiana Biologica, 24, 199 - 204." type="journal article" year="1994">Seipp &amp; Klemmer 1994</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA1FFB90B41FE15D4ECB7D0" author="Gerard" box="[857,1001,404,428]" pageId="31" pageNumber="32" refString="Gerard, P. (1999) Rhacodactylus ciliatus. The enigmatic crested gecko. Reptilia (GB), 9 (October), 27 - 33." type="journal article" year="1999">Gerard 1999</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA1FFB90BEDFE15D3EEB7D0" author="Seipp" box="[1013,1259,404,428]" pageId="31" pageNumber="32" refString="Vences, M., Henkel, F. - W. &amp; Seipp, R. (2001) Molekulare Untersuchungen zur Phylogenie und Taxonomie der Neukaledonischen Geckos der Gattung Rhacodactylus (Reptilia: Gekkonidae). Salamandra, 37, 73 - 82." type="journal article" year="2001">Seipp &amp; Henkel 2001</bibRefCitation>
,
<bibRefCitation id="B0774B06FFA1FFB90CEEFE15D235B7D0" author="Henkel" box="[1270,1328,404,428]" pageId="31" pageNumber="32" refString="Seipp, R. &amp; Henkel, F. - W. (2011) Rhacodactylus: Biology, Natural History and Husbandry, 2 nd ed. Edition Chimaira, Frankfurt am Main, 173 pp." type="book" year="2011">2011</bibRefCitation>
; Bauer &amp; Sadlier 2000; de
<bibRefCitation id="B0774B06FFA1FFB90942FE36D52AB7B3" author="Vosjoli" box="[346,559,439,464]" pageId="31" pageNumber="32" refString="de Vosjoli, P., Fast, F. &amp; Repashy, A. (2003) Rhacodactylus, the Complete Guide to their Selelction and Care. Advanced Visions, Inc., Vista, California. 296 pp." type="book" year="2003">
Vosjoli
<emphasis id="E692EAE5FFA1FFB909A9FE38D6EEB7AC" box="[433,491,440,464]" italics="true" pageId="31" pageNumber="32">et al.</emphasis>
2003
</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA1FFB90A23FE36D5BCB7B3" author="Bach" box="[571,697,439,463]" pageId="31" pageNumber="32" refString="Bach, S. (2006) Der Kronengecko, Rhacodactylus ciliatus. Natur und Tier-Verlag, Munster. 64 pp." type="book" year="2006">Bach 2006</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA1FFB90ADEFE36D4E5B7B3" author="Henkel" box="[710,992,439,463]" pageId="31" pageNumber="32" refString="Henkel, F. - W. &amp; Schmidt, W. (2007) Rhacodactylus ciliatus und auriculatus. Herpeton, Offenbach, 94 pp." type="book" year="2007">Henkel &amp; Schmidt 2007</bibRefCitation>
;
<bibRefCitation id="B0774B06FFA1FFB90BF4FE39D38FB7B3" author="Hamper" box="[1004,1162,439,463]" pageId="31" pageNumber="32" refString="Hamper, R. (2003) The Crested Gecko in Captivity. ECO Publishing, Lansing, Michigan, [10] + 69 pp." type="book" year="2003">Hamper 2003</bibRefCitation>
; Wirth &amp; Peukert 2009;
<bibRefCitation id="B0774B06FFA1FFB9088FFE5DD63FB788" author="Sommer" box="[151,314,476,500]" pageId="31" pageNumber="32" refString="Sommer, L. (2009) Introduktion til Rhacodactylus ciliatus - Kronegekko. Nordisk Herpetologisk Forening, 51,178 - 185." type="book" year="2009">Sommer 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>