366 lines
55 KiB
XML
366 lines
55 KiB
XML
<document ID-CLB-Dataset="21747" ID-DOI="http://dx.doi.org/10.3897/zookeys.938.51234" ID-GBIF-Dataset="0223fbe1-35f8-4987-b029-29b62cdce2e6" ID-PMC="PMC7289900" ID-Pensoft-Pub="1313-2970-938-137" ID-Pensoft-UUID="258F1856E4025778B55F9D603A15DD8C" ID-PubMed="32550789" ID-ZooBank="C57570A14A484EA3889E0646632DCA9B" ModsDocID="1313-2970-938-137" checkinTime="1591376751164" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I., Thach, Phanara, Chhuoy, Samol, Ngor, Peng Bun, Srisonchai, Ruttapon, Sutcharit, Chirasak & Panha, Somsak" docDate="2020" docId="5C16212414185C3B96ED0B45CC259ED9" docLanguage="en" docName="ZooKeys 938: 137-151" docOrigin="ZooKeys 938" docSource="http://dx.doi.org/10.3897/zookeys.938.51234" docTitle="Plusioglyphiulus khmer Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha 2020, sp. nov." docType="treatment" docUuid="7721A428-FA76-4226-9E3E-42CF2100475B" docUuidSource="ZooBank" docVersion="6" id="258F1856E4025778B55F9D603A15DD8C" lastPageNumber="137" masterDocId="258F1856E4025778B55F9D603A15DD8C" masterDocTitle="Two new species of the millipede genus Plusioglyphiulus Silvestri, 1923 from Cambodia (Diplopoda, Spirostreptida)" masterLastPageNumber="151" masterPageNumber="137" pageNumber="137" updateTime="1732798675588" updateUser="ExternalLinkService">
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<mods:title id="16F2448E45C55D0682FEC6AC337D1017">Two new species of the millipede genus Plusioglyphiulus Silvestri, 1923 from Cambodia (Diplopoda, Spirostreptida)</mods:title>
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<mods:name id="0DAC434B822B34AFBD7AD19835548907" type="personal">
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<mods:roleTerm id="7F65339527BBA60919505FFFC0F26E1E">Author</mods:roleTerm>
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<mods:namePart id="FDA9B070936AAA4250E3289D5558B3C4">Likhitrakarn, Natdanai</mods:namePart>
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<mods:affiliation id="44BA05EDA69608167AAA466AE37380F1">Division of Plant Protection, Faculty of Agricultural Production, Maejo University, Chiang Mai 50290, Thailand</mods:affiliation>
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<mods:namePart id="5FC83277C482A4776D793D2828D0D12E">Golovatch, Sergei I.</mods:namePart>
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<mods:affiliation id="E9DB9867A16DA2475409FEC66C682C39">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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</mods:name>
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<mods:name id="6597D4D66257D5336AFD41BE04AC6596" type="personal">
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<mods:roleTerm id="18A1E112D65FB366DF478CFE988C35D9">Author</mods:roleTerm>
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<mods:namePart id="600ABD2551C0A1BAC90228C17C912E1A">Thach, Phanara</mods:namePart>
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<mods:affiliation id="DAE194D7110ED399598CB21998CAC6A7">Inland Fisheries Research and Development Institute (IFReDI) No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<mods:name id="CC8477C132A472C25AE47738CD7A8AFD" type="personal">
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<mods:roleTerm id="04D704C36085A9E61B55FB8074BB8B85">Author</mods:roleTerm>
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<mods:namePart id="B7482071CF8134DF062379388ADD36A2">Chhuoy, Samol</mods:namePart>
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<mods:affiliation id="CC8E82DC48D78BE6E6C465A68A1D1B5A">Inland Fisheries Research and Development Institute (IFReDI) No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<mods:name id="E0525B251B32C8480611352D5E1E0FA5" type="personal">
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<mods:roleTerm id="40AE24742CE6332D7B7C963FB967115F">Author</mods:roleTerm>
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<mods:namePart id="372F91C90635B8E75CD848AECBECB7DB">Ngor, Peng Bun</mods:namePart>
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<mods:affiliation id="CB382091EBE30CBEBF84022A9BEED5E0">Inland Fisheries Research and Development Institute (IFReDI) No. 86, Norodom Blvd., PO Box 582, Phnom Penh, Cambodia</mods:affiliation>
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<mods:nameIdentifier id="82E50E81DC5F5DED9F603F506B599A11" type="orcid">https://orcid.org/0000-0002-3659-6577</mods:nameIdentifier>
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<mods:namePart id="FB9188D111E7CBA90EDD9662EF161242">Srisonchai, Ruttapon</mods:namePart>
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<mods:affiliation id="86AF555DECC8069EE278A79824698552">Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen, 40002, Thailand</mods:affiliation>
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</mods:name>
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<mods:name id="B27BEC42249C7DBE9D0C46BC3D44EC5C" type="personal">
