158 lines
18 KiB
XML
158 lines
18 KiB
XML
<document id="6CB48BCB106D52BB1E4B9E5328869A04" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFE0615BFF0263CAFDBBFAA4" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Madecorphnus brunneus Frolov 2010" docType="treatment" docVersion="16" lastPageNumber="50" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="50" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:mods id="A492AD796980B6AA461FB5962D7073C4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="9A72A0CB0F52CF58F234516F66C49F9D">
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<mods:title id="54472B711995901153FC2C1B6E3DBD32">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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</mods:titleInfo>
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<mods:name id="C183B89EE87474C55E68C3C7D800F463" type="personal">
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<mods:role id="080101F29070D942EA148A7850A79845">
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<mods:roleTerm id="40856718A8F51D7977EB4747C8D64BB0">Author</mods:roleTerm>
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<mods:namePart id="5EB3662A07E5C108DB6AF19B5ABE18EE">Frolov, Andrey V.</mods:namePart>
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<mods:name id="CCC51E1A3AAA094A65BEFA48003BABB5" type="personal">
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<mods:roleTerm id="6244B90937AF86729DF635DA85B571DB">Author</mods:roleTerm>
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<mods:namePart id="02C6E6199B31E20ABC90D46F982AE543">Montreuil, Olivier</mods:namePart>
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</mods:name>
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<mods:name id="89460970F91420E6CC935C79B1016EB4" type="personal">
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<mods:roleTerm id="5A6519DDF1B294FE5D150E42A4112684">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="CE893E1DABA2F2257EE2B203149A36CC">Akhmetova, Lilia A.</mods:namePart>
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<mods:typeOfResource id="EA251BEFA6200274DFE20795907D835F">text</mods:typeOfResource>
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<mods:titleInfo id="050C3698D2D51CBBC4622F29694985D4">
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<mods:title id="72165BE02783BF9C769AFC850C55369E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="6543A991916255A020EA989CE21BC136">
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<mods:date id="89DF485E4993C6FAD0BF173EFC97874D">2016</mods:date>
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<mods:detail id="CAB7FB098DDA3F498165806FDCCEB071" type="volume">
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<mods:number id="76C674272ABFB6427D0AE1CED7793C43">4207</mods:number>
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<mods:detail id="9EBB24819D57A284F5C1867DA66C3076" type="issue">
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<mods:number id="CBBFBEB78F5C0F3B4C1A90C8138DCAE8">1</mods:number>
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<mods:end id="B54D15A70E2BA6440AE4D93F5406CE50">93</mods:end>
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<mods:classification id="CBD9E3A71642E128B946377EC75C3C62">journal article</mods:classification>
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<mods:identifier id="6F987D9895A31300923BF2C1BA38B005" type="CLB-Dataset">37549</mods:identifier>
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<mods:identifier id="59724F86872AAB527DD772ED745A5E51" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
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<mods:identifier id="AFD52234E174FCA5E5CDBE2AA96AF626" type="GBIF-Dataset">cef35771-ff95-4a41-9af6-196dea899a64</mods:identifier>
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<mods:identifier id="65B801C4472E84151F5007F61E6520DD" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5B4FBD00FFE0615BFF0263CAFDBBFAA4" ID-DOI="http://doi.org/10.5281/zenodo.6057668" ID-GBIF-Taxon="125107282" ID-Zenodo-Dep="6057668" LSID="urn:lsid:plazi:treatment:5B4FBD00FFE0615BFF0263CAFDBBFAA4" httpUri="http://treatment.plazi.org/id/5B4FBD00FFE0615BFF0263CAFDBBFAA4" lastPageNumber="50" pageId="49" pageNumber="50">
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<subSubSection id="9BFC5F9DFFE0615BFF0263CAFEBCFE27" pageId="49" pageNumber="50" type="nomenclature">
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<paragraph id="D3590C16FFE0615BFF0263CAFDE0FE45" blockId="49.[151,618,406,466]" box="[151,618,406,432]" pageId="49" pageNumber="50">
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<heading id="8811BB7AFFE0615BFF0263CAFDE0FE45" bold="true" box="[151,618,406,432]" fontSize="11" level="1" pageId="49" pageNumber="50" reason="1">
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<taxonomicName id="14E67795FFE0615BFF0263CAFDE0FE45" authority="Frolov, 2010" authorityName="Frolov" authorityYear="2010" box="[151,618,406,432]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
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||
<emphasis id="E192D004FFE0615BFF0263CAFDE0FE45" bold="true" box="[151,618,406,432]" pageId="49" pageNumber="50">
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<emphasis id="E192D004FFE0615BFF0263CAFE43FE45" bold="true" box="[151,457,406,432]" italics="true" pageId="49" pageNumber="50">Madecorphnus brunneus</emphasis>
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Frolov, 2010
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="D3590C16FFE0615BFF0263E5FEBCFE27" blockId="49.[151,618,406,466]" box="[151,310,441,466]" pageId="49" pageNumber="50">
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(
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||
<figureCitation id="4BDD1093FFE0615BFF0A63E5FF73FE27" box="[159,249,441,466]" captionStart="FIGURE 25" captionStartId="50.[151,250,1700,1722]" captionTargetBox="[196,1383,217,1666]" captionTargetId="figure@50.[196,1390,193,1678]" captionTargetPageId="50" captionText="FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200468/files/figure.png" pageId="49" pageNumber="50">Figs. 25</figureCitation>
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A–F)
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFE0615BFF02605DFDE6FDE2" box="[151,620,513,535]" pageId="49" pageNumber="50" type="reference_group">
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||
<paragraph id="D3590C16FFE0615BFF02605DFDE6FDE2" blockId="49.[151,620,513,535]" box="[151,620,513,535]" pageId="49" pageNumber="50">
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<treatmentCitationGroup id="F3F62B38FFE0615BFF02605DFDE6FDE2" box="[151,620,513,535]" pageId="49" pageNumber="50">
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<taxonomicName id="14E67795FFE0615BFF02605DFDE2FDE2" authority="Frolov, 2010c: 1108" authorityName="Frolov" authorityPageNumber="1108" authorityYear="2010" box="[151,616,513,535]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="E192D004FFE0615BFF02605DFE19FDE2" box="[151,403,513,535]" italics="true" pageId="49" pageNumber="50">Madecorphnus brunneus</emphasis>
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<treatmentCitation id="52472A07FFE0615BFE0F605DFDE2FDE2" author="Frolov" box="[410,616,513,535]" page="1108" pageId="49" pageNumber="50" year="2010">
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<bibRefCitation id="B77771E7FFE0615BFE0F605DFDA0FDE2" author="Frolov" box="[410,554,513,535]" pageId="49" pageNumber="50" refString="Frolov, A. V. (2010 c) Revision of the Madagascan genus Madecorphnus Paulian (Coleoptera, Scarabaeidae, Orphninae). Journal of Natural History, 44, 1095 - 1111." type="journal article" year="2010" yearSuffix="c">Frolov, 2010c</bibRefCitation>
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: 1108
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFE0615BFF026018FC78FD75" pageId="49" pageNumber="50" type="materials_examined">
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<paragraph id="D3590C16FFE0615BFF026018FC78FD75" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">
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<emphasis id="E192D004FFE0615BFF026018FE48FDA8" bold="true" box="[151,450,580,605]" pageId="49" pageNumber="50">Type material examined.</emphasis>
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Holotype (
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<figureCitation id="4BDD1093FFE0615BFDD16018FD2BFDA8" box="[580,673,580,605]" captionStart="FIGURE 25" captionStartId="50.[151,250,1700,1722]" captionTargetBox="[196,1383,217,1666]" captionTargetId="figure@50.[196,1390,193,1678]" captionTargetPageId="50" captionText="FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200468/files/figure.png" pageId="49" pageNumber="50">Figs. 25</figureCitation>
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A–E), male, “ANDRINGITRA Est Anjavidilava
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<quantity id="141EA1F3FFE0615BFB476018FAE4FDA9" box="[1234,1390,580,605]" metricMagnitude="3" metricUnit="m" metricValue="1.9" metricValueMax="1.95" metricValueMin="1.85" pageId="49" pageNumber="50" unit="m" value="1900.0" valueMax="1950.0" valueMin="1850.0">1850–1950 m</quantity>
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<date id="A7582AD6FFE0615BFAE26018FF4BFD74" bridgedPair="-" pageId="49" pageNumber="50" value="1918-12">18- XII</date>
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/
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<date id="A7582AD6FFE0615BFF5F6034FEBCFD74" box="[202,310,616,641]" pageId="49" pageNumber="50" value="1971-01-15">15-I-1971</date>
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/ Holotypus
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<taxonomicName id="14E67795FFE0615BFE526034FCFAFD75" authority="Frolov 2009" authorityName="Frolov" authorityYear="2009" box="[455,880,616,641]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="E192D004FFE0615BFE526034FD54FD74" box="[455,734,616,641]" italics="true" pageId="49" pageNumber="50">Madecorphnus brunneus</emphasis>
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<bibRefCitation id="B77771E7FFE0615BFD706034FCFAFD75" author="Frolov" box="[741,880,616,641]" pageId="49" pageNumber="50" refString="Frolov, A. V. & Montreuil, O. (2009) A new genus of Orphninae (Coleoptera, Scarabaeidae) from Madagascar. Zoosystematica Rossica, 18, 65 - 69." type="journal article" year="2009">Frolov 2009</bibRefCitation>
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</taxonomicName>
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” (MNHN).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFE0615BFF5260D0FBFCFD3D" pageId="49" pageNumber="50" type="diagnosis">
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||
<paragraph id="D3590C16FFE0615BFF5260D0FBFCFD3D" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">
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<emphasis id="E192D004FFE0615BFF5260D0FECBFD50" bold="true" box="[199,321,652,677]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
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<taxonomicName id="14E67795FFE0615BFEDF60D0FDE9FD50" box="[330,611,652,677]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="E192D004FFE0615BFEDF60D0FDE9FD50" box="[330,611,652,677]" italics="true" pageId="49" pageNumber="50">Madecorphnus brunneus</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="14E67795FFE0615BFD6860D1FC21FD51" box="[765,939,653,676]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="perinetensis">
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<emphasis id="E192D004FFE0615BFD6860D1FC21FD51" box="[765,939,653,676]" italics="true" pageId="49" pageNumber="50">M. perinetensis</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="14E67795FFE0615BFC7360D1FBF4FD51" box="[998,1150,653,676]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="peyrierasi">
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<emphasis id="E192D004FFE0615BFC7360D1FBF4FD51" box="[998,1150,653,676]" italics="true" pageId="49" pageNumber="50">M. peyrierasi</emphasis>
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</taxonomicName>
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but differs from them in having the parameres less acute in lateral view and in the different internal sac armature.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFE0615BFF526088FE28FAFC" pageId="49" pageNumber="50" type="description">
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<paragraph id="D3590C16FFE0615BFF526088FA95FD18" blockId="49.[151,1437,580,1361]" box="[199,1311,724,749]" pageId="49" pageNumber="50">
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<emphasis id="E192D004FFE0615BFF526088FE29FD18" bold="true" box="[199,419,724,749]" pageId="49" pageNumber="50">Description. Male.</emphasis>
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Body length
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<quantity id="141EA1F3FFE0615BFDAE6088FD1EFD19" box="[571,660,724,749]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="49" pageNumber="50" unit="mm" value="6.2">6.2 mm</quantity>
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. Color brown, pronotum and head dark brown (
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<figureCitation id="4BDD1093FFE0615BFB3C6088FB76FD18" box="[1193,1276,724,749]" captionStart="FIGURE 25" captionStartId="50.[151,250,1700,1722]" captionTargetBox="[196,1383,217,1666]" captionTargetId="figure@50.[196,1390,193,1678]" captionTargetPageId="50" captionText="FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200468/files/figure.png" pageId="49" pageNumber="50">Fig. 25</figureCitation>
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A).
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</paragraph>
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<paragraph id="D3590C16FFE0615BFF5260A4FD8CFCC0" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">Right mandible about 2 times longer than left, without tooth basad of apex. Labrum trapezoidal, its length about 1/5 width (in dorsal view).</paragraph>
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<paragraph id="D3590C16FFE0615BFF52611CFD45FC89" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">Frontoclypeus asymmetrical, slightly convex anteriorly, obtuse, with 2 long setae. Frontoclypeus slightly depressed in the middle anteriorly, finely punctate.</paragraph>
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<paragraph id="D3590C16FFE0615BFF5261D5FCA8FC31" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">Pronotum 1.9 times wider than long, widest medially. Margins with relatively wide border, lateral margins with 4 long setae. Pronotum evenly convex, minutely punctate.</paragraph>
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<paragraph id="D3590C16FFE0615BFF52618CFA9BFBC4" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">Elytra convex, with distinct humeral and apical tubercules. Maximum width approximately at basal 1/3. First stria distinct and reaching the apex of elytron, other striae indistinct. Disc of elytra with sparse punctation. Epipleura with long, sparse, brown setae. Base of elytra with border from scutellum to humeral callus.</paragraph>
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<paragraph id="D3590C16FFE0615BFF526660FB3EFB68" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">Protibiae with 3 outer teeth, lateral margin basad of outer teeth not crenulate. Apex with robust, spur-like seta and a few smaller setae basally. Mesothoracic and metathoracic legs similar in shape. Longer tibial spur shorter than 2 basal tarsomeres in mesothoracic legs and as long as 2 tarsomeres in metathoracic legs.</paragraph>
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<paragraph id="D3590C16FFE0615BFF5266F5FD6DFB10" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">
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Parameres acutely rounded in lateral view (
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<figureCitation id="4BDD1093FFE0615BFD3A66F4FC88FB34" box="[687,770,1192,1217]" captionStart="FIGURE 25" captionStartId="50.[151,250,1700,1722]" captionTargetBox="[196,1383,217,1666]" captionTargetId="figure@50.[196,1390,193,1678]" captionTargetPageId="50" captionText="FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200468/files/figure.png" pageId="49" pageNumber="50">Fig. 25</figureCitation>
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B). Internal sac of the aedeagus with 1 longer sclerite and 1 smaller, somewhat bifurcated sclerite (
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<figureCitation id="4BDD1093FFE0615BFDD26690FD2EFB10" box="[583,676,1228,1253]" captionStart="FIGURE 25" captionStartId="50.[151,250,1700,1722]" captionTargetBox="[196,1383,217,1666]" captionTargetId="figure@50.[196,1390,193,1678]" captionTargetPageId="50" captionText="FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200468/files/figure.png" pageId="49" pageNumber="50">Figs. 25</figureCitation>
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D–E).
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</paragraph>
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<paragraph id="D3590C16FFE0615BFF5266ACFE28FAFC" blockId="49.[151,1437,580,1361]" box="[199,418,1264,1289]" pageId="49" pageNumber="50">
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<emphasis id="E192D004FFE0615BFF5266ACFEAFFAFC" bold="true" box="[199,293,1264,1289]" pageId="49" pageNumber="50">Female.</emphasis>
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Unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFFE0615BFF526748FDBBFAA4" pageId="49" pageNumber="50" type="distribution">
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<paragraph id="D3590C16FFE0615BFF526748FDBBFAA4" blockId="49.[151,1437,580,1361]" pageId="49" pageNumber="50">
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||
<emphasis id="E192D004FFE0615BFF526748FE70FAD8" bold="true" box="[199,506,1300,1325]" pageId="49" pageNumber="50">Distribution and habitat.</emphasis>
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<taxonomicName id="14E67795FFE0615BFD936748FCABFAD8" box="[518,801,1300,1325]" class="Insecta" family="Orphnidae" genus="Madecorphnus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brunneus">
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<emphasis id="E192D004FFE0615BFD936748FCABFAD8" box="[518,801,1300,1325]" italics="true" pageId="49" pageNumber="50">Madecorphnus brunneus</emphasis>
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</taxonomicName>
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is known from one locality in the Adringitra Massif, southeastern
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<collectingCountry id="ABF14C86FFE0615BFEBF6764FE3BFAA4" box="[298,433,1336,1361]" name="Madagascar" pageId="49" pageNumber="50">Madagascar</collectingCountry>
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(
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<figureCitation id="4BDD1093FFE0615BFE556764FD99FAA4" box="[448,531,1336,1361]" captionStart="FIGURE 25" captionStartId="50.[151,250,1700,1722]" captionTargetBox="[196,1383,217,1666]" captionTargetId="figure@50.[196,1390,193,1678]" captionTargetPageId="50" captionText="FIGURE 25. Madecorphnus brunneus, holotype. Habitus in dorsal view (A), aedeagus in lateral view and parameres in dorsal view (B), labels (C), internal sac of aedeagus (D—photograph, E—schematic representation of armature), distributional records (F)." httpUri="https://zenodo.org/record/200468/files/figure.png" pageId="49" pageNumber="50">Fig. 25</figureCitation>
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F).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |