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<document id="34F417D40CB1EB38C9A3EC1D18AD3C6D" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFE86153FF0263E4FEE3F95C" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triodontus" docType="key" docVersion="16" lastPageNumber="58" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="58" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:title id="25F094615BDD4DC61195429D7B26ABC2">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:namePart id="B82A2876E152CEF2307612D19FCEA06B">Akhmetova, Lilia A.</mods:namePart>
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<treatment id="5B4FBD00FFE86153FF0263E4FEE3F95C" ID-DOI="http://doi.org/10.5281/zenodo.6057680" ID-GBIF-Taxon="125107261" ID-Zenodo-Dep="6057680" LSID="urn:lsid:plazi:treatment:5B4FBD00FFE86153FF0263E4FEE3F95C" httpUri="http://treatment.plazi.org/id/5B4FBD00FFE86153FF0263E4FEE3F95C" lastPageNumber="58" pageId="57" pageNumber="58">
<subSubSection id="9BFC5F9DFFE86153FF0263E4FDBEFE27" box="[151,564,440,466]" pageId="57" pageNumber="58" type="nomenclature">
<paragraph id="D3590C16FFE86153FF0263E4FDBEFE27" blockId="57.[151,564,440,466]" box="[151,564,440,466]" pageId="57" pageNumber="58">
<heading id="8811BB7AFFE86153FF0263E4FDBEFE27" bold="true" box="[151,564,440,466]" fontSize="11" level="1" pageId="57" pageNumber="58" reason="1">
<emphasis id="E192D004FFE86153FF0263E4FDBEFE27" bold="true" box="[151,564,440,466]" pageId="57" pageNumber="58">
Key to
<taxonomicName id="14E67795FFE86153FF6563E4FEFEFE27" box="[240,372,440,466]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="genus">
<emphasis id="E192D004FFE86153FF6563E4FEFEFE27" bold="true" box="[240,372,440,466]" italics="true" pageId="57" pageNumber="58">Triodontus</emphasis>
</taxonomicName>
species (males)
</emphasis>
</heading>
</paragraph>
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<subSubSection id="9BFC5F9DFFE86153FF0263A2FEE3F95C" pageId="57" pageNumber="58" type="key">
<keyStep id="681214B3FFE86153FF0263A2FA11FD9E" pageId="57" pageNumber="58">
<paragraph id="D3590C16FFE86153FF0263A2FA16FDC5" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
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1 Central pronotal bulge with 3 more-or-less similar tubercles (
<figureCitation id="4BDD1093FFE86153FCD063A2FC19FDE6" box="[837,915,510,531]" captionStart="FIGURE 3" captionStartId="9.[151,250,1798,1820]" captionTargetBox="[156,1428,193,1776]" captionTargetId="figure@9.[156,1431,193,1777]" captionTargetPageId="9" captionText="FIGURE 3. Triodontus itremoi. Protibia in ventral view (A), metacoxa in dorsal view (B), metatibia in apical view (C), mesotibia in apical view (D), procoxa in ventral view (E), wing (F); pars. str — stridulatory field, procx. fv — hollow of procoxa; 1 a and 2 a—anal cells; AA 1 + 2, AAP, AA 3 b + (AA 4 + AP 1 + 2), AP 4 — anal veins; aas—antero-apical sclerotization; cas—costo-apical sclerotization; CuA—anterior cubital vein; js—jugal sclerotization; MP 3 + 4 — posterior medial vein; rc—radial cell; RMPfused posterior radial (RP 3 + 4), anterior medial and posterior medial (MP 1 + 2) veins; SV 2 — secondary “ vein ”. Wing venation terminology follows Fedorenko (2009). Not to scale." httpUri="https://zenodo.org/record/200447/files/figure.png" pageId="57" pageNumber="58">Figs. 3 1</figureCitation>
A–B); outer lobes of parameres rather large, mostly covering inner lobes (in lateral view:
<figureCitation id="4BDD1093FFE86153FDDD6047FD01FDC5" box="[584,651,539,560]" captionStart="FIGURE 31" captionStartId="61.[151,250,1844,1866]" captionTargetBox="[153,1433,201,1819]" captionTargetId="figure@61.[152,1435,193,1822]" captionTargetPageId="61" captionText="FIGURE 31. Triodontus nitidulus. Habitus in dorsal view (A—male, C—female), head and pronotum of male in dorsolateral view (B), internal sac of aedeagus (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200474/files/figure.png" pageId="57" pageNumber="58">Fig. 31</figureCitation>
E).............................................................. 2
</keyLead>
</paragraph>
<paragraph id="D3590C16FFE86153FF026064FA11FD9E" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF026064FA11FD9E" pageId="57" pageNumber="58">
<emphasis id="E192D004FFE86153FF026064FF15FDBB" bold="true" box="[151,159,568,590]" pageId="57" pageNumber="58">-</emphasis>
Central pronotal bulge with 2 tubercles (
<figureCitation id="4BDD1093FFE86153FDF36065FD20FDBB" box="[614,682,569,590]" captionStart="FIGURE 35" captionStartId="70.[151,250,1908,1930]" captionTargetBox="[176,1391,269,1883]" captionTargetId="figure@70.[176,1411,193,1886]" captionTargetPageId="70" captionText="FIGURE 35. Triodontus lemoulti¸ holotype male (A, B, D, E), paratype male (F, G), paratype female (C). Habitus in dorsal (A, C) and dorsolateral view (B), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200478/files/figure.png" pageId="57" pageNumber="58">Fig. 35</figureCitation>
A); outer lobes of parameres smaller, not covering inner lobes (in lateral view:
<figureCitation id="4BDD1093FFE86153FF74600AFEAFFD9E" box="[225,293,598,619]" captionStart="FIGURE 35" captionStartId="70.[151,250,1908,1930]" captionTargetBox="[176,1391,269,1883]" captionTargetId="figure@70.[176,1411,193,1886]" captionTargetPageId="70" captionText="FIGURE 35. Triodontus lemoulti¸ holotype male (A, B, D, E), paratype male (F, G), paratype female (C). Habitus in dorsal (A, C) and dorsolateral view (B), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200478/files/figure.png" pageId="57" pageNumber="58">Fig. 35</figureCitation>
E).......................................................................................... 4
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<keyStep id="681214B3FFE86153FF026028FA11FD53" pageId="57" pageNumber="58">
<paragraph id="D3590C16FFE86153FF026028FA16FD7C" blockId="57.[151,1436,510,1705]" box="[151,1436,628,649]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF026028FA16FD7C" box="[151,1436,628,649]" pageId="57" pageNumber="58">2 Body length less than 9.0 mm........................................................................... 3</keyLead>
</paragraph>
<paragraph id="D3590C16FFE86153FF0260CDFA11FD53" blockId="57.[151,1436,510,1705]" box="[151,1435,657,678]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF0260CDFA11FD53" box="[151,1435,657,678]" pageId="57" pageNumber="58">
- Body length 9.5–12.0 mm................................................................
<taxonomicName id="14E67795FFE86153FB7760CDFA11FD53" authority="Fairmaire" authorityName="Fairmaire" box="[1250,1435,657,678]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hova">
<emphasis id="E192D004FFE86153FB7760CDFAA3FD53" box="[1250,1321,657,678]" italics="true" pageId="57" pageNumber="58">T. hova</emphasis>
(Fairmaire)
</taxonomicName>
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</paragraph>
</keyStep>
<keyStep id="681214B3FFE86153FF0260F2FA11FCEE" pageId="57" pageNumber="58">
<paragraph id="D3590C16FFE86153FF0260F2FA11FD14" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF0260F2FA11FD14" pageId="57" pageNumber="58">
3 Abdominal sternite 8 narrower and more sinuate medially, without oval transverse concavity (
<figureCitation id="4BDD1093FFE86153FB0360F2FB6AFD36" box="[1174,1248,686,707]" captionStart="FIGURE 31" captionStartId="61.[151,250,1844,1866]" captionTargetBox="[153,1433,201,1819]" captionTargetId="figure@61.[152,1435,193,1822]" captionTargetPageId="61" captionText="FIGURE 31. Triodontus nitidulus. Habitus in dorsal view (A—male, C—female), head and pronotum of male in dorsolateral view (B), internal sac of aedeagus (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200474/files/figure.png" pageId="57" pageNumber="58">Fig. 31</figureCitation>
F); lateral lobes of parameres with small, keel-shaped processes on proximal parts (
<figureCitation id="4BDD1093FFE86153FCDF6090FC04FD14" box="[842,910,716,737]" captionStart="FIGURE 31" captionStartId="61.[151,250,1844,1866]" captionTargetBox="[153,1433,201,1819]" captionTargetId="figure@61.[152,1435,193,1822]" captionTargetPageId="61" captionText="FIGURE 31. Triodontus nitidulus. Habitus in dorsal view (A—male, C—female), head and pronotum of male in dorsolateral view (B), internal sac of aedeagus (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200474/files/figure.png" pageId="57" pageNumber="58">Fig. 31</figureCitation>
E).................
<taxonomicName id="14E67795FFE86153FBFB6090FA11FD14" authority="Guerin-Meneville" authorityName="Guerin-Meneville" box="[1134,1435,716,737]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="nitidulus">
<emphasis id="E192D004FFE86153FBFB6090FBF0FD14" box="[1134,1146,716,737]" italics="true" pageId="57" pageNumber="58">T</emphasis>
.
<emphasis id="E192D004FFE86153FB136090FB50FD14" box="[1158,1242,716,737]" italics="true" pageId="57" pageNumber="58">nitidulus</emphasis>
(Guérin-Méneville)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3590C16FFE86153FF0260B5FA11FCEE" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF0260B5FA11FCEE" pageId="57" pageNumber="58">
- Abdominal sternite 8 wider and less sinuate medially, with oval transverse concavity (
<figureCitation id="4BDD1093FFE86153FBA560B5FBF2FD0B" box="[1072,1144,745,766]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="57" pageNumber="58">Fig. 33</figureCitation>
G); lateral lobes of parameres without small, keel-shaped processes on proximal parts (
<figureCitation id="4BDD1093FFE86153FC95615AFCCFFCEE" box="[768,837,774,795]" captionStart="FIGURE 33" captionStartId="66.[151,250,1908,1930]" captionTargetBox="[176,1399,268,1883]" captionTargetId="figure@66.[176,1411,193,1886]" captionTargetPageId="66" captionText="FIGURE 33. Triodontus bicavatus. Habitus in dorsal view (A, B—male [A—holotype], C—female), labels of holotype (D), internal sac of aedeagus (E), aedeagus in lateral view and parameres in dorsal and ventral view (F), abdomen of holotype (G), distributional records (H)." httpUri="https://zenodo.org/record/200476/files/figure.png" pageId="57" pageNumber="58">Fig. 33</figureCitation>
F).............................
<taxonomicName id="14E67795FFE86153FB24615AFA11FCEE" authority="Fairmaire" authorityName="Fairmaire" box="[1201,1435,774,795]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="bicavatus">
<emphasis id="E192D004FFE86153FB24615AFAA2FCEE" box="[1201,1320,774,795]" italics="true" pageId="57" pageNumber="58">T. bicavatus</emphasis>
(Fairmaire)
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="681214B3FFE86153FF026178FA16FCA3" pageId="57" pageNumber="58">
<paragraph id="D3590C16FFE86153FF026178FA16FCCC" blockId="57.[151,1436,510,1705]" box="[151,1436,804,825]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF026178FA16FCCC" box="[151,1436,804,825]" pageId="57" pageNumber="58">
4 Abdominal sternite 8 with more-or-less developed tubercule medially (
<figureCitation id="4BDD1093FFE86153FC1C6178FC45FCCC" box="[905,975,804,825]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="57" pageNumber="58">Fig. 34</figureCitation>
F)................................... 5
</keyLead>
</paragraph>
<paragraph id="D3590C16FFE86153FF02611DFA16FCA3" blockId="57.[151,1436,510,1705]" box="[151,1436,833,854]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF02611DFA16FCA3" box="[151,1436,833,854]" pageId="57" pageNumber="58">
- Abdominal sternite 8 without tubercule medially (
<figureCitation id="4BDD1093FFE86153FD54611DFC8CFCA3" box="[705,774,833,854]" captionStart="FIGURE 36" captionStartId="72.[151,250,1910,1932]" captionTargetBox="[184,1401,253,1881]" captionTargetId="figure@72.[183,1404,195,1889]" captionTargetPageId="72" captionText="FIGURE 36. Triodontus modestus, lectotype male (A, B, D), paralectotype male (E, F), paralectotype female (C). Habitus in dorsal (A, C) and dorsolateral view (B), labels of lectotype (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200479/files/figure.png" pageId="57" pageNumber="58">Fig. 36</figureCitation>
G)............................................. ………. 6
</keyLead>
</paragraph>
</keyStep>
<keyStep id="681214B3FFE86153FF026102FA16FC3E" pageId="57" pageNumber="58">
<paragraph id="D3590C16FFE86153FF026102FA16FC64" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF026102FA16FC64" pageId="57" pageNumber="58">
5 Tubercules of abdominal sternite 8 rounded apically (
<figureCitation id="4BDD1093FFE86153FD4A6102FCAFFC86" box="[735,805,862,883]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="57" pageNumber="58">Fig. 34</figureCitation>
F); lateral lobes of parameres with keel-shaped processes on prox- imal parts (in dorsal view,
<figureCitation id="4BDD1093FFE86153FE756120FDAFFC64" box="[480,549,892,913]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="57" pageNumber="58">Fig. 34</figureCitation>
D)..............................
