treatments-xml/data/5B/56/E8/5B56E872FF984A52FF328FA0FD20EDB4.xml

236 lines
24 KiB
XML

<document id="5E4C85632B9946301826D361D4CBF968" ID-CLB-Dataset="26920" ID-DOI="10.11646/zootaxa.4599.1.1" ID-GBIF-Dataset="ea727a9b-324b-4c6b-a625-bc7cdd204473" ID-ISSN="1175-5326" ID-Zenodo-Dep="2668058" ID-ZooBank="7CDD24FF-5BA3-4D06-898D-A3E220515D4C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1557127209543" checkinUser="plazi" docAuthor="Longino, John T." docDate="2019" docId="5B56E872FF984A52FF328FA0FD20EDB4" docLanguage="en" docName="zootaxa.4599.1.1.pdf.imf" docOrigin="Zootaxa 4599 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pheidole belonorte Longino 2019, new species" docType="treatment" docVersion="22" lastPageNumber="26" masterDocId="A76F900AFF804A4BFFA58B6FFFF4EF5B" masterDocTitle="Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest" masterLastPageNumber="126" masterPageNumber="1" pageNumber="25" updateTime="1732580243751" updateUser="ExternalLinkService">
<mods:mods id="2A24F61CD6FF52645C08F6E889656969" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8FCC31E25A8DEE1424AF69C56B226669">
<mods:title id="CE4E9E864E5C24BBE0D695468E741A21">Pheidole (Hymenoptera, Formicidae) of Middle American Wet Forest</mods:title>
</mods:titleInfo>
<mods:name id="6DB7BF8AA3A7824B03A018845F742AB6" type="personal">
<mods:role id="4E23804DF266DC2904AAA9F7DB051725">
<mods:roleTerm id="B22061064EA1DBB37EDE790F4309E809">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B15B55B6C8CC995F7824856523211B88">Longino, John T.</mods:namePart>
</mods:name>
<mods:typeOfResource id="6CC93D4822A1A0D44C42AC890574105C">text</mods:typeOfResource>
<mods:relatedItem id="91C19668303EA794ED79A282C985F936" type="host">
<mods:titleInfo id="35B17EEAD38F190F2DB4A475291C8EEB">
<mods:title id="D86A19BC7E375C3D8B56781799B94D2F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="672C2EB6AC8F60188D0635DB632502D3">
<mods:date id="6EEC36CBDA668D900FFA6EFE760CE3C5">2019</mods:date>
<mods:detail id="487C0E6C089AB6CD0C9C7BDF47BB1840" type="pubDate">
<mods:number id="ED97372508BCFCE63D9E2252BA70F9FB">2019-05-06</mods:number>
</mods:detail>
<mods:detail id="51EEE8B76A42EE59C5CB4ECAE3C6CD34" type="volume">
<mods:number id="8FBA3146131D51E869204E975540C8F1">4599</mods:number>
</mods:detail>
<mods:detail id="F37C0B5743922AB12E0CD48D64FAAED8" type="issue">
<mods:number id="F6E19FB703BA839843E814D3D6E90866">1</mods:number>
</mods:detail>
<mods:extent id="86F3E081BF187964DBE56FB131585C72" unit="page">
<mods:start id="861E49E82106CC263A22769AF970554D">1</mods:start>
<mods:end id="569CDC6F1B0FF9BCA49D91C20F02DCDB">126</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5E5853BB5C58849DFE2A301060E2A979">journal article</mods:classification>
<mods:identifier id="E168342867B59332CBB236273D839398" type="CLB-Dataset">26920</mods:identifier>
<mods:identifier id="449588C6E42885671A4C54DE95AC5157" type="DOI">10.11646/zootaxa.4599.1.1</mods:identifier>
<mods:identifier id="C5A61279B9B877413D150A71A171E282" type="GBIF-Dataset">ea727a9b-324b-4c6b-a625-bc7cdd204473</mods:identifier>
<mods:identifier id="FB5438460AFDE8A0A1BA551D79CD0CC6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="FB33402421D1EB7C0B2889216378C500" type="Zenodo-Dep">2668058</mods:identifier>
<mods:identifier id="D7DE36C8F3EF4B3D59D24F31B9F147F4" type="ZooBank">7CDD24FF-5BA3-4D06-898D-A3E220515D4C</mods:identifier>
</mods:mods>
<treatment id="5B56E872FF984A52FF328FA0FD20EDB4" ID-GBIF-Taxon="157062590" LSID="urn:lsid:plazi:treatment:5B56E872FF984A52FF328FA0FD20EDB4" httpUri="http://treatment.plazi.org/id/5B56E872FF984A52FF328FA0FD20EDB4" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" scope_family="Formicidae" scope_order="Hymenoptera">
<subSubSection id="9BE50AEFFF984A53FF328FA0FF02EA50" pageId="24" pageNumber="25" type="nomenclature">
<paragraph id="D3405964FF984A53FF328FA0FDE5EBB1" blockId="24.[151,529,1231,1291]" box="[151,529,1231,1258]" pageId="24" pageNumber="25">
<heading id="8808EE08FF984A53FF328FA0FDE5EBB1" bold="true" box="[151,529,1231,1258]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<emphasis id="E18B8576FF984A53FF328FA0FDE5EBB1" bold="true" box="[151,529,1231,1258]" pageId="24" pageNumber="25">
<taxonomicName id="14FF22E7FF984A53FF328FA0FE8EEBB2" ID-CoL="CMKWM" authority="Longino, 2019" authorityName="Longino" authorityYear="2019" box="[151,378,1231,1257]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="belonorte" status="sp. nov.">
<emphasis id="E18B8576FF984A53FF328FA0FE8EEBB2" bold="true" box="[151,378,1231,1257]" italics="true" pageId="24" pageNumber="25">Pheidole belonorte</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAB8380DFF984A53FE248FBFFDE5EBB1" box="[385,529,1232,1258]" pageId="24" pageNumber="25" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D3405964FF984A53FF328F9DFF02EA50" blockId="24.[151,529,1231,1291]" box="[151,246,1266,1291]" pageId="24" pageNumber="25">
(
<figureCitation id="4BC445E1FF984A53FF3B8F9DFF19EA50" box="[158,237,1266,1291]" captionStart="PLATE 8" captionStartId="74.[151,238,1954,1977]" captionTargetId="figure@74.[202,1384,194,1933]" captionTargetPageId="74" captionText="PLATE 8." httpUri="https://zenodo.org/record/2668243/files/figure.png" pageId="24" pageNumber="25">Plate 8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF984A53FF328E55FC6FEA08" box="[151,923,1338,1363]" pageId="24" pageNumber="25" type="description">
<paragraph id="D3405964FF984A53FF328E55FC6FEA08" blockId="24.[151,1437,1338,2011]" box="[151,923,1338,1363]" pageId="24" pageNumber="25">
<taxonomicName id="14FF22E7FF984A53FF328E55FF0FEA08" authorityName="Biolleyi Forel" authorityYear="1908" box="[151,251,1338,1363]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="genus">
<emphasis id="E18B8576FF984A53FF328E55FF0FEA08" box="[151,251,1338,1363]" italics="true" pageId="24" pageNumber="25">Pheidole</emphasis>
</taxonomicName>
JTL-225: morphospecies code previously used on AntWeb.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF984A53FF628E30FD54E95C" pageId="24" pageNumber="25" type="materials_examined">
<paragraph id="D3405964FF984A53FF628E30FE98EAB8" blockId="24.[151,1437,1338,2011]" pageId="24" pageNumber="25">
<materialsCitation id="63975339FF984A53FF628E30FE87EAE4" ID-GBIF-Occurrence="2237851478" collectingDate="2002-04-04" collectingDateMax="2010-05-10" collectingDateMin="2002-04-04" collectionCode="MCZC" country="Honduras" elevation="0" location="Catacamas" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Olancho" typeStatus="holotype">
<typeStatus id="0C44E7C6FF984A53FF628E30FEABEA2C" box="[199,351,1375,1399]" pageId="24" pageNumber="25" type="holotype">HOLOTYPE</typeStatus>
: 1 major worker,
<collectingCountry id="ABE819F4FF984A53FD8F8E31FD6EEA2C" box="[554,666,1374,1399]" name="Honduras" pageId="24" pageNumber="25">Honduras</collectingCountry>
,
<collectingRegion id="113B9786FF984A53FD028E31FCFEEA2C" box="[679,778,1374,1399]" country="Honduras" name="Olancho" pageId="24" pageNumber="25">Olancho</collectingRegion>
:
<quantity id="1407F481FF984A53FCB28E31FCBCEA2C" box="[791,840,1374,1399]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="24" pageNumber="25" unit="km" value="9.0">9km</quantity>
N
<location id="D6200FBFFF984A53FCCB8E30FC1EEA2C" LSID="urn:lsid:plazi:treatment:5B56E872FF984A52FF328FA0FD20EDB4:D6200FBFFF984A53FCCB8E30FC1EEA2C" box="[878,1002,1375,1399]" country="Honduras" name="Catacamas" pageId="24" pageNumber="25" stateProvince="Olancho">Catacamas</location>
, 14.93833 -85.90315 ±
<elevation id="58D2BE57FF984A53FB518E31FA63EA2D" box="[1268,1431,1374,1399]" isEnumeration="true" pageId="24" pageNumber="25">
<quantity id="1407F481FF984A53FB518E31FAC5EA2D" box="[1268,1329,1374,1399]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="24" pageNumber="25" unit="m" value="50.0">50 m</quantity>
,
<quantity id="1407F481FF984A53FA9B8E30FA63EA2D" box="[1342,1431,1375,1399]" metricMagnitude="3" metricUnit="m" metricValue="1.29" pageId="24" pageNumber="25" unit="m" value="1290.0">1290 m</quantity>
</elevation>
,
<date id="A7417FA4FF984A53FF328EECFEC0EAC0" box="[151,308,1411,1435]" pageId="24" pageNumber="25" value="2010-05-10">
<collectingDate id="B705864CFF984A53FF328EECFEC0EAC0" box="[151,308,1411,1435]" pageId="24" pageNumber="25" value="2010-05-10">10-May-2010</collectingDate>
</date>
, mixed hardwood forest, at bait (LLAMA, Ba-C-
<date id="A7417FA4FF984A53FCD08EECFC04EAC0" box="[885,1008,1411,1435]" pageId="24" pageNumber="25" value="2002-04-04" valueMax="2002-12-04" valueMin="2002-04-04">
<collectingDate id="B705864CFF984A53FCD08EECFC04EAC0" box="[885,1008,1411,1435]" pageId="24" pageNumber="25" value="2002-04-04" valueMax="2002-12-04" valueMin="2002-04-04">02-4-04-12</collectingDate>
</date>
) [
<collectionCode id="B5EEC1A1FF984A53FBAC8EECFBAAEAC0" box="[1033,1118,1411,1435]" pageId="24" pageNumber="25">MCZC</collectionCode>
, unique specimen identifier CASENT0613794]
</materialsCitation>
.
<typeStatus id="0C44E7C6FF984A53FED98EC8FDEFEAE4" box="[380,539,1447,1471]" pageId="24" pageNumber="25" type="paratype">PARATYPES</typeStatus>
: major, minor workers:
<materialsCitation id="63975339FF984A53FC838EC8FB73EAE4" ID-GBIF-Occurrence="2237851519" box="[806,1159,1446,1471]" collectingDate="2002-04-04" collectingDateMax="2010-05-10" collectingDateMin="2002-04-04" collectionCode="MCZC" country="Honduras" elevation="0" location="Catacamas" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Olancho" typeStatus="paratype">
same data as holotype [
<collectionCode id="B5EEC1A1FF984A53FB8C8EC8FB75EAE4" box="[1065,1153,1447,1471]" pageId="24" pageNumber="25">MCZC</collectionCode>
]
</materialsCitation>
;
<materialsCitation id="63975339FF984A53FB378EC8FE9CEAB8" ID-GBIF-Occurrence="2237851572" collectingDate="2003-02-04" collectingDateMax="2010-05-10" collectingDateMin="2002-04-04" collectionCode="CAS" country="Honduras" elevation="0" location="Catacamas" pageId="24" pageNumber="25" specimenCount="1" stateProvince="Olancho" typeStatus="paratype">
same data except Ba-C-
<date id="A7417FA4FF984A53FF328EA4FEE1EAB8" box="[151,277,1483,1507]" pageId="24" pageNumber="25" value="2003-02-04" valueMax="2003-02-04" valueMin="2003-02-04">
<collectingDate id="B705864CFF984A53FF328EA4FEE1EAB8" box="[151,277,1483,1507]" pageId="24" pageNumber="25" value="2003-02-04" valueMax="2003-02-04" valueMin="2003-02-04">02-4-04-03</collectingDate>
</date>
[
<collectionCode id="B5EEC1A1FF984A53FE868EA4FE95EAB8" box="[291,353,1483,1507]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="24" pageNumber="25" type="Museum">CAS</collectionCode>
]
</materialsCitation>
.
</paragraph>
<paragraph id="D3405964FF984A53FF628E81FD54E95C" blockId="24.[151,1437,1338,2011]" box="[199,672,1518,1543]" pageId="24" pageNumber="25">
<emphasis id="E18B8576FF984A53FF628E81FE51E95C" bold="true" box="[199,421,1518,1543]" pageId="24" pageNumber="25">Geographic range.</emphasis>
<collectingCountry id="ABE819F4FF984A53FE098E81FDF5E95C" box="[428,513,1518,1543]" name="Mexico" pageId="24" pageNumber="25">Mexico</collectingCountry>
to
<collectingCountry id="ABE819F4FF984A53FD808E81FD68E95C" box="[549,668,1518,1543]" name="Nicaragua" pageId="24" pageNumber="25">Nicaragua</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF984A53FF628D7DFF22E8C9" pageId="24" pageNumber="25" type="diagnosis">
<paragraph id="D3405964FF984A53FF628D7DFF22E8C9" blockId="24.[151,1437,1338,2011]" pageId="24" pageNumber="25">
<emphasis id="E18B8576FF984A53FF628D7DFE6FE970" bold="true" box="[199,411,1554,1579]" pageId="24" pageNumber="25">
Diagnosis.
<emphasis id="E18B8576FF984A53FEF78D7DFE6FE970" bold="true" box="[338,411,1554,1579]" italics="true" pageId="24" pageNumber="25">Minor</emphasis>
</emphasis>
: face foveolate, overlain with reticulate rugulae; posterior margin of head weakly emarginate in face view; promesonotal groove absent; pronotal dorsum with faint foveolation overlain with weak rugulae; lateral pronotum smooth and shiny; katepisternum foveolate; propodeal spines short, upturned, about one fourth length of posterior face of propodeum; gaster smooth and shining; erect setae on mesosomal dorsum and gaster; tibiae without erect setae; color yellow brown.
<emphasis id="E18B8576FF984A53FCB18DCDFCA8E9E0" bold="true" box="[788,860,1698,1723]" italics="true" pageId="24" pageNumber="25">Major</emphasis>
: head elongate, subrectangular; inner hypostomal teeth stout, closely spaced; scape base terete; face with very shallow scrobal impressions, not delimited; anterior half of face with faint foveolation, overlain with subparallel, longitudinal carinae between eye and antennal fossa and between frontal carinae, posterior face and vertex lobes smooth and shiny; propodeal spines short, upturned, about one fifth length of posterior face of propodeum; gastral dorsum smooth and shining; sides of head with abundant short erect setae; abundant short erect setae on mesosomal dorsum and gaster; tibiae with 0-3 suberect setae.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF984A52FF628CF1FD9BEF8E" lastPageId="25" lastPageNumber="26" pageId="24" pageNumber="25" type="description">
<paragraph id="D3405964FF984A53FF628CF1FD89E880" blockId="24.[151,1437,1338,2011]" pageId="24" pageNumber="25">
<emphasis id="E18B8576FF984A53FF628CF1FDD9E8EC" bold="true" box="[199,557,1950,1975]" pageId="24" pageNumber="25">Measurements, minor worker</emphasis>
: HW 0.43, HL 0.48, SL 0.36, EL 0.07, WL 0.52, PSL 0.05, PTW 0.08, PPW 0.13, CI 89, SI 85, PSLI 12, PPI 157 (n=5).
</paragraph>
<paragraph id="D3405964FF994A52FF628BF8FD9BEF8E" blockId="25.[151,1436,151,752]" pageId="25" pageNumber="26">
<emphasis id="E18B8576FF994A52FF628BF8FDD9EFEB" bold="true" box="[199,557,151,176]" pageId="25" pageNumber="26">Measurements, major worker</emphasis>
: HW 0.79, HL 0.97, SL 0.40, EL 0.11, WL 0.76, PSL 0.06, PTW 0.15, PPW 0.30, CI 81, SI 51, PSLI 8, PPI 196 (n=5).
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF994A52FF628BB0FF27EE1B" pageId="25" pageNumber="26" type="biology_ecology">
<paragraph id="D3405964FF994A52FF628BB0FF27EE1B" blockId="25.[151,1436,151,752]" pageId="25" pageNumber="26">
<emphasis id="E18B8576FF994A52FF628BB0FED3EFA3" bold="true" box="[199,295,223,248]" pageId="25" pageNumber="26">Biology.</emphasis>
This species inhabits mature lowland wet forest and lower cloud forest. Minor and major workers recruit to ground baits and are collected in Winkler samples of sifted leaf litter and rotten wood from the forest floor.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF994A52FF628A23FB66ED96" pageId="25" pageNumber="26" type="discussion">
<paragraph id="D3405964FF994A52FF628A23FB66ED96" blockId="25.[151,1436,151,752]" pageId="25" pageNumber="26">
<emphasis id="E18B8576FF994A52FF628A23FEBCEE3E" bold="true" box="[199,328,332,357]" pageId="25" pageNumber="26">Comments</emphasis>
.
<taxonomicName id="14FF22E7FF994A52FEFC8A23FDC5EE3E" authorityName="Longino" authorityYear="2019" box="[345,561,332,357]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="belonorte">
<emphasis id="E18B8576FF994A52FEFC8A23FDC5EE3E" box="[345,561,332,357]" italics="true" pageId="25" pageNumber="26">Pheidole belonorte</emphasis>
</taxonomicName>
is a northern version of
<taxonomicName id="14FF22E7FF994A52FCF08A22FC22EE3E" authorityName="Wilson" authorityYear="2003" box="[853,982,332,357]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FCF08A22FC22EE3E" box="[853,982,332,357]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
. DNA barcoding puts Costa Rican (La Selva)
<taxonomicName id="14FF22E7FF994A52FF408A1EFE97EED3" authorityName="Wilson" authorityYear="2003" box="[229,355,367,392]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FF408A1EFE97EED3" box="[229,355,367,392]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
and Nicaraguan
<taxonomicName id="14FF22E7FF994A52FD878A1EFD5EEED3" authorityName="Longino" authorityYear="2019" box="[546,682,367,392]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="belonorte">
<emphasis id="E18B8576FF994A52FD878A1EFD5EEED3" box="[546,682,367,392]" italics="true" pageId="25" pageNumber="26">P. belonorte</emphasis>
</taxonomicName>
in different clusters.
<taxonomicName id="14FF22E7FF994A52FC3D8A00FB9AEED3" authorityName="Longino" authorityYear="2019" box="[920,1134,367,392]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="belonorte">
<emphasis id="E18B8576FF994A52FC3D8A00FB9AEED3" box="[920,1134,367,392]" italics="true" pageId="25" pageNumber="26">Pheidole belonorte</emphasis>
</taxonomicName>
has the major worker with larger, more rectangular head, with vertex lobes smooth and shiny. This species occurs elsewhere in
<collectingCountry id="ABE819F4FF994A52FA878AFBFA6CEEF6" box="[1314,1432,404,429]" name="Nicaragua" pageId="25" pageNumber="26">Nicaragua</collectingCountry>
,
<collectingCountry id="ABE819F4FF994A52FF328AD8FEF3EE8B" box="[151,263,439,464]" name="Honduras" pageId="25" pageNumber="26">Honduras</collectingCountry>
, and
<collectingRegion id="113B9786FF994A52FEE18AD8FE55EE8B" box="[324,417,439,464]" country="Mexico" name="Chiapas" pageId="25" pageNumber="26">Chiapas</collectingRegion>
,
<collectingCountry id="ABE819F4FF994A52FE0B8AD8FDF3EE8B" box="[430,519,439,464]" name="Mexico" pageId="25" pageNumber="26">Mexico</collectingCountry>
, but so far appears rare in
<collectingCountry id="ABE819F4FF994A52FC9E8AD8FC43EE8B" box="[827,951,439,464]" name="Guatemala" pageId="25" pageNumber="26">Guatemala</collectingCountry>
, with only one tentative record of a minor worker. The minors are indistinguishable from Costa Rican
<taxonomicName id="14FF22E7FF994A52FC968AB2FC44EEAE" authorityName="Wilson" authorityYear="2003" box="[819,944,476,501]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FC968AB2FC44EEAE" box="[819,944,476,501]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
. Among the mounted material, the only site where both species are known to occur is Cerro Musún in southern
<collectingCountry id="ABE819F4FF994A52FC0E8A90FBD6ED43" box="[939,1058,511,536]" name="Nicaragua" pageId="25" pageNumber="26">Nicaragua</collectingCountry>
. During quantitative sampling in 2011,
<taxonomicName id="14FF22E7FF994A52FF7B894AFEAAED66" authorityName="Wilson" authorityYear="2003" box="[222,350,548,573]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FF7B894AFEAAED66" box="[222,350,548,573]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
was in the miniWinkler transects at
<quantity id="1407F481FF994A52FCAC894BFC8FED67" box="[777,891,548,572]" metricMagnitude="2" metricUnit="m" metricValue="7.5" metricValueMax="8.0" metricValueMin="7.0" pageId="25" pageNumber="26" unit="m" value="750.0" valueMax="800.0" valueMin="700.0">700-800m</quantity>
elevation;
<taxonomicName id="14FF22E7FF994A52FC58894AFB73ED66" authorityName="Longino" authorityYear="2019" box="[1021,1159,548,573]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="belonorte">
<emphasis id="E18B8576FF994A52FC58894AFB73ED66" box="[1021,1159,548,573]" italics="true" pageId="25" pageNumber="26">P. belonorte</emphasis>
</taxonomicName>
was in higher elevation maxiWinklers,
<quantity id="1407F481FF994A52FEE18927FE23ED3B" box="[324,471,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.4" metricValueMin="1.0" pageId="25" pageNumber="26" unit="m" value="1200.0" valueMax="1400.0" valueMin="1000.0">1000-1400m</quantity>
. Previously, all specimens were identified as a more broadly defined
<taxonomicName id="14FF22E7FF994A52FB498926FA9EED3B" authorityName="Wilson" authorityYear="2003" box="[1260,1386,583,608]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FB498926FA9EED3B" box="[1260,1386,583,608]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
. On AntWeb, I reidentified all material north of Musún as
<taxonomicName id="14FF22E7FF994A52FD598902FC75EDDE" authorityName="Longino" authorityYear="2019" box="[764,897,620,645]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="belonorte">
<emphasis id="E18B8576FF994A52FD598902FC75EDDE" box="[764,897,620,645]" italics="true" pageId="25" pageNumber="26">P. belonorte</emphasis>
</taxonomicName>
, including material in ethanol that I did not reexamine. For Musún, I identified all material below
<quantity id="1407F481FF994A52FCA389FFFCB1EDF3" box="[774,837,656,680]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="25" pageNumber="26" unit="m" value="800.0">800m</quantity>
as
<taxonomicName id="14FF22E7FF994A52FCD189FEFC02EDF3" authorityName="Wilson" authorityYear="2003" box="[884,1014,655,680]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FCD189FEFC02EDF3" box="[884,1014,655,680]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
; all material above as
<taxonomicName id="14FF22E7FF994A52FAAF89FEFA66EDF3" authorityName="Longino" authorityYear="2019" box="[1290,1426,655,680]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="belonorte">
<emphasis id="E18B8576FF994A52FAAF89FEFA66EDF3" box="[1290,1426,655,680]" italics="true" pageId="25" pageNumber="26">P. belonorte</emphasis>
</taxonomicName>
. Ethanol material should be re-examined, although minors cannot be separated at this point.
</paragraph>
</subSubSection>
<subSubSection id="9BE50AEFFF994A52FF6289B8FD20EDB4" box="[199,724,727,752]" pageId="25" pageNumber="26" type="etymology">
<paragraph id="D3405964FF994A52FF6289B8FD20EDB4" blockId="25.[151,1436,151,752]" box="[199,724,727,752]" pageId="25" pageNumber="26">
<emphasis id="E18B8576FF994A52FF6289B8FEBCEDAB" bold="true" box="[199,328,727,752]" pageId="25" pageNumber="26">Etymology</emphasis>
: A northern version of
<taxonomicName id="14FF22E7FF994A52FDEA89B6FD39EDAB" authorityName="Wilson" authorityYear="2003" box="[591,717,727,752]" class="Insecta" family="Formicidae" genus="Pheidole" kingdom="Animalia" order="Hymenoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="beloceps">
<emphasis id="E18B8576FF994A52FDEA89B6FD39EDAB" box="[591,717,727,752]" italics="true" pageId="25" pageNumber="26">P. beloceps</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>