181 lines
18 KiB
XML
181 lines
18 KiB
XML
<document id="153EA1CEECA87D2A3D5A1D201A5B47F7" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FFAF4A7BFF328E66FC52EE3E" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole musinermis Longino 2019, new species" docType="treatment" docVersion="22" lastPageNumber="49" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="48" updateTime="1732580243751" updateUser="ExternalLinkService">
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<mods:mods id="F244283919FD403DCDC11DF6B83CF2F1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="44AC05C62E5FCB0D970C0D77A32C1211">
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<mods:title id="61A509B1C964167F14191F2941BD836C">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
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</mods:titleInfo>
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<mods:name id="A9D83C52F8E6F4E05B9FED4346B8AEB3" type="personal">
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<mods:role id="57A960DB18DBD036AFFAF25192BA034F">
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<mods:roleTerm id="A4D4EC48CD4F2336CD8A6DEA6D0FAAC2">Author</mods:roleTerm>
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<mods:namePart id="B7C1BE00A9337590BD18FF95D09FE7A6">Longino, John T.</mods:namePart>
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<mods:typeOfResource id="A5602DF2EB0D5D505037BE448414A6A9">text</mods:typeOfResource>
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<mods:title id="B5509C0117E2A164CF8CA507DBD2F5B1">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="0C172DB3EA2A253C69C17331CAD61462">
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<mods:date id="B4F93491EF638D0D5A657C4D74CA6D93">2019</mods:date>
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<mods:detail id="7B4CC8A8B4CAF6B7D226FE9FBFC42E45" type="pubDate">
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<mods:number id="29C5581BA5866A3813BE5AA91056F7FE">2019-05-06</mods:number>
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<mods:detail id="07131FA1F8962CD77F1E4C701A3C0D31" type="volume">
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<mods:number id="3876F8E0B6F3C25E6E1E0D5E11AABD3B">4599</mods:number>
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<mods:detail id="6983F2E5D95A32B9C30C8D9772ED71EF" type="issue">
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<mods:number id="0C84A84E2B98DB9401B02704E83945DF">1</mods:number>
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<mods:classification id="B8949AFF3CFFB927C5F0856AD8B473FF">journal article</mods:classification>
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<mods:identifier id="B0F5AEC0BDE1BEA9BB981B2E977B92BF" type="CLB-Dataset">26920</mods:identifier>
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<mods:identifier id="F8B5E566AE4E18937DFC7C3E3C40F9B2" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
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<mods:identifier id="AE9A203825D458BAAEB9CB20829E7190" type="GBIF-Dataset">ea727a9b-324b-4c6b-a625-bc7cdd204473</mods:identifier>
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<mods:identifier id="E9501AC6BF0D709B17671C60394D49DF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="A5A717A33E52DB5A6B6B122795815DAF" type="Zenodo-Dep">2668058</mods:identifier>
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<mods:identifier id="E99C84F9173D94843FAB29F17F0EAA1E" type="ZooBank">7CDD24FF-5BA3-4D06-898D-A3E220515D4C</mods:identifier>
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</mods:mods>
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<treatment id="5B56E872FFAF4A7BFF328E66FC52EE3E" ID-GBIF-Taxon="157062630" LSID="urn:lsid:plazi:treatment:5B56E872FFAF4A7BFF328E66FC52EE3E" httpUri="http://treatment.plazi.org/id/5B56E872FFAF4A7BFF328E66FC52EE3E" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" scope_family="Formicidae" scope_order="Hymenoptera">
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<subSubSection id="9BE50AEFFFAF4A64FF328E66FEF1EA1F" pageId="47" pageNumber="48" type="nomenclature">
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<paragraph id="D3405964FFAF4A64FF328E66FDD9EA78" blockId="47.[151,557,1289,1349]" box="[151,557,1289,1315]" pageId="47" pageNumber="48">
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<heading id="8808EE08FFAF4A64FF328E66FDD9EA78" bold="true" box="[151,557,1289,1315]" fontSize="11" level="1" pageId="47" pageNumber="48" reason="1">
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<emphasis id="E18B8576FFAF4A64FF328E66FDD9EA78" bold="true" box="[151,557,1289,1315]" pageId="47" pageNumber="48">
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<taxonomicName id="14FF22E7FFAF4A64FF328E66FE62EA78" ID-CoL="CMLKC" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,406,1289,1315]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="musinermis" status="sp. nov.">
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<emphasis id="E18B8576FFAF4A64FF328E66FE62EA78" bold="true" box="[151,406,1289,1315]" italics="true" pageId="47" pageNumber="48">Pheidole musinermis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FAB8380DFFAF4A64FE388E66FDD9EA78" box="[413,557,1289,1315]" pageId="47" pageNumber="48" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D3405964FFAF4A64FF328E43FEF1EA1F" blockId="47.[151,557,1289,1349]" box="[151,261,1324,1349]" pageId="47" pageNumber="48">
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(
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<figureCitation id="4BC445E1FFAF4A64FF3B8E43FF09EA1F" box="[158,253,1324,1349]" captionStart="PLATE 42" captionStartId="108.[151,238,1954,1977]" captionTargetId="figure@108.[202,1384,194,1933]" captionTargetPageId="108" captionText="PLATE 42." httpUri="https://zenodo.org/record/2668317/files/figure.png" pageId="47" pageNumber="48">Plate 42</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF328E1BFC6FEAD6" box="[151,923,1396,1421]" pageId="47" pageNumber="48" type="description">
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<paragraph id="D3405964FFAF4A64FF328E1BFC6FEAD6" blockId="47.[151,1436,1396,2033]" box="[151,923,1396,1421]" pageId="47" pageNumber="48">
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<taxonomicName id="14FF22E7FFAF4A64FF328E1BFF0FEAD6" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,1396,1421]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="genus">
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<emphasis id="E18B8576FFAF4A64FF328E1BFF0FEAD6" box="[151,251,1396,1421]" italics="true" pageId="47" pageNumber="48">Pheidole</emphasis>
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</taxonomicName>
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JTL-123: morphospecies code previously used on AntWeb.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF628EF7FD61E964" pageId="47" pageNumber="48" type="materials_examined">
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<paragraph id="D3405964FFAF4A64FF628EF7FEADE947" blockId="47.[151,1436,1396,2033]" pageId="47" pageNumber="48">
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<materialsCitation id="63975339FFAF4A64FF628EF7FE87EAA3" ID-GBIF-Occurrence="2237851595" collectingDate="2007-02-06" collectingDateMax="2015-07-06" collectingDateMin="2007-02-06" collectionCode="MUCR" country="Costa Rica" elevation="0" location="Cerro Plano" pageId="47" pageNumber="48" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
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<typeStatus id="0C44E7C6FFAF4A64FF628EF7FEABEAEB" box="[199,351,1432,1456]" pageId="47" pageNumber="48" type="holotype">HOLOTYPE</typeStatus>
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: 1 major worker,
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<collectingCountry id="ABE819F4FFAF4A64FD8C8EF7FD53EAEB" box="[553,679,1431,1456]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
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,
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<collectingRegion id="113B9786FFAF4A64FD118EF7FCE8EAEB" box="[692,796,1432,1456]" country="Costa Rica" name="San Jose" pageId="47" pageNumber="48">San José</collectingRegion>
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:
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<location id="D6200FBFFFAF4A64FC828EF7FC46EAEB" LSID="urn:lsid:plazi:treatment:5B56E872FFAF4A7BFF328E66FC52EE3E:D6200FBFFFAF4A64FC828EF7FC46EAEB" box="[807,946,1431,1456]" country="Costa Rica" name="Cerro Plano" pageId="47" pageNumber="48" stateProvince="San Jose">Cerro Plano</location>
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, 9.48117 -83.96363 ±
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<elevation id="58D2BE57FFAF4A64FB0E8EF8FABAEAF4" box="[1195,1358,1431,1456]" isEnumeration="true" pageId="47" pageNumber="48">
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<quantity id="1407F481FFAF4A64FB0E8EF8FB1CEAF4" box="[1195,1256,1431,1456]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="47" pageNumber="48" unit="m" value="50.0">50 m</quantity>
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,
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<quantity id="1407F481FFAF4A64FB508EF8FABAEAF4" box="[1269,1358,1431,1456]" metricMagnitude="3" metricUnit="m" metricValue="1.06" pageId="47" pageNumber="48" unit="m" value="1060.0">1060 m</quantity>
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</elevation>
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,
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<date id="A7417FA4FFAF4A64FAFE8EF8FF26EA8E" pageId="47" pageNumber="48" value="2015-07-06">
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<collectingDate id="B705864CFFAF4A64FAFE8EF8FF26EA8E" pageId="47" pageNumber="48" value="2015-07-06">6-Jul- 2015</collectingDate>
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</date>
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, ridgetop cloud forest, at bait (ADMAC, Ba-E-
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<date id="A7417FA4FFAF4A64FCE38ED3FC36EA8E" box="[838,962,1468,1493]" pageId="47" pageNumber="48" value="2007-02-06" valueMax="2007-04-06" valueMin="2007-02-06">
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<collectingDate id="B705864CFFAF4A64FCE38ED3FC36EA8E" box="[838,962,1468,1493]" pageId="47" pageNumber="48" value="2007-02-06" valueMax="2007-04-06" valueMin="2007-02-06">06-2-04-07</collectingDate>
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</date>
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) [
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<collectionCode id="B5EEC1A1FFAF4A64FC408ED3FBC9EA8F" box="[997,1085,1468,1492]" country="Costa Rica" name="Museo de Insectos" pageId="47" pageNumber="48">MUCR</collectionCode>
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, unique specimen identifier CASENT0631280]
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</materialsCitation>
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.
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<materialsCitation id="63975339FFAF4A64FE248E8FFEA1E947" ID-GBIF-Occurrence="2237851544" collectingDate="2007-02-06" collectingDateMax="2015-07-06" collectingDateMin="2007-02-06" collectionCode="CAS, DZUP, JTLC, MCZC, MUCR, USNM" country="Costa Rica" elevation="0" location="Cerro Plano" pageId="47" pageNumber="48" specimenCount="1" stateProvince="San Jose" typeStatus="paratype">
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<typeStatus id="0C44E7C6FFAF4A64FE248E8FFDD4EAA3" box="[385,544,1504,1528]" pageId="47" pageNumber="48" type="paratype">PARATYPES</typeStatus>
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: major, minor workers: same data as holotype [
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<collectionCode id="B5EEC1A1FFAF4A64FBF48E8FFB78EAA3" box="[1105,1164,1504,1528]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="47" pageNumber="48" type="Museum">CAS</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FB398E8FFB11EAAC" box="[1180,1253,1504,1527]" country="Brazil" httpUri="http://grbio.org/cool/5xp9-edpx" name="Universidade Federal do Parana, Colecao de Entomologia Pe. Jesus Santiago Moure" pageId="47" pageNumber="48" type="University or college">DZUP</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FB518E8FFAC3EAA3" box="[1268,1335,1504,1528]" pageId="47" pageNumber="48">JTLC</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FAE38E8FFA6CEAA3" box="[1350,1432,1504,1528]" pageId="47" pageNumber="48">MCZC</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FF328D6BFF19E947" box="[151,237,1540,1564]" country="Costa Rica" name="Museo de Insectos" pageId="47" pageNumber="48">MUCR</collectionCode>
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,
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<collectionCode id="B5EEC1A1FFAF4A64FF528D6BFEBAE947" box="[247,334,1540,1564]" country="USA" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="47" pageNumber="48" type="Museum">USNM</collectionCode>
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]
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="D3405964FFAF4A64FF628D48FD61E964" blockId="47.[151,1436,1396,2033]" box="[199,661,1575,1600]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628D48FE51E91B" bold="true" box="[199,421,1575,1600]" pageId="47" pageNumber="48">Geographic range.</emphasis>
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<collectingCountry id="ABE819F4FFAF4A64FE098D47FDDDE91B" box="[428,553,1575,1600]" name="Costa Rica" pageId="47" pageNumber="48">Costa Rica</collectingCountry>
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,
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<collectingCountry id="ABE819F4FFAF4A64FD918D47FD64E964" box="[564,656,1576,1599]" name="Panama" pageId="47" pageNumber="48">Panama</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF628D23FDD6E83B" pageId="47" pageNumber="48" type="diagnosis">
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<paragraph id="D3405964FFAF4A64FF628D23FDD6E83B" blockId="47.[151,1436,1396,2033]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628D23FE62E93F" bold="true" box="[199,406,1611,1637]" pageId="47" pageNumber="48">
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Diagnosis.
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<emphasis id="E18B8576FFAF4A64FEE88D24FE62E93F" bold="true" box="[333,406,1611,1636]" italics="true" pageId="47" pageNumber="48">Minor</emphasis>
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</emphasis>
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: face smooth and shining; head strongly tapering posteriorly; occipital carina strongly developed; promesonotal groove present, distinctly impressed; lateral pronotum smooth and shiny; katepisternum faintly foveate; propodeal spines nearly absent; gaster smooth and shining; abundant, flexuous, erect setae on mesosomal dorsum, tibiae, and gastral dorsum; color yellow.
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<emphasis id="E18B8576FFAF4A64FCDD8DD8FC34E98B" bold="true" box="[888,960,1719,1744]" italics="true" pageId="47" pageNumber="48">Major</emphasis>
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: inner hypostomal teeth widely spaced, acicular; scape base subterete, narrower than apical portion; face with irregular rugulae anteriorly, entire face with faint foveolate sculpture; promesonotal groove impressed; propodeal spines forming obtuse angles, not dentiform; gastral dorsum foveolate on anterior half; abundant, erect, flexuous setae on face, sides of head, mesosomal dorsum, tibiae, and gastral dorsum.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFAF4A64FF628C03FD9BE8AB" pageId="47" pageNumber="48" type="description">
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<paragraph id="D3405964FFAF4A64FF628C03FD89E8F3" blockId="47.[151,1436,1396,2033]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628C03FDD9E8DE" bold="true" box="[199,557,1900,1925]" pageId="47" pageNumber="48">Measurements, minor worker</emphasis>
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: HW 0.60, HL 0.87, SL 1.37, EL 0.17, WL 1.23, PSL 0.00, PTW 0.13, PPW 0.21, CI 69, SI 229, PSLI 0, PPI 154 (n=3).
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</paragraph>
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<paragraph id="D3405964FFAF4A64FF628CDBFD9BE8AB" blockId="47.[151,1436,1396,2033]" pageId="47" pageNumber="48">
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<emphasis id="E18B8576FFAF4A64FF628CDBFDD9E896" bold="true" box="[199,557,1972,1997]" pageId="47" pageNumber="48">Measurements, major worker</emphasis>
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: HW 1.59, HL 1.70, SL 1.32, EL 0.25, WL 1.78, PSL 0.02, PTW 0.31, PPW 0.45, CI 94, SI 83, PSLI 2, PPI 147 (n=2).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFB04A7BFF628BF8FA77EF8E" pageId="48" pageNumber="49" type="biology_ecology">
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<paragraph id="D3405964FFB04A7BFF628BF8FA77EF8E" blockId="48.[151,1436,151,357]" pageId="48" pageNumber="49">
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<emphasis id="E18B8576FFB04A7BFF628BF8FED3EFEB" bold="true" box="[199,295,151,176]" pageId="48" pageNumber="49">Biology.</emphasis>
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This species occurs in lower montane wet forest. Most collections are from ground baits, to which minors and majors recruit. Minor workers have been collected in a pan trap and a Winkler sample of sifted litter.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BE50AEFFFB04A7BFF628BB0FE94EE1B" pageId="48" pageNumber="49" type="discussion">
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<paragraph id="D3405964FFB04A7BFF628BB0FE94EE1B" blockId="48.[151,1436,151,357]" pageId="48" pageNumber="49">
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<emphasis id="E18B8576FFB04A7BFF628BB0FEBCEFA3" bold="true" box="[199,328,223,248]" pageId="48" pageNumber="49">Comments</emphasis>
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. This species is part of a complex of large, yellow species that occur in
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<collectingCountry id="ABE819F4FFB04A7BFBD08B8FFB07EFA3" box="[1141,1267,223,248]" name="Costa Rica" pageId="48" pageNumber="49">Costa Rica</collectingCountry>
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. It is similar to
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<taxonomicName id="14FF22E7FFB04A7BFF328A6AFEB7EE46" authorityName="Longino" authorityYear="2019" box="[151,323,260,285]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="familiaparra">
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<emphasis id="E18B8576FFB04A7BFF328A6AFEB7EE46" box="[151,323,260,285]" italics="true" pageId="48" pageNumber="49">P. familiaparra</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="14FF22E7FFB04A7BFEF78A6AFE36EE47" authorityName="Longino" authorityYear="2019" box="[338,450,261,284]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="savegre">
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<emphasis id="E18B8576FFB04A7BFEF78A6AFE36EE47" box="[338,450,261,284]" italics="true" pageId="48" pageNumber="49">P. savegre</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="14FF22E7FFB04A7BFE718A6AFD92EE46" authorityName="Longino" authorityYear="2019" box="[468,614,260,285]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="musacolor">
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<emphasis id="E18B8576FFB04A7BFE718A6AFD92EE46" box="[468,614,260,285]" italics="true" pageId="48" pageNumber="49">P. musacolor</emphasis>
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</taxonomicName>
|
|
, and
|
|
<taxonomicName id="14FF22E7FFB04A7BFD038A6AFCE5EE47" authorityName="Longino" authorityYear="2019" box="[678,785,261,284]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="tinamu">
|
|
<emphasis id="E18B8576FFB04A7BFD038A6AFCE5EE47" box="[678,785,261,284]" italics="true" pageId="48" pageNumber="49">P. tinamu</emphasis>
|
|
</taxonomicName>
|
|
, but is distinguishable by the lack of propodeal spines in the minor worker.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="9BE50AEFFFB04A7BFF628A23FC52EE3E" box="[199,934,332,357]" pageId="48" pageNumber="49" type="etymology">
|
|
<paragraph id="D3405964FFB04A7BFF628A23FC52EE3E" blockId="48.[151,1436,151,357]" box="[199,934,332,357]" pageId="48" pageNumber="49">
|
|
<emphasis id="E18B8576FFB04A7BFF628A23FEBCEE3E" bold="true" box="[199,328,332,357]" pageId="48" pageNumber="49">Etymology</emphasis>
|
|
: The color of a banana, and lacking propodeal spines.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |