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<document id="6A5FE1327AFE94FD423D4CB5A46E0EC9" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFFC6145FF026640FA17FF02" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madecorphnus hanskii Frolov 2012" docType="treatment" docVersion="16" lastPageNumber="47" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="46" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="9F658E7B2865C7A8504120C038971E18">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:namePart id="1D2637055AAB7A59251BE13F073EBBF5">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="8219FCB73C5BB00725F35D5E31F935F8">Montreuil, Olivier</mods:namePart>
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<mods:namePart id="0C0A8AC883D6850C3B7B0977207D6ADF">Akhmetova, Lilia A.</mods:namePart>
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<subSubSection id="9BFC5F9DFFFC6147FF026640FEBCFBA2" pageId="45" pageNumber="46" type="nomenclature">
<paragraph id="D3590C16FFFC6147FF026640FDDAFBC3" blockId="45.[151,592,1052,1111]" box="[151,592,1052,1078]" pageId="45" pageNumber="46">
<heading id="8811BB7AFFFC6147FF026640FDDAFBC3" bold="true" box="[151,592,1052,1078]" fontSize="11" level="1" pageId="45" pageNumber="46" reason="1">
<taxonomicName id="14E67795FFFC6147FF026640FDDAFBC3" authority="Frolov, 2012" authorityName="Frolov" authorityYear="2012" box="[151,592,1052,1078]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFC6147FF026640FDDAFBC3" bold="true" box="[151,592,1052,1078]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF026640FE24FBC3" bold="true" box="[151,430,1052,1078]" italics="true" pageId="45" pageNumber="46">Madecorphnus hanskii</emphasis>
Frolov, 2012
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="D3590C16FFFC6147FF026662FEBCFBA2" blockId="45.[151,592,1052,1111]" box="[151,310,1086,1111]" pageId="45" pageNumber="46">
(
<figureCitation id="4BDD1093FFFC6147FF0A6662FF73FBA2" box="[159,249,1086,1111]" captionStart="FIGURE 23" captionStartId="46.[151,250,1740,1762]" captionTargetBox="[241,1341,405,1690]" captionTargetId="figure@46.[227,1360,265,1718]" captionTargetPageId="46" captionText="FIGURE 23. Madecorphnus hanskii, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="45" pageNumber="46">Figs. 23</figureCitation>
AF)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6147FF0266DAFDC7FB69" box="[151,589,1158,1180]" pageId="45" pageNumber="46" type="reference_group">
<paragraph id="D3590C16FFFC6147FF0266DAFDC7FB69" blockId="45.[151,589,1158,1180]" box="[151,589,1158,1180]" pageId="45" pageNumber="46">
<treatmentCitationGroup id="F3F62B38FFFC6147FF0266DAFDC7FB69" box="[151,589,1158,1180]" pageId="45" pageNumber="46">
<taxonomicName id="14E67795FFFC6147FF0266DAFDC3FB69" authority="Frolov, 2012b: 160" authorityName="Frolov" authorityPageNumber="160" authorityYear="2012" box="[151,585,1158,1180]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFC6147FF0266DAFEF4FB69" box="[151,382,1158,1180]" italics="true" pageId="45" pageNumber="46">Madecorphnus hanskii</emphasis>
<treatmentCitation id="52472A07FFFC6147FE1166DAFDC3FB69" author="Frolov" box="[388,585,1158,1180]" page="160" pageId="45" pageNumber="46" year="2012">
<bibRefCitation id="B77771E7FFFC6147FE1166DAFD9CFB69" author="Frolov" box="[388,534,1158,1180]" pageId="45" pageNumber="46" refString="Frolov, A. V. (2012 b) Three new species of scarab beetle genus Madecorphnus Paulian, 1992 (Coleoptera: Scarabaeidae: Orphninae) from Madagascar. Proceedings of the Zoological Institute RAS, 316, 159 - 165." type="journal article" year="2012" yearSuffix="b">Frolov, 2012b</bibRefCitation>
: 160
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6147FF026695FB17FAF3" pageId="45" pageNumber="46" type="materials_examined">
<paragraph id="D3590C16FFFC6147FF026695FB17FAF3" blockId="45.[151,1436,1225,2006]" pageId="45" pageNumber="46">
<materialsCitation id="638E064BFFFC6147FF026695FB13FAF3" ID-GBIF-Occurrence="1416610297" collectingDate="2005-03-24" collectionCode="ZIN" collectorName="I. Hanski-group &amp; Madec." country="Madagascar" latitude="-18.166666" location="Madagascar" longLatPrecision="1276" longitude="47.283333" pageId="45" pageNumber="46" specimenCount="3" typeStatus="holotype">
<emphasis id="E192D004FFFC6147FF026695FE41FB17" bold="true" box="[151,459,1225,1250]" pageId="45" pageNumber="46">
<typeStatus id="0C5DB2B4FFFC6147FF026695FF58FB17" box="[151,210,1225,1250]" pageId="45" pageNumber="46">Type</typeStatus>
material examined.
</emphasis>
<typeStatus id="0C5DB2B4FFFC6147FE4D6695FDCAFB17" box="[472,576,1225,1250]" pageId="45" pageNumber="46" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FFFC6147FDC36695FD32FB17" box="[598,696,1225,1250]" captionStart="FIGURE 23" captionStartId="46.[151,250,1740,1762]" captionTargetBox="[241,1341,405,1690]" captionTargetId="figure@46.[227,1360,265,1718]" captionTargetPageId="46" captionText="FIGURE 23. Madecorphnus hanskii, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="45" pageNumber="46">Figs. 23</figureCitation>
AE), male, “
<collectingCountry id="ABF14C86FFFC6147FCC86695FC6DFB17" box="[861,999,1225,1250]" name="Madagascar" pageId="45" pageNumber="46">Madagascar</collectingCountry>
Ambohitantely [
<geoCoordinate id="B6D26AD1FFFC6147FB246696FA93FB17" box="[1201,1305,1226,1250]" direction="south" orientation="latitude" pageId="45" pageNumber="46" precision="925" value="-18.166666">18°10' S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FFFC6147FAB96696FA1FFB14" box="[1324,1429,1226,1250]" direction="east" orientation="longitude" pageId="45" pageNumber="46" precision="925" value="47.283333">47°17' E</geoCoordinate>
]
<collectingDate id="B71CD33EFFFC6147FF0266B1FE80FAF3" box="[151,266,1261,1286]" pageId="45" pageNumber="46" value="2005-03-24">
<date id="A7582AD6FFFC6147FF0266B1FE80FAF3" box="[151,266,1261,1286]" pageId="45" pageNumber="46" value="2005-03-24">24.3.2005</date>
</collectingDate>
, wet forest,
<collectorName id="7E1369C0FFFC6147FE0766B2FDC9FAF3" box="[402,579,1261,1286]" pageId="45" pageNumber="46">I. Hanski-group</collectorName>
leg. /
<typeStatus id="0C5DB2B4FFFC6147FD1C66B1FD74FAF3" box="[649,766,1261,1286]" pageId="45" pageNumber="46" type="holotype">Holotypus</typeStatus>
<emphasis id="E192D004FFFC6147FC9066B1FCD3FAF3" box="[773,857,1261,1286]" italics="true" pageId="45" pageNumber="46">
<collectorName id="7E1369C0FFFC6147FC9066B1FCD3FAF3" box="[773,857,1261,1286]" pageId="45" pageNumber="46">Madec.</collectorName>
</emphasis>
<taxonomicName id="14E67795FFFC6147FCCA66B1FBC9FAF3" authority="Frolov 2009" authorityName="Frolov" authorityYear="2009" box="[863,1091,1261,1286]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFC6147FCCA66B1FC3AFAF3" box="[863,944,1261,1286]" italics="true" pageId="45" pageNumber="46">hanskii</emphasis>
<bibRefCitation id="B77771E7FFFC6147FC2266B1FBC9FAF3" author="Frolov" box="[951,1091,1261,1286]" pageId="45" pageNumber="46" refString="Frolov, A. V. &amp; Montreuil, O. (2009) A new genus of Orphninae (Coleoptera, Scarabaeidae) from Madagascar. Zoosystematica Rossica, 18, 65 - 69." type="journal article" year="2009">Frolov 2009</bibRefCitation>
</taxonomicName>
” (
<collectionCode id="B5F794D3FFFC6147FBC966B2FB1BFAF3" box="[1116,1169,1262,1286]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="45" pageNumber="46">ZIN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6147FF52674DFBA0FA60" pageId="45" pageNumber="46" type="diagnosis">
<paragraph id="D3590C16FFFC6147FF52674DFBA0FA60" blockId="45.[151,1436,1225,2006]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF52674DFECBFADF" bold="true" box="[199,321,1297,1322]" pageId="45" pageNumber="46">Diagnosis.</emphasis>
<taxonomicName id="14E67795FFFC6147FED9674DFDDAFADF" box="[332,592,1297,1322]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFC6147FED9674DFDDAFADF" box="[332,592,1297,1322]" italics="true" pageId="45" pageNumber="46">Madecorphnus hanskii</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="14E67795FFFC6147FD64674EFCF8FADF" box="[753,882,1297,1322]" class="Insecta" family="Scarabaeidae" genus="Sissantobius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="falciger">
<emphasis id="E192D004FFFC6147FD64674EFCF8FADF" box="[753,882,1297,1322]" italics="true" pageId="45" pageNumber="46">M. falciger</emphasis>
</taxonomicName>
and
<taxonomicName id="14E67795FFFC6147FC25674EFBCCFADC" box="[944,1094,1298,1321]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="punctatus">
<emphasis id="E192D004FFFC6147FC25674EFBCCFADC" box="[944,1094,1298,1321]" italics="true" pageId="45" pageNumber="46">M. punctatus</emphasis>
</taxonomicName>
in having internal sac of the aedeagus with 3 sclerites: 1 relatively long, spur-like, and 2 comma-shaped or somewhat bifurcate (
<figureCitation id="4BDD1093FFFC6147FA996769FAEDFABB" box="[1292,1383,1333,1358]" captionStart="FIGURE 16" captionStartId="35.[151,250,1841,1863]" captionTargetBox="[217,1388,194,1811]" captionTargetId="figure@35.[191,1396,193,1820]" captionTargetPageId="35" captionText="FIGURE 16. Madecorphnus punctatus, holotype. Habitus (A) and head (B) in dorsal view, aedeagus in lateral view and parameres in dorsal view (C), labels (D), internal sac of aedeagus (E—photograph, F—schematic representation of armature), distributional records (G)." httpUri="https://zenodo.org/record/272642/files/figure.png" pageId="45" pageNumber="46">Figs. 16</figureCitation>
EF, 22DE, 23DE). It can be separated from these species by the shape and relative size of these sclerites and, from the latter, by the shape of parameres and sparse punctation of head and pronotum.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFC6144FF5267FDFE28F83E" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46" type="description">
<paragraph id="D3590C16FFFC6147FF5267FDFBA2FA4F" blockId="45.[151,1436,1225,2006]" box="[199,1064,1441,1466]" pageId="45" pageNumber="46">
<emphasis id="E192D004FFFC6147FF5267FDFE29FA4F" bold="true" box="[199,419,1441,1466]" pageId="45" pageNumber="46">Description. Male.</emphasis>
Body length
<quantity id="141EA1F3FFFC6147FDAE67FDFD1EFA4C" box="[571,660,1441,1466]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" pageId="45" pageNumber="46" unit="mm" value="5.3">5.3 mm</quantity>
. Color uniformly brown (
<figureCitation id="4BDD1093FFFC6147FC2667FDFB8FFA4F" box="[947,1029,1441,1466]" captionStart="FIGURE 23" captionStartId="46.[151,250,1740,1762]" captionTargetBox="[241,1341,405,1690]" captionTargetId="figure@46.[227,1360,265,1718]" captionTargetPageId="46" captionText="FIGURE 23. Madecorphnus hanskii, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="45" pageNumber="46">Fig. 23</figureCitation>
A).
</paragraph>
<paragraph id="D3590C16FFFC6147FF526799FF75F99B" blockId="45.[151,1436,1225,2006]" pageId="45" pageNumber="46">Right mandible as long as left, without tooth behind apex. Labrum trapezoidal, with broadly rounded sides, length about 1/6 width (in dorsal view). Frontoclypeus slightly asymmetrical, apically obtuse, with 2 long and 6 shorter setae on the apical margin. Canthus and frontoclypeus almost flat, minutely depressed apicomedially. Head without traces of frontoclypeal suture, finely punctate with minute punctures separated by greater than 5 times their diameter.</paragraph>
<paragraph id="D3590C16FFFC6147FF526426FE7FF90B" blockId="45.[151,1436,1225,2006]" pageId="45" pageNumber="46">Pronotum approximately 1.6 times wider than long, widest medially. Disc of pronotum convex, without any depressions, tubercles, or ridges. Punctation on pronotum similar to that on head. Margins with relatively wide border, lateral margins with 4 long setae: 1 seta on basal angle, 1 seta approximately in the middle of lateral margin, and 2 setae on the apical angle.</paragraph>
<paragraph id="D3590C16FFFC6147FF526555FC56F8D7" blockId="45.[151,1436,1225,2006]" box="[199,988,1801,1826]" pageId="45" pageNumber="46">Scutellum triangular, right angled apically, about 1/13 length of elytra.</paragraph>
<paragraph id="D3590C16FFFC6147FF526571FC4FF87B" blockId="45.[151,1436,1225,2006]" pageId="45" pageNumber="46">Elytra convex, with distinct humeral humps, widest at basal third, glabrous. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with a few relatively large punctures. Epipleura with long, sparse, brown setae. Base of elytron with border. Wings fully developed.</paragraph>
<paragraph id="D3590C16FFFC6144FF5265C5FD56FF20" blockId="45.[151,1436,1225,2006]" lastBlockId="46.[151,1436,151,213]" lastPageId="46" lastPageNumber="47" pageId="45" pageNumber="46">Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Protibial apex with robust, spurlike seta and 2 smaller setae basally. Mesothoracic and metathoracic legs similar in shape to each other, metathoracic legs somewhat longer. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and longer than 2 basal tarsomeres in metathoracic legs.</paragraph>
<caption id="87995C9EFFFF6144FF026490FE87F8EA" httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="46" pageNumber="47" targetBox="[241,1341,405,1690]" targetPageId="46">
<paragraph id="D3590C16FFFF6144FF026490FE87F8EA" blockId="46.[151,1436,1740,1823]" pageId="46" pageNumber="47">
<emphasis id="E192D004FFFF6144FF026490FEABF917" bold="true" box="[151,289,1740,1762]" pageId="46" pageNumber="47">FIGURE 23.</emphasis>
<taxonomicName id="14E67795FFFF6144FEBD6490FD9BF917" box="[296,529,1740,1762]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFF6144FEBD6490FD9BF917" box="[296,529,1740,1762]" italics="true" pageId="46" pageNumber="47">Madecorphnus hanskii</emphasis>
</taxonomicName>
, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F).