<taxonomicName id="14E67795FFE86153FC3E6120FA16FC64" authority="Frolov, Montreuil &amp; Akhmetova" authorityName="Frolov, Montreuil &amp; Akhmetova" box="[939,1436,892,913]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="ankarafantsikae">
<emphasis id="E192D004FFE86153FC3E6120FC3DFC64" box="[939,951,892,913]" italics="true" pageId="57" pageNumber="58">T</emphasis>
.
<emphasis id="E192D004FFE86153FC566120FBD4FC64" box="[963,1118,892,913]" italics="true" pageId="57" pageNumber="58">ankarafantsikae</emphasis>
Frolov, Montreuil &amp; Akhmetova
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3590C16FFE86153FF0261C5FA16FC3E" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
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- Tubercule of abdominal sternite 8 with concavity apically (
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G); lateral lobes of parameres with semicircular notches on proximal parts (in dorsal view,
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E).................................
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<emphasis id="E192D004FFE86153FC6461EAFBD6FC3E" box="[1009,1116,950,971]" italics="true" pageId="57" pageNumber="58">T. lemoulti</emphasis>
Frolov, Montreuil &amp; Akhmetova
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6 Body length less than
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.........................................................
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.
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(Benderitter)
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<keyLead id="6817AF23FFE86153FF0261ADFA16FBF3" box="[151,1436,1009,1030]" pageId="57" pageNumber="58">- Body length 9.0–12.0 mm.............................................................................. 7</keyLead>
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7 Outer lobes of parameres strongly sclerotized, with distinct lateral notches and teeth situated more-or-less apically (
<figureCitation id="4BDD1093FFE86153FAA16652FAF3FBD6" box="[1332,1401,1038,1059]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStart-3="FIGURE 6" captionStart-4="FIGURE 7" captionStartId-0="9.[151,250,1798,1820]" captionStartId-1="10.[151,250,1673,1695]" captionStartId-2="13.[151,250,1941,1963]" captionStartId-3="15.[151,250,1814,1836]" captionStartId-4="17.[151,250,1807,1829]" captionTargetBox-0="[156,1428,193,1776]" captionTargetBox-1="[163,1418,185,1644]" captionTargetBox-2="[243,1371,374,1916]" captionTargetBox-3="[216,1403,212,1792]" captionTargetBox-4="[220,1364,197,1782]" captionTargetId-0="figure@9.[156,1431,193,1777]" captionTargetId-1="figure@10.[161,1425,185,1652]" captionTargetId-2="figure@13.[216,1371,373,1920]" captionTargetId-3="figure@15.[181,1405,193,1793]" captionTargetId-4="figure@17.[206,1380,193,1796]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="13" captionTargetPageId-3="15" captionTargetPageId-4="17" captionText-0="FIGURE 3. Triodontus itremoi. Protibia in ventral view (A), metacoxa in dorsal view (B), metatibia in apical view (C), mesotibia in apical view (D), procoxa in ventral view (E), wing (F); pars. str — stridulatory field, procx. fv — hollow of procoxa; 1 a and 2 a—anal cells; AA 1 + 2, AAP, AA 3 b + (AA 4 + AP 1 + 2), AP 4 — anal veins; aas—antero-apical sclerotization; cas—costo-apical sclerotization; CuA—anterior cubital vein; js—jugal sclerotization; MP 3 + 4 — posterior medial vein; rc—radial cell; RMPfused posterior radial (RP 3 + 4), anterior medial and posterior medial (MP 1 + 2) veins; SV 2 — secondary “ vein ”. Wing venation terminology follows Fedorenko (2009). Not to scale." captionText-1="FIGURE 4. Triodontus itremoi, abdomen (A—dorsal, B—ventral view): cav. mtcox — metacoxal cavity, plec — plectrum, propyg — propygidium, pyg — pygidium, stern 2 – stern 8 — abdominal sternites 2 – 8, terg 1 – terg 6 — abdominal tergites 1 – 6; Triodontus lemoulti, spiculum gastrale (C); Triodontus itremoi, female genitalia (D): burs. cop — bursa copulatrix, coxt — coxite, gl. acc — vaginal gland, gl. rec — spermatheca gland, ovd — oviduct, parapr — paraproct, rec. sem — spermatheca, scoxt — subcoxite, styl — stylus. Not to scale." captionText-2="FIGURE 5. Pseudorphnus carinatus. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), labels of holotype (D), head in dorsal view (E), head and pronotum in dorso-lateral view (F), distributional records (G)." captionText-3="FIGURE 6. Pseudorphnus olsoufieffi. Habitus (A—dorsal, B—lateral view), aedeagus in lateral view and parameres in dorsal view (C), head in dorsal view (D), head and pronotum in dorsolateral view (E), labels of holotype (F), distributional records (G)." captionText-4="FIGURE 7. Pseudorphnus hiboni. Habitus in dorsal view (A—male, F—female), head and pronotum in dorsolateral view (B), aedeagus in lateral view and parameres in dorsal view (C), protibia (D, male left, female right), base of elytra, SEM (E), labels of holotype (G), distributional records (H)." httpUri-0="https://zenodo.org/record/200447/files/figure.png" httpUri-1="https://zenodo.org/record/200448/files/figure.png" httpUri-2="https://zenodo.org/record/200449/files/figure.png" httpUri-3="https://zenodo.org/record/200450/files/figure.png" httpUri-4="https://zenodo.org/record/200451/files/figure.png" pageId="57" pageNumber="58">Fig. 3 7</figureCitation>
D)................................................................................................... 8
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- Outer lobes of parameres less sclerotized, somewhat rolled up in most species; lateral notches, if present, situated more basally (
<figureCitation id="4BDD1093FFE86153FF7D663AFEA7FB8E" box="[232,301,1126,1147]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="57" pageNumber="58">Fig. 41</figureCitation>
E)......................................................................................... 10
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<paragraph id="D3590C16FFE86153FF0266D8FA10FB6C" blockId="57.[151,1436,510,1705]" box="[151,1434,1156,1177]" pageId="57" pageNumber="58">
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8 Outer lobes of parameres slender in lateral and dorsal view (
<figureCitation id="4BDD1093FFE86153FCB666D8FCE3FB6C" box="[803,873,1156,1177]" captionStart="FIGURE 37" captionStartId="74.[151,250,1864,1886]" captionTargetBox="[176,1383,233,1839]" captionTargetId="figure@74.[176,1410,193,1843]" captionTargetPageId="74" captionText="FIGURE 37. Triodontus owas, holotype male (A, B, D), male (E, F), female (C). Habitus in dorsal view (A, C), labels of holotype (B), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200480/files/figure.png" pageId="57" pageNumber="58">Fig. 37</figureCitation>
D).............................
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.
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Westwood
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<keyLead id="6817AF23FFE86153FF0266FDFA16FB43" box="[151,1436,1185,1206]" pageId="57" pageNumber="58">
- Outer lobes of parameres wider, at lease in dorsal view (
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E, 40E)......................................... 9
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9 Outer lobes of parameres with slit-shaped lateral notches (
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E)............
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.
<emphasis id="E192D004FFE86153FBB766E2FBD7FB26" box="[1058,1117,1214,1235]" italics="true" pageId="57" pageNumber="58">viettei</emphasis>
Frolov, Montreuil &amp; Akhmetova
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<keyLead id="6817AF23FFE86153FF026680FA11FB04" box="[151,1435,1244,1265]" pageId="57" pageNumber="58">
- Outer lobes of parameres with wide lateral notches (
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E)............................
<taxonomicName id="14E67795FFE86153FB016680FA11FB04" authority="Fairmaire" authorityName="Fairmaire" box="[1172,1435,1244,1265]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FFE86153FB016680FAA2FB04" box="[1172,1320,1244,1265]" italics="true" pageId="57" pageNumber="58">T. hildebrandtii</emphasis>
(Fairmaire)
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10 Outer lobes of parameres long, curved downward and inward, with more-or-less separated strongly sclerotized lateral parts and membranous, overlapping medial parts; lateral notches absent (
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E)...........................
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.
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Paulian
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<keyLead id="6817AF23FFE86153FF026768FA10FA93" pageId="57" pageNumber="58">- Outer lobes of parameres of different shape, without distinctly separated lateral and medial, membranous parts; lateral notches absent or present..................................................................................... 11</keyLead>
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11 Outer lobes of parameres without lateral notches (
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F)................
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<emphasis id="E192D004FFE86153FC706732FBD6FA76" box="[997,1116,1390,1411]" italics="true" pageId="57" pageNumber="58">T. fairmairei</emphasis>
Frolov, Montreuil &amp; Akhmetova
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<keyLead id="6817AF23FFE86153FF0267D0FA16FA54" box="[151,1436,1420,1441]" pageId="57" pageNumber="58">
- Outer lobes of parameres with more-or-less distinct lateral notches and teeth (
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F, 44D, 45E, 46D).............. 12
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12 Outer lobes of parameres with distinct, slit-shaped lateral notches (in lateral view,
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F, 44D)....................13
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- Outer lobes of parameres less distinct, wider lateral notches (in lateral view,
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E, 46D)......................... 14
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<paragraph id="D3590C16FFE86153FF0267B8FA10F9E3" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF0267B8FA10F9E3" pageId="57" pageNumber="58">
13 Abdominal sternite 8 without fossa (
<figureCitation id="4BDD1093FFE86153FDAA67B8FD0FFA0C" box="[575,645,1508,1529]" captionStart="FIGURE 43" captionStartId="83.[151,250,1887,1909]" captionTargetBox="[170,1387,200,1836]" captionTargetId="figure@83.[169,1418,193,1865]" captionTargetPageId="83" captionText="FIGURE 43. Triodontus itremoi, holotype male (A, B, C), paratype male (E right, F, G, H), paratype female (E left, D). Habitus in dorsal (A, D) and dorsolateral view (B), labels (C), abdomen (E, H), aedeagus in lateral view and parameres in dorsal and ventral view (F), internal sac of aedeagus (G), distributional records (I)." httpUri="https://zenodo.org/record/200486/files/figure.png" pageId="57" pageNumber="58">Fig. 43</figureCitation>
E); slit-shaped lateral notches of outer lobes of parameres longer (
<figureCitation id="4BDD1093FFE86153FB6967B8FACEFA0C" box="[1276,1348,1508,1529]" captionStart="FIGURE 43" captionStartId="83.[151,250,1887,1909]" captionTargetBox="[170,1387,200,1836]" captionTargetId="figure@83.[169,1418,193,1865]" captionTargetPageId="83" captionText="FIGURE 43. Triodontus itremoi, holotype male (A, B, C), paratype male (E right, F, G, H), paratype female (E left, D). Habitus in dorsal (A, D) and dorsolateral view (B), labels (C), abdomen (E, H), aedeagus in lateral view and parameres in dorsal and ventral view (F), internal sac of aedeagus (G), distributional records (I)." httpUri="https://zenodo.org/record/200486/files/figure.png" pageId="57" pageNumber="58">Fig. 43</figureCitation>
F)..................................................................................................
<taxonomicName id="14E67795FFE86153FAD5645DFA10F9E3" box="[1344,1434,1537,1558]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="itremoi">
<emphasis id="E192D004FFE86153FAD5645DFA10F9E3" box="[1344,1434,1537,1558]" italics="true" pageId="57" pageNumber="58">T. itremoi</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="D3590C16FFE86153FF026442FA16F9A4" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF026442FA16F9A4" pageId="57" pageNumber="58">
- Abdominal sternite 8 with shallow but distinct, somewhat transversal fossa (
<figureCitation id="4BDD1093FFE86153FC4D6442FBA8F9C6" box="[984,1058,1566,1587]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="57" pageNumber="58">Fig. 44</figureCitation>
E); slit-shaped lateral notches of outer lobes of parameres shorter (
<figureCitation id="4BDD1093FFE86153FE7E6460FDBBF9A4" box="[491,561,1596,1617]" captionStart="FIGURE 44" captionStartId="86.[151,250,1871,1893]" captionTargetBox="[185,1396,298,1847]" captionTargetId="figure@86.[184,1403,229,1850]" captionTargetPageId="86" captionText="FIGURE 44. Triodontus copridoides, holotype male (A, B, D, E), male (C). Habitus in dorsal view (A, C), labels (D), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200487/files/figure.png" pageId="57" pageNumber="58">Fig. 44</figureCitation>
D).....................................................
<taxonomicName id="14E67795FFE86153FB506460FA16F9A4" authority="Paulian" authorityName="Paulian" box="[1221,1436,1596,1617]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="copridoides">
<emphasis id="E192D004FFE86153FB506460FAC7F9A4" box="[1221,1357,1596,1617]" italics="true" pageId="57" pageNumber="58">T. copridoides</emphasis>
Paulian
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="681214B3FFE86153FF026405FEE3F95C" pageId="57" pageNumber="58">
<paragraph id="D3590C16FFE86153FF026405FA11F99B" blockId="57.[151,1436,510,1705]" box="[151,1435,1625,1646]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF026405FA11F99B" box="[151,1435,1625,1646]" pageId="57" pageNumber="58">
14 Outer lobes of parameres 1.3 times longer than inner lobes; inner lobes smaller (in lateral view) (
<figureCitation id="4BDD1093FFE86153FB146405FB4CF99B" box="[1153,1222,1625,1646]" captionStart="FIGURE 45" captionStartId="88.[151,250,1946,1968]" captionTargetBox="[219,1367,312,1926]" captionTargetId="figure@88.[219,1368,244,1929]" captionTargetPageId="88" captionText="FIGURE 45. Triodontus hanskii, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), internal sac of aedeagus (D), aedeagus in lateral view and parameres in dorsal and ventral view (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200488/files/figure.png" pageId="57" pageNumber="58">Fig. 45</figureCitation>
E)..
<taxonomicName id="14E67795FFE86153FB6C6405FA11F99B" authority="Frolov" authorityName="Frolov" box="[1273,1435,1625,1646]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="hanskii">
<emphasis id="E192D004FFE86153FB6C6405FADCF99B" box="[1273,1366,1625,1646]" italics="true" pageId="57" pageNumber="58">T. hanskii</emphasis>
Frolov
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="D3590C16FFE86153FF02642AFEE3F95C" blockId="57.[151,1436,510,1705]" pageId="57" pageNumber="58">
<keyLead id="6817AF23FFE86153FF02642AFEE3F95C" pageId="57" pageNumber="58">
- Outer lobes of parameres as long as inner lobes; inner lobes larger (in lateral view,
<figureCitation id="4BDD1093FFE86153FC69642AFBCBF97E" box="[1020,1089,1654,1675]" captionStart="FIGURE 46" captionStartId="89.[151,250,1858,1880]" captionTargetBox="[220,1389,271,1824]" captionTargetId="figure@89.[195,1392,193,1825]" captionTargetPageId="89" captionText="FIGURE 46. Triodontus inexpectatus, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), abdomen (E), distributional records (F)." httpUri="https://zenodo.org/record/200489/files/figure.png" pageId="57" pageNumber="58">Fig. 46</figureCitation>
D)
<taxonomicName id="14E67795FFE86153FBC9642AFEE3F95C" authority="Frolov, Montreuil &amp; Akhmetova" authorityName="Frolov, Montreuil &amp; Akhmetova" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="57" pageNumber="58" phylum="Arthropoda" rank="species" species="inexpectatus">
<emphasis id="E192D004FFE86153FBC9642AFB60F97E" box="[1116,1258,1654,1675]" italics="true" pageId="57" pageNumber="58">T. inexpectatus</emphasis>
Frolov, Montreuil &amp; Akhmetova
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</paragraph>
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</subSubSection>
</treatment>
</document>
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