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<mods:namePart id="C28B5D2CCB9FE37F26F2B1741082E786">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="944E9EFD4E1A6CC99FC3717AB2CFDAAC">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="C3329FCB5BC71CF79B38281426023180">Panha, Somsak</mods:namePart>
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<mods:affiliation id="425F169410E0403D794904396C12E554">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="8525FA004D41B48478C17057AED7D8D3" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title id="F5C16AD56A51385904A98DE81D6E0C5C">ZooKeys</mods:title>
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<mods:date id="1F6D6CDDDC2A819B6CDCCF8D5524E4B9">2020</mods:date>
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<mods:number id="73503D168600E99781E1175649A65536">938</mods:number>
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<mods:url id="9FBDC38C1AAF5C9142073EFBB5672B24">http://dx.doi.org/10.3897/zookeys.938.51234</mods:url>
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<mods:classification id="D8E977CBBB7C5D39394233B0E862E659">journal article</mods:classification>
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<mods:identifier id="07A011FB866AE415ECD0DEFBC992FAFB" type="DOI">http://dx.doi.org/10.3897/zookeys.938.51234</mods:identifier>
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<mods:identifier id="64BE54BEF479253AF2F240E15D02BD40" type="Pensoft-Pub">1313-2970-938-137</mods:identifier>
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<treatment id="5C16212414185C3B96ED0B45CC259ED9" ID-GBIF-Taxon="164479804" LSID="urn:lsid:zoobank.org:act:7721A428-FA76-4226-9E3E-42CF2100475B" httpUri="http://treatment.plazi.org/id/5C16212414185C3B96ED0B45CC259ED9" lastPageNumber="137" pageId="0" pageNumber="137" scope_class="Diplopoda" scope_family="Cambalopsidae" scope_order="Spirostreptida">
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<subSubSection id="840556CFBA46F84FD39AD923DFA42761" pageId="0" pageNumber="137" type="nomenclature">
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<paragraph id="84DC3461733C20B9CE0B52F63F765DC2" pageId="0" pageNumber="137">
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<taxonomicName id="2DD0B85DFFF3E7322666408B2039F289" ID-CoL="8NB9G" LSID="5C162124-1418-5C3B-96ED-0B45CC259ED9" authority="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha, 2020" authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusioglyphiulus khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer" status="sp. nov.">Plusioglyphiulus khmer</taxonomicName>
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<taxonomicNameLabel id="B044E237AC0B6568C3CBF728EB78230F" pageId="0" pageNumber="137">sp. nov.</taxonomicNameLabel>
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<figureCitation id="61BBD8CBB151DD41CB612DBA0C56A95F" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">Figures 1C</figureCitation>
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<figureCitation id="984DC47BFF03459809F3B79D92A55F3F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">, 4</figureCitation>
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<figureCitation id="6D08CE5C88CF9076B3DCCC3532A5F82F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">, 5</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="6D484F934E7BBB75D8623CB5D3DBAA83" pageId="0" pageNumber="137" type="type material">
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<paragraph id="878F377D3F47C14D127A5838214C4023" pageId="0" pageNumber="137">Type material.</paragraph>
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<paragraph id="17B4958D8AD5C038F14B9497FAA76F86" pageId="0" pageNumber="137">
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<emphasis id="4489F806F057A980122F6C3677CC7561" bold="true" italics="true" pageId="0" pageNumber="137">Holotype</emphasis>
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♂ (CUMZ-CAM186), Cambodia, Kep Province, Damnak
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<normalizedToken id="6E3273F5C12474267BAEFE610E4F8CD1" originalValue="Chang’aeur">Chang'aeur</normalizedToken>
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District, Ou Krasar, Phnom Sorsia Temple (locality code C047),
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<geoCoordinate id="89E504860EC5CF15D74001DA2BC351B5" degrees="10" direction="north" minutes="33" orientation="latitude" precision="15" seconds="54" value="10.565001">10°33'54"N</geoCoordinate>
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,
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<geoCoordinate id="E7E3BC0B07D8B59004F0AF71F61E8F38" degrees="104" direction="east" minutes="17" orientation="longitude" precision="15" seconds="2" value="104.28389">104°17'2"E</geoCoordinate>
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, 16.09.2019, leg. E. Jeratthitikul and R. Srisonchai.