</paragraph>
</caption>
<paragraph id="D3590C16FFFF6144FF526514FF33F852" blockId="46.[151,1436,1862,2032]" pageId="46" pageNumber="47">
Parameres with teeth laterally, narrowly rounded apically in dorsal view and curved downwards in lateral view (
<figureCitation id="4BDD1093FFFF6144FF0A6536FF78F876" box="[159,242,1898,1923]" captionStart="FIGURE 23" captionStartId="46.[151,250,1740,1762]" captionTargetBox="[241,1341,405,1690]" captionTargetId="figure@46.[227,1360,265,1718]" captionTargetPageId="46" captionText="FIGURE 23. Madecorphnus hanskii, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="46" pageNumber="47">Fig. 23</figureCitation>
B). Internal sac with a long, spur-like sclerite and 2 smaller, somewhat comma-shaped sclerites (
<figureCitation id="4BDD1093FFFF6144FA8A6536FAF3F876" box="[1311,1401,1898,1923]" captionStart="FIGURE 23" captionStartId="46.[151,250,1740,1762]" captionTargetBox="[241,1341,405,1690]" captionTargetId="figure@46.[227,1360,265,1718]" captionTargetPageId="46" captionText="FIGURE 23. Madecorphnus hanskii, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="46" pageNumber="47">Figs. 23</figureCitation>
D E).
</paragraph>
<paragraph id="D3590C16FFFF6144FF5265EEFE28F83E" blockId="46.[151,1436,1862,2032]" box="[199,418,1970,1995]" pageId="46" pageNumber="47">
<emphasis id="E192D004FFFF6144FF5265EEFEAFF83E" bold="true" box="[199,293,1970,1995]" pageId="46" pageNumber="47">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFFFF6145FF52658AFA17FF02" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="distribution">
<paragraph id="D3590C16FFFF6145FF52658AFA17FF02" blockId="46.[151,1436,1862,2032]" lastBlockId="47.[151,1437,151,248]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
<emphasis id="E192D004FFFF6144FF52658AFE72F81A" bold="true" box="[199,504,2006,2031]" pageId="46" pageNumber="47">Distribution and habitat.</emphasis>
The
<typeStatus id="0C5DB2B4FFFF6144FDAC6584FDE3F805" box="[569,617,2008,2032]" pageId="46" pageNumber="47">type</typeStatus>
specimen of
<taxonomicName id="14E67795FFFF6144FC9E6584FC0FF81A" box="[779,901,2006,2031]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFF6144FC9E6584FC0FF81A" box="[779,901,2006,2031]" italics="true" pageId="46" pageNumber="47">M. hanskii</emphasis>
</taxonomicName>
was collected in wet forest in Ambohitantely special reserve (
<figureCitation id="4BDD1093FFFE6145FEC762CBFE2DFF45" box="[338,423,151,176]" captionStart="FIGURE 23" captionStartId="46.[151,250,1740,1762]" captionTargetBox="[241,1341,405,1690]" captionTargetId="figure@46.[227,1360,265,1718]" captionTargetPageId="46" captionText="FIGURE 23. Madecorphnus hanskii, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200466/files/figure.png" pageId="47" pageNumber="48">Fig. 23</figureCitation>
F). The reserve is situated at altitudes of about
<quantity id="141EA1F3FFFE6145FC5862CBFBAFFF5A" box="[973,1061,151,176]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="47" pageNumber="48" unit="m" value="1500.0">1500 m</quantity>
and houses the last remnants of indigenous forest on the central plateau of
<collectingCountry id="ABF14C86FFFE6145FDEF62E0FC8EFF20" box="[634,772,188,213]" name="Madagascar" pageId="47" pageNumber="48">Madagascar</collectingCountry>
.
<taxonomicName id="14E67795FFFE6145FC9A62E0FB85FF20" box="[783,1039,188,213]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFFE6145FC9A62E0FB85FF20" box="[783,1039,188,213]" italics="true" pageId="47" pageNumber="48">Madecorphnus hanskii</emphasis>
</taxonomicName>
is apparently a forest litter dweller and, due to severe deforestation of the central plateau, its current range may be limited to this small forest remnant.
</paragraph>
</subSubSection>
</treatment>
</document>