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</paragraph>
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<paragraph id="134514A1ED63F1ADF93BC522456CFE3A" pageId="0" pageNumber="137">
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<emphasis id="405E60BDA0115863B32E0AF378C386F5" bold="true" pageId="0" pageNumber="137">
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<emphasis id="83E5C6DC5811B2FC93FCF6A83173EA61" italics="true" pageId="0" pageNumber="137">Paratypes</emphasis>
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.
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</emphasis>
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16 ♂, 17 ♀, 6 juveniles (CUMZ-CAM186), 2 ♂, 2 ♀ (ZMUM), 2 ♂, 2 ♀ (NHMD), 2 ♂, 2 ♀ (ZRC), same locality, together with holotype.
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</paragraph>
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</subSubSection>
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<subSubSection id="DA91572CE4AFFE2007421C89F7873626" pageId="0" pageNumber="137" type="name">
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<paragraph id="AB744D2AF001909235A83D95D29FE19D" pageId="0" pageNumber="137">Name.</paragraph>
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<paragraph id="62F8A97AADE85018CED721ED442A9DA3" pageId="0" pageNumber="137">
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To emphasize "
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<taxonomicName id="6272B572798BD44450B8E9474DCE22EF" authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer">khmer</taxonomicName>
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", referring to the main people of Cambodia; a noun in apposition.
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</paragraph>
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</subSubSection>
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<subSubSection id="A331E34625A3250224CB6BA569C8F5E0" pageId="0" pageNumber="137" type="diagnosis">
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<paragraph id="798DBC4E9298AE99D6329EB83A5D1118" pageId="0" pageNumber="137">Diagnosis.</paragraph>
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<paragraph id="6CB64B06A6A2CB4A372DD61A236DEAEE" pageId="0" pageNumber="137">
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This new species differs from all congeners by all crests on the collum being undivided, mostly complete (Fig.
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<figureCitation id="5661A52EAD3DC34ED3A288D483922090" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5B</figureCitation>
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) and male legs 1 with very long and one-segmented telopodites (Fig.
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<figureCitation id="6CFA03D04E8733FC33ECD59CEE995CA5" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5D, E</figureCitation>
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), as well as by the presence of 2+2 paramedian, characteristically long, slender, coxosternal processes of the anterior gonopods, a shorter and only apically curved posterior (
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<emphasis id="F270ECB6989972831550766F40F80DBE" bold="true" pageId="0" pageNumber="137">cxp1</emphasis>
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) pair, and a longer, mesally micropapillate and regularly curved anterior (
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<emphasis id="EF7269297BA912948482F6B338BFA259" bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
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) one (Fig.
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<figureCitation id="3BAE27843C1625EB4A7C1094DFCB0B8A" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5H, I</figureCitation>
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); the posterior gonopods are oblong-subtrapeziform, membranous, each with a small apical spike (
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<emphasis id="CCF8C573D8DDC3C5291A3A860235A582" bold="true" pageId="0" pageNumber="137">d</emphasis>
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) and a bifid process (
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<emphasis id="E871A8E8080A69B8DA3BDA2483638B7A" bold="true" pageId="0" pageNumber="137">k</emphasis>
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) at about 1/3 telopodite length on anterior face (Fig.
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<figureCitation id="59FFBE243618D17EA15F45E5A32FCAC0" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5J, K</figureCitation>
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).
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</paragraph>
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</subSubSection>
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||
<subSubSection id="47B72958E872E9A884F6BB432BF0DEEC" pageId="0" pageNumber="137" type="description">
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<paragraph id="901B7ABDFE74B949F1E7B363117C5F80" pageId="0" pageNumber="137">Description.</paragraph>
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<paragraph id="E3333C2C8F971903A0FB8C544062CD6D" pageId="0" pageNumber="137">
|
||
<emphasis id="EFE7BF8B90BC561A132D516D3BE6FD49" bold="true" italics="true" pageId="0" pageNumber="137">Length</emphasis>
|
||
of holotype ca 26 mm; adult paratypes 19.3-25.4 (♂) or 14.1-27.0 mm (♀), juveniles 8.5-11.5 mm long; midbody segments round in cross-section (Fig.
|
||
<figureCitation id="D831752CFF5F8E7B5722555F505593A4" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4F</figureCitation>
|
||
), their width (horizontal diameter) shorter than height (vertical diameter), width in holotype 1.2 mm; paratypes 1.1-1.5 (♂), 1.1-1.6 mm (♀) or 0.8-1.1 mm (juveniles).
|
||
</paragraph>
|
||
<caption id="4F8355D0FCDD28189250CCAC019A09B3" doi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137" start="Figure 4" startId="F4">
|
||
<paragraph id="CC8422266A691CB7C487D7592494EABB" pageId="0" pageNumber="137">
|
||
<emphasis id="83B28FCDF578D167465C8316EE45BC96" bold="true" pageId="0" pageNumber="137">Figure 4.</emphasis>
|
||
<taxonomicName id="9ABC6B2FC30A825634C5A4DBB207E048" authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusioglyphiulus khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer">
|
||
<emphasis id="7D00B948F9A6E1BB1590D4FC21F6A867" italics="true" pageId="0" pageNumber="137">Plusioglyphiulus khmer</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., ♂ paratype
|
||
<emphasis id="026323ACDDE95E36DBA6AA4A6E322E42" bold="true" pageId="0" pageNumber="137">A-C</emphasis>
|
||
anterior part of body, lateral, dorsal and ventral views, respectively
|
||
<emphasis id="3015B1A2D3529B2CB181364134BCAB7B" bold="true" pageId="0" pageNumber="137">D, E</emphasis>
|
||
midbody segments, dorsal and lateral views, respectively
|
||
<emphasis id="A5CBFD6E4122933C215453225042D92C" bold="true" pageId="0" pageNumber="137">F</emphasis>
|
||
cross-section of a midbody segment
|
||
<emphasis id="131B93430065766F7FEB2EDEB09588F6" bold="true" pageId="0" pageNumber="137">G-I</emphasis>
|
||
posterior part of body, lateral, dorsal and ventral views, respectively.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="A3617F20010E7D48AC43361463BE1529" pageId="0" pageNumber="137">
|
||
<emphasis id="EF0220324152C7F60A68FE25C433EE04" bold="true" italics="true" pageId="0" pageNumber="137">Coloration</emphasis>
|
||
of live animals light brown to chocolate brown (Fig.
|
||
<figureCitation id="2C4CFDC8DC242122C8D992CF1DD5967E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">1C</figureCitation>
|
||
) with blackish or dark brown anterior and posterior parts of body; antennae light brown to brown; venter and legs light yellowish to yellowish white; rather contrasting dark brownish, lateral, longitudinal stripes above ozopores on each side, both interrupted mid-dorsally by a light wide axial stripe; ommatidia brown to blackish (Fig.
|
||
<figureCitation id="F2490BA6CAEDA0DE266A6819277FCB0F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, C</figureCitation>
|
||
); coloration in alcohol, after six months of preservation (Fig.
|
||
<figureCitation id="838F0938393DBD82E4031AC6D1F79A98" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4</figureCitation>
|
||
), similar to live one, but body red-brownish or dark castaneous brown to grey-brown; vertex dark brown to yellow-brown; antennae dark brown to blackish; venter and legs light yellowish to yellowish red (Fig.
|
||
<figureCitation id="CD5DF86A2E1844EE7495AA420AB4AC54" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A-C, E-G, I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="1DA4A73EFF7BC48F35FC9AC4D282EFC2" pageId="0" pageNumber="137">
|
||
<emphasis id="8DC8F68E01F306D831FF6636F0D0C36A" bold="true" italics="true" pageId="0" pageNumber="137">Adult body</emphasis>
|
||
with 53p+2a+T (holotype); paratypes with 46-57p+1-3a+T (♂), 44-55p+2-4a+T (♀) or 35-42+3-4a +T (juveniles). Eye patches transversely ovoid, with 11-18 ommatidia arranged in three or four longitudinal rows (adult) (Fig.
|
||
<figureCitation id="7299D1ADEF5FE4DE46101660A34645B2" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, C</figureCitation>
|
||
) or with 6-10 ommatidia in three longitudinal rows (juveniles). Clypeus with three teeth anteromedially (Fig.
|
||
<figureCitation id="8E05696634D3D7FDCE692F366DE7139F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6B0B438733D2156C917AB49E12D2E7B2" pageId="0" pageNumber="137">
|
||
<emphasis id="78B4DD42DCBE654214936B1A22F3DD6C" bold="true" italics="true" pageId="0" pageNumber="137">Antennae</emphasis>
|
||
short and clavate (Figs
|
||
<figureCitation id="1529E255F0652BBDA8607EC2FE8E1140" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">1C</figureCitation>
|
||
,
|
||
<figureCitation id="34D0A6B1BB2B8085D8AA5F0872B070E6" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, C</figureCitation>
|
||
,
|
||
<figureCitation id="9B777324BE9C2D1E0FA85180C4457F11" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5C</figureCitation>
|
||
), extending behind segment 3 laterally, antennomeres 5 and 6 each with a small apicodorsal field or corolla of bacilliform sensilla (Fig.
|
||
<figureCitation id="225BB53A8EDF71F57AE376249BA21A6D" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5C</figureCitation>
|
||
). Gnathochilarium oligotrichous, each lamella lingualis with two or three setae; promentum bare, separated from eumentum by a distinct transverse suture (
|
||
<emphasis id="6E7BF745AF99C5472FC99C16AE50C45E" italics="true" pageId="0" pageNumber="137">n</emphasis>
|
||
= 2) (Fig.
|
||
<figureCitation id="3F87CFB1E234C82A4867A2E5EAEDAA0B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="1E043BD167717300561B7D757A8294B5" doi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137" start="Figure 5" startId="F5">
|
||
<paragraph id="1DE0FEDCF5C5C14E00A54DF0982A3A11" pageId="0" pageNumber="137">
|
||
<emphasis id="20EF286C557B745E31AAFCE83B5A5A83" bold="true" pageId="0" pageNumber="137">Figure 5.</emphasis>
|
||
<taxonomicName id="272D49DAA8CA0B7A6DFC088B7370F0A6" authorityName="Likhitrakarn & Golovatch & Thach & Chhuoy & Ngor & Srisonchai & Sutcharit & Panha" authorityYear="2020" class="Diplopoda" family="Cambalopsidae" genus="Plusioglyphiulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Plusioglyphiulus khmer" order="Spirostreptida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="khmer">
|
||
<emphasis id="9266A5C0A2648FDC0F509A8D1BAA1BAF" italics="true" pageId="0" pageNumber="137">Plusioglyphiulus khmer</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.
|
||
<emphasis id="8588F4C1811E4EF86F86697627F5A06D" bold="true" pageId="0" pageNumber="137">A, B</emphasis>
|
||
♂ paratype
|
||
<emphasis id="394BB622A0DAD22339D9E8C26D279F9F" bold="true" pageId="0" pageNumber="137">C-L</emphasis>
|
||
♂ holotype
|
||
<emphasis id="FD6EF77D428967156C1D5D40AC0C6B60" bold="true" pageId="0" pageNumber="137">A</emphasis>
|
||
gnathochilarium, ventral view
|
||
<emphasis id="C8E394F9F300D37D66676F9A3CE69DAD" bold="true" pageId="0" pageNumber="137">B</emphasis>
|
||
collum, dorsal view
|
||
<emphasis id="75060B4D095BDA1A49616964E010E26C" bold="true" pageId="0" pageNumber="137">C</emphasis>
|
||
antenna, lateral view
|
||
<emphasis id="6D32CC9E8EAA4C4A0A18F6CB48304A23" bold="true" pageId="0" pageNumber="137">D, E</emphasis>
|
||
♂ legs 1, anterior and posterior views, respectively
|
||
<emphasis id="31665B8BF6AA79F2E98A82CB3B0B93B4" bold="true" pageId="0" pageNumber="137">F</emphasis>
|
||
♂ legs 2, posterior view
|
||
<emphasis id="12AB3808C28B72B2463A20808ACBEC2C" bold="true" pageId="0" pageNumber="137">G</emphasis>
|
||
♂ legs 3, posterior view
|
||
<emphasis id="090EC9CC625145838B31D898D8BAC1E4" bold="true" pageId="0" pageNumber="137">H, I</emphasis>
|
||
anterior gonopods, anterior and posterior views, respectively
|
||
<emphasis id="E4425C700C632E15353E5E0D69C3EE6E" bold="true" pageId="0" pageNumber="137">J, K</emphasis>
|
||
posterior gonopods, posterior and anterior views, respectively
|
||
<emphasis id="614EF187CE3B69A48456A1A4543DA920" bold="true" pageId="0" pageNumber="137">L</emphasis>
|
||
midbody leg, anterior view. Abbreviations:
|
||
<emphasis id="E0DC2542CC633F12746477D1F5D26A2D" bold="true" pageId="0" pageNumber="137">cxp1</emphasis>
|
||
1st coxosternal process
|
||
<emphasis id="9300A2808322A07205F38907EF7C249C" bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
|
||
2nd coxosternal process
|
||
<emphasis id="EE98AB44DC88AE6AB4781098DCFD0397" bold="true" pageId="0" pageNumber="137">te</emphasis>
|
||
telopodites
|
||
<emphasis id="33990AF91BC25004352B829B9C3C25D5" bold="true" pageId="0" pageNumber="137">d</emphasis>
|
||
terminal medial spike
|
||
<emphasis id="88E56008F0E303D13EB00D8A406BA82B" bold="true" pageId="0" pageNumber="137">k</emphasis>
|
||
a small bifid process. Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="FF880A9E8381098E270A108ABBF1AE71" pageId="0" pageNumber="137">
|
||
<taxonomicName id="B3DCA29E617D023EC1E74B58689BA501" genus="Postcollum" lsidName="Postcollum" pageId="0" pageNumber="137" rank="genus">
|
||
<emphasis id="666E36A2E5E105979A8F0B93E3670050" bold="true" italics="true" pageId="0" pageNumber="137">Postcollum</emphasis>
|
||
</taxonomicName>
|
||
constriction evident, but collum only moderately enlarged (Figs
|
||
<figureCitation id="C06D86406C69E4F08FE7BCAE92816AAC" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus, live coloration A Plusioglyphiulus boutini Mauries, 1970, ♂ from Prasat Phnom Totong Temple B Plusioglyphiulus biserratus sp. nov., ♀ paratype C Plusioglyphiulus khmer sp. nov., ♂ paratype. All pictures by R. Srisonchai, not taken to scale." figureDoi="10.3897/zookeys.938.51234.figure1" httpUri="https://binary.pensoft.net/fig/418283" pageId="0" pageNumber="137">1C</figureCitation>
|
||
,
|
||
<figureCitation id="BFD23E32F9D680DAF08C35A511C0646E" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A-C</figureCitation>
|
||
). Carinotaxic formula of collum: 1+ta+2+3+4+ta+5(5p/t)+ta+P+ma (Figs
|
||
<figureCitation id="39586900E82A95DC5845128EBC015227" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A-C</figureCitation>
|
||
,
|
||
<figureCitation id="FB1629FCD395B72EBD3677CB99762F4B" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5B</figureCitation>
|
||
). Carinotaxy of metatergum 2, 8/8+m/m+8/8; of metaterga 3 and 4, 7/7+m/m+7/7 (Fig.
|
||
<figureCitation id="510F8F5407306773D3750898FE4CF0DA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B</figureCitation>
|
||
); formula of metaterga 5 and following metaterga, except last few, usually 3/3+I/i+3/3/3+m/m+3/3/3+I/i+3/3 (Fig.
|
||
<figureCitation id="0725FB8F27FECE570A0D86F18A93FC42" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B, D-H</figureCitation>
|
||
); of legless segments, usually 7+m+7 (Fig.
|
||
<figureCitation id="6F4CCCB32BD2918CCCF84311B8E260F7" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4G, H</figureCitation>
|
||
); all crests and tubercles, including poriferous cones, rather low. Dorsal crests on several posteriormost segments lower than others (Fig.
|
||
<figureCitation id="14D501F3618C1B6A28A8166F6DB35EFB" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B, D-H</figureCitation>
|
||
). Porosteles large, low, conical, round, directed caudolaterad, wider than high (Fig.
|
||
<figureCitation id="46F528E7D34EFB6B40BC726C70D30AED" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4D-H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0DD92BB1F6A57750C8F71F2FD962AE48" pageId="0" pageNumber="137">
|
||
<emphasis id="6EDCB19771B4D5ABD496A3503C32D8EC" bold="true" italics="true" pageId="0" pageNumber="137">Tegument</emphasis>
|
||
finely alveolate-areolate (Fig.
|
||
<figureCitation id="DA42112363C7C50FDAC6E4272EB8A59B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A, B, D, E, G, H</figureCitation>
|
||
), dull throughout. Metatergal setae absent. Pleural regions of segments 2-4 conspicuously elongated, flap-shaped, especially well so on segment 3 (Fig.
|
||
<figureCitation id="C17C19107EDD814297F247F3D6A0EB4A" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4A</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="CE3EC3B85BD39F344A9AA9307CA1F6E8" pageId="0" pageNumber="137">
|
||
<taxonomicName id="7DD79AD6D452200530ED91A8284615D3" genus="Epiproct" lsidName="Epiproct" pageId="0" pageNumber="137" rank="genus">
|
||
<emphasis id="99A2A6BD42325C678871E7B72755FC96" bold="true" italics="true" pageId="0" pageNumber="137">Epiproct</emphasis>
|
||
</taxonomicName>
|
||
(Fig.
|
||
<figureCitation id="8CEA8FCAD185A096953652DD6824A255" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4G-I</figureCitation>
|
||
) broadly rounded apically, with 2+2 small, paramedian tubercles at midway. Paraprocts rather clearly flattened, each with a faint premarginal sulcus medially (Fig.
|
||
<figureCitation id="783960ED35F8B6AA4BC4A34DFA439663" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4G, I</figureCitation>
|
||
). Hypoproct clearly emarginate at caudal margin (Fig.
|
||
<figureCitation id="378794738274BCB6975A7A4127A5D52D" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4I</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="04AFB2B4D61A6F53860CC77F4D775DFD" pageId="0" pageNumber="137">
|
||
<emphasis id="82325AC2B8312AB72C85BD53742E60FA" bold="true" italics="true" pageId="0" pageNumber="137">Ventral</emphasis>
|
||
flaps behind gonopod aperture on male segment 7 evident, distinguishable as rather high swellings with rounded flaps bent abruptly caudad (Fig.
|
||
<figureCitation id="FB96A62CE6A31598B76554EE74271166" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="681B2AEB01CFA52CB42D979B15370323" pageId="0" pageNumber="137">
|
||
<emphasis id="242772DD026E2A62152E1B16BA0AF744" bold="true" italics="true" pageId="0" pageNumber="137">Legs</emphasis>
|
||
short, on midbody segments about 1/3 as long as body diameter (Figs
|
||
<figureCitation id="44188EAEE754F6EF2A2FF2C8B321184F" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Plusioglyphiulus khmer sp. nov., ♂ paratype A-C anterior part of body, lateral, dorsal and ventral views, respectively D, E midbody segments, dorsal and lateral views, respectively F cross-section of a midbody segment G-I posterior part of body, lateral, dorsal and ventral views, respectively." figureDoi="10.3897/zookeys.938.51234.figure4" httpUri="https://binary.pensoft.net/fig/418286" pageId="0" pageNumber="137">4E-G</figureCitation>
|
||
,
|
||
<figureCitation id="EAC5B9BE70AE6C36000D359A99DFC3ED" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5L</figureCitation>
|
||
), claw at base with a strong, spiniform, accessory claw about 1/3 as long as claw itself (Fig.
|
||
<figureCitation id="ECA99CE34BFDBB06FAF402CC86FBD396" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5L</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="0220FA8C2F245AAA368F6C349B51E35E" pageId="0" pageNumber="137">
|
||
Male legs 1 with a usual, strong, central hook (actually a double structure of tightly appressed hooks), regularly curved forward; a pair of 1-segmented, sac-shaped and very long telopodites, the latter almost as long as central hook (Fig.
|
||
<figureCitation id="A1C1BDA388400CEC9F098C8BFC966B7F" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5D, E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="76F2666853BDBE8390C00DF08F54F814" pageId="0" pageNumber="137">
|
||
Male legs 2 strongly enlarged, with very high and large coxae; telopodites hirsute on anterior face; penes subconical, truncate apically, fused at base (Fig.
|
||
<figureCitation id="68E85242B2D8F24A9A4467EF3A3A667C" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="6F423EA3563AD29E1578E42F171B57FE" pageId="0" pageNumber="137">
|
||
Male legs 3 modified as usual, with particularly elongate and slender coxae and shortened telopodites (Fig.
|
||
<figureCitation id="3A849740DB8D4E8769AE6D407509B757" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="680E183E7040FA904E186734B673653E" pageId="0" pageNumber="137">
|
||
<emphasis id="927D0D1B66DAC77250F45606E542DE2C" bold="true" italics="true" pageId="0" pageNumber="137">Anterior gonopods</emphasis>
|
||
(Fig.
|
||
<figureCitation id="9D26E8D7CFF78166B57ADDF4943D7300" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5H, I</figureCitation>
|
||
) rather complex, with 2+2 paramedian, long, slender, coxosternal processes: a shorter and only apically curved posterior (
|
||
<emphasis id="778E4B7FC926416781245217731A1D49" bold="true" pageId="0" pageNumber="137">cxp1</emphasis>
|
||
) pair, and a longer, mesally micropapillate and regularly curved anterior (
|
||
<emphasis id="BD840557566953BE93F72CD2D916CB05" bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
|
||
) one (Fig.
|
||
<figureCitation id="EA7BE5CB927855FAD0162E5C15627DE7" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5H, I</figureCitation>
|
||
); telopodites (
|
||
<emphasis id="69E00E6E5BF856BCDDA9D26B990AA973" bold="true" pageId="0" pageNumber="137">te</emphasis>
|
||
) finger-shaped, subcylindrical, long, apically setose, about as high as
|
||
<emphasis id="1EC3972B456E8DC039A57A532F2EA584" bold="true" pageId="0" pageNumber="137">cxp2</emphasis>
|
||
, attached to coxal region caudolaterally, probably capable of movement.
|
||
</paragraph>
|
||
<paragraph id="9BB6DEBB7820EBCCCA1EDD82A08F2D16" pageId="0" pageNumber="137">
|
||
<emphasis id="86EABA8B747B28DADBECD27D6DAF53DF" bold="true" italics="true" pageId="0" pageNumber="137">Posterior gonopods</emphasis>
|
||
(Fig.
|
||
<figureCitation id="EDB583CBD3960425F889589A0AACE6F6" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Plusioglyphiulus khmer sp. nov. A, B ♂ paratype C-L ♂ holotype A gnathochilarium, ventral view B collum, dorsal view C antenna, lateral view D, E ♂ legs 1, anterior and posterior views, respectively F ♂ legs 2, posterior view G ♂ legs 3, posterior view H, I anterior gonopods, anterior and posterior views, respectively J, K posterior gonopods, posterior and anterior views, respectively L midbody leg, anterior view. Abbreviations: cxp 1 1 st coxosternal process cxp 2 2 nd coxosternal process te telopodites d terminal medial spike k a small bifid process. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.938.51234.figure5" httpUri="https://binary.pensoft.net/fig/418287" pageId="0" pageNumber="137">5J, K</figureCitation>
|
||
) rather long (high), simple; coxites well-separated from sternum, fused only basally, oblong-subtrapeziform, membranous, with traces of telopodites in the form of a small, latero-parabasal field of microsetae on posterior face of each gonopod; each coxite with a terminal medial spike (
|
||
<emphasis id="D483C7F4317B91FE0BCBA8E6E7A56A9B" bold="true" pageId="0" pageNumber="137">d</emphasis>
|
||
), a subterminal lobule at
|
||
<emphasis id="B54481E33B2A67D996B3A9844B3BDA82" bold="true" pageId="0" pageNumber="137">d</emphasis>
|
||
base, and a small bifid process (
|
||
<emphasis id="E042C787C96C23AF33CABE5A23C86A98" bold="true" pageId="0" pageNumber="137">k</emphasis>
|
||
) at about 1/3 coxite length on frontal face; distal half lamellose, fringed and with a deep fovea subapically.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5703334D8A00FC101F68459058AA679F" pageId="0" pageNumber="137" type="remarks">
|
||
<paragraph id="B22D8AAF5A7D41D647B2DE9ECFB7AF84" pageId="0" pageNumber="137">Remarks.</paragraph>
|
||
<paragraph id="0171C5B00B132428A0B3147468A09594" pageId="0" pageNumber="137">
|
||
This species was found together with a single male specimen of
|
||
<taxonomicName id="70B559D144614212A4FBEDB4696B4CBB" baseAuthorityName="Saussure" baseAuthorityYear="1860" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha coarctata" order="Polydesmida" pageId="0" pageNumber="137" phylum="Arthropoda" rank="species" species="coarctata">
|
||
<emphasis id="B4F4DCA65911796B278E932192CC3D48" italics="true" pageId="0" pageNumber="137">Orthomorpha coarctata</emphasis>
|
||
</taxonomicName>
|
||
(De Saussure, 1860).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |