129 lines
14 KiB
XML
129 lines
14 KiB
XML
<document id="AF9DD8B7640EBEEBC4E38D1D8AE90D67" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FFDA6166FF02646AFCD0FD70" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Pseudorphnus Benderitter 1913" docType="treatment" docVersion="16" lastPageNumber="13" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="12" updateTime="1698433759527" updateUser="ExternalLinkService">
|
||
<mods:mods id="693C5584753E4504BE8D41E80016FEFA" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="3398A78916471BFE5F70586ACF407829">
|
||
<mods:title id="ED65456E69491EBB2295BB6BFD7421A3">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="A5A973FFEF4D604789B699F8C9331061" type="personal">
|
||
<mods:role id="3AA55E1CF5CC6C4C5CD1B78041B480E8">
|
||
<mods:roleTerm id="4E58C91F54C9438D178F38DE120143D7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="D52B91889E98EE422263CCBD6F09393B">Frolov, Andrey V.</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="A7D40AE940761F769D4FB1021A0DAD66" type="personal">
|
||
<mods:role id="ED88F567F98EEDBA72CCE22C730A4749">
|
||
<mods:roleTerm id="E806FA5C7C2216C8213DB4C6A8FA7D79">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="A71D88DC02D19CF429E53461301C346A">Montreuil, Olivier</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="6DA47A2B2254CE7E5C51E6B9C89A3D99" type="personal">
|
||
<mods:role id="1E171D6E261BF02C45A1FCF78A34EC92">
|
||
<mods:roleTerm id="D9482922D499D5B1E332902E9BCFDC19">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="15DF98228E3FABD2588A418B780B4AEA">Akhmetova, Lilia A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="DB8AA744586C1953D07F79D3E2472CCC">text</mods:typeOfResource>
|
||
<mods:relatedItem id="7921FFDA8BC5A50CE20D8F28F9AA29F0" type="host">
|
||
<mods:titleInfo id="01239A9D8BCE00C3C9514B322A204BD4">
|
||
<mods:title id="14470DD723B071968583A7221561042C">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="060DC8C3E27A102E750E162CF06E8F81">
|
||
<mods:date id="3383357E176303EC58FE626E3C1F7283">2016</mods:date>
|
||
<mods:detail id="78AFDA10DCBD2DCEC176F36631A8F64D" type="volume">
|
||
<mods:number id="0D555848F6B2F0CFC4DD5066B6331511">4207</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="B6039B8F28F7E6EE8BF8D4AB2AF63DE0" type="issue">
|
||
<mods:number id="229B9AE50D0D5B1DAD21D606F965B39C">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="87CBD49D60BC51D2894BF2404D62102F" unit="page">
|
||
<mods:start id="FD22109D3ADAC2F7DD8BD29D0986A2A2">1</mods:start>
|
||
<mods:end id="EDBE053892996969844C7830A3F4F3B1">93</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F5B931D70BE53548F1750EEA444EB9EF">journal article</mods:classification>
|
||
<mods:identifier id="DAC9A20E0981FFD64D1A58852DA9EBA0" type="CLB-Dataset">37549</mods:identifier>
|
||
<mods:identifier id="EAD052E19C039890307CFBBC39B169D0" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
|
||
<mods:identifier id="EF40A33802D6A4CDF181677DEC52F557" type="GBIF-Dataset">cef35771-ff95-4a41-9af6-196dea899a64</mods:identifier>
|
||
<mods:identifier id="10908FE31AD0140E18DF84F6718DBBD4" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="79277AA444211CC4179D26AEA95ACFBE" type="Zenodo-Dep">200444</mods:identifier>
|
||
<mods:identifier id="4380E30DFF92CE3C90AFFE123FBE4C27" type="ZooBank">F2875582-31E2-496F-AEEF-1D657DD86C33</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="5B4FBD00FFDA6166FF02646AFCD0FD70" ID-DOI="http://doi.org/10.5281/zenodo.6057630" ID-GBIF-Taxon="125107266" ID-Zenodo-Dep="6057630" LSID="urn:lsid:plazi:treatment:5B4FBD00FFDA6166FF02646AFCD0FD70" httpUri="http://treatment.plazi.org/id/5B4FBD00FFDA6166FF02646AFCD0FD70" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<subSubSection id="9BFC5F9DFFDA6161FF02646AFDF1F9A5" box="[151,635,1590,1616]" pageId="11" pageNumber="12" type="nomenclature">
|
||
<paragraph id="D3590C16FFDA6161FF02646AFDF1F9A5" blockId="11.[151,635,1590,1616]" box="[151,635,1590,1616]" pageId="11" pageNumber="12">
|
||
<heading id="8811BB7AFFDA6161FF02646AFDF1F9A5" bold="true" box="[151,635,1590,1616]" fontSize="11" level="1" pageId="11" pageNumber="12" reason="1">
|
||
<emphasis id="E192D004FFDA6161FF02646AFDF1F9A5" bold="true" box="[151,635,1590,1616]" pageId="11" pageNumber="12">
|
||
Genus
|
||
<taxonomicName id="14E67795FFDA6161FF7B646AFDF1F9A5" authority="Benderitter, 1913" authorityName="Benderitter" authorityYear="1913" box="[238,635,1590,1616]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E192D004FFDA6161FF7B646AFE16F9A5" bold="true" box="[238,412,1590,1616]" italics="true" pageId="11" pageNumber="12">Pseudorphnus</emphasis>
|
||
<bibRefCitation id="B77771E7FFDA6161FE31646AFDF1F9A5" author="Benderitter" box="[420,635,1590,1616]" pageId="11" pageNumber="12" refString="Benderitter, E. (1913) Etude sur le genre Orphnus et descriptions de deux Orphnus et d'un Phaeochrous nouveaus (Col. Scarabaeidae). Bulletin de la Societe entomologique de France, 1913, 82 - 85." type="journal article" year="1913">Benderitter, 1913</bibRefCitation>
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFFDA6161FF026421FD86F966" box="[151,524,1661,1683]" pageId="11" pageNumber="12" type="reference_group">
|
||
<paragraph id="D3590C16FFDA6161FF026421FD86F966" blockId="11.[151,814,1661,1714]" box="[151,524,1661,1683]" pageId="11" pageNumber="12">
|
||
<treatmentCitationGroup id="F3F62B38FFDA6161FF026421FD86F966" box="[151,524,1661,1683]" pageId="11" pageNumber="12">
|
||
<taxonomicName id="14E67795FFDA6161FF026421FD8DF966" authority="Benderitter, 1913: 83" authorityName="Benderitter" authorityPageNumber="83" authorityYear="1913" box="[151,519,1661,1683]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E192D004FFDA6161FF026421FEA2F966" box="[151,296,1661,1683]" italics="true" pageId="11" pageNumber="12">Pseudorphnus</emphasis>
|
||
<treatmentCitation id="52472A07FFDA6161FEBB6421FD8DF966" author="Benderitter" box="[302,519,1661,1683]" page="83" pageId="11" pageNumber="12" year="1913">
|
||
<bibRefCitation id="B77771E7FFDA6161FEBB6421FE6BF966" author="Benderitter" box="[302,481,1661,1683]" pageId="11" pageNumber="12" refString="Benderitter, E. (1913) Etude sur le genre Orphnus et descriptions de deux Orphnus et d'un Phaeochrous nouveaus (Col. Scarabaeidae). Bulletin de la Societe entomologique de France, 1913, 82 - 85." type="journal article" year="1913">Benderitter, 1913</bibRefCitation>
|
||
: 83
|
||
</treatmentCitation>
|
||
</taxonomicName>
|
||
.
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFFDA6161FF0264C0FCA4F947" box="[151,814,1692,1714]" pageId="11" pageNumber="12" type="materials_examined">
|
||
<paragraph id="D3590C16FFDA6161FF0264C0FCA4F947" blockId="11.[151,814,1661,1714]" box="[151,814,1692,1714]" pageId="11" pageNumber="12">
|
||
<emphasis id="E192D004FFDA6161FF0264C0FEA8F947" bold="true" box="[151,290,1692,1714]" pageId="11" pageNumber="12">Type species.</emphasis>
|
||
<taxonomicName id="14E67795FFDA6161FEBC64C0FD12F947" authority="Fairmaire, 1868" authorityName="Fairmaire" authorityYear="1868" box="[297,664,1692,1714]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="coquerelii">
|
||
<emphasis id="E192D004FFDA6161FEBC64C0FE67F947" box="[297,493,1692,1714]" italics="true" pageId="11" pageNumber="12">Orphnus coquerelii</emphasis>
|
||
<bibRefCitation id="B77771E7FFDA6161FE6164C0FD12F947" author="Fairmaire" box="[500,664,1692,1714]" pageId="11" pageNumber="12" refString="Fairmaire, L. (1868) Notes sur les Coleopteres receuillis par Charles Coquerel a Madagascar et sur les cotes d'Afrique. Annales de la Societe entomologique de France, 4 (8), 753 - 820." type="journal article" year="1868">Fairmaire, 1868</bibRefCitation>
|
||
</taxonomicName>
|
||
, by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFFDA6166FF026482FD96FE58" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
|
||
<paragraph id="D3590C16FFDA6166FF026482FE79FECA" blockId="11.[151,1437,1758,2035]" lastBlockId="12.[151,1437,151,645]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
<emphasis id="E192D004FFDA6161FF026482FE80F902" bold="true" box="[151,266,1758,1783]" pageId="11" pageNumber="12">Diagnosis</emphasis>
|
||
. Body length
|
||
<quantity id="141EA1F3FFDA6161FE2B6482FDC2F903" box="[446,584,1758,1783]" metricMagnitude="-3" metricUnit="m" metricValue="9.75" metricValueMax="13.0" metricValueMin="6.5" pageId="11" pageNumber="12" unit="mm" value="9.75" valueMax="13.0" valueMin="6.5">6.5–13 mm</quantity>
|
||
, brown to dark brown. Mandibles subsymmetrical. Frontoclypeus with rounded anterior margin, with tubercle or longitudinal ridge in males; females have frontoclypeus without tubercles or ridges. Pronotum with deep excavation in the middle and with 2 horn or ridge-shaped lateral processes bordering excavation near anterior margin in males; females with pronotum convex or slightly impressed anteromedially. The shape of the head and prothoracic armature in males is subject to moderate allometric variability: they vary from fully developed as described above to only a small tubercle on the frontoclypeus and a shallow fossa on the disc of the pronotum anteriorly. Propleurae smoothly convex, without carinae separating anterolateral areas from basal area. Scutellum triangular, narrowly rounded apically, about 1/10 length of elytra. Elytra convex, with feebly marked humeral humps. Sculpture of elytron varies. Wings fully developed. Metepisternum triangular, tapering caudally, with slightly widened anterodorsal angle slightly overlapping epipleuron. Protibiae with 2 strong outer teeth and 1 small tooth located basally. Phallobase strongly sclerotized dorsally and with a thin membrane ventrally. Parameres relatively long apices tapering or curved downwards, without setae. Internal sac of aedeagus with species-specific armature.
|
||
</paragraph>
|
||
<paragraph id="D3590C16FFDD6166FF526310FD96FE58" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="13">
|
||
<emphasis id="E192D004FFDD6166FF526310FE5AFE90" bold="true" box="[199,464,332,357]" pageId="12" pageNumber="13">Diagnostic characters.</emphasis>
|
||
The most reliable characters to differentiate
|
||
<taxonomicName id="14E67795FFDD6166FC506310FBEFFE90" box="[965,1125,332,357]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E192D004FFDD6166FC506310FBEFFE90" box="[965,1125,332,357]" italics="true" pageId="12" pageNumber="13">Pseudorphnus</emphasis>
|
||
</taxonomicName>
|
||
species are the shape of the parameres and the shape of the frontoclypeal and prothoracic armature (especially when fully developed). Body size also varies among the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFFDD6166FF5263EBFCD0FD70" pageId="12" pageNumber="13" type="description">
|
||
<paragraph id="D3590C16FFDD6166FF5263EBFCD0FD70" blockId="12.[151,1437,151,645]" pageId="12" pageNumber="13">
|
||
<materialsCitation id="638E064BFFDD6166FF5263EBFCD0FD70" ID-GBIF-Occurrence="1416610197" country="Madagascar" location="Central Escarpment" pageId="12" pageNumber="13" specimenCount="1">
|
||
<emphasis id="E192D004FFDD6166FF5263EBFD64FE25" bold="true" box="[199,750,439,464]" pageId="12" pageNumber="13">Species composition, distribution, and habitat.</emphasis>
|
||
<taxonomicName id="14E67795FFDD6166FD6163EBFC1EFE25" box="[756,916,439,464]" class="Insecta" family="Orphnidae" genus="Pseudorphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="E192D004FFDD6166FD6163EBFC1EFE25" box="[756,916,439,464]" italics="true" pageId="12" pageNumber="13">Pseudorphnus</emphasis>
|
||
</taxonomicName>
|
||
is endemic to
|
||
<collectingCountry id="ABF14C86FFDD6166FBA363EBFB4AFE25" box="[1078,1216,439,464]" name="Madagascar" pageId="12" pageNumber="13">Madagascar</collectingCountry>
|
||
. Four species of the genus are known so far. The species are distributed chiefly along the eastern slopes of the
|
||
<location id="D6395ACDFFDD6166FB2E6380FA16FE01" LSID="urn:lsid:plazi:treatment:5B4FBD00FFDA6166FF02646AFCD0FD70:D6395ACDFFDD6166FB2E6380FA16FE01" box="[1211,1436,476,501]" country="Madagascar" name="Central Escarpment" pageId="12" pageNumber="13">Central Escarpment</location>
|
||
corresponding to the distribution of the Malagasy tropical wet forests. All species are apparently forest litter dwellers feeding on different rotten substances. One species,
|
||
<taxonomicName id="14E67795FFDD6166FCF46079FC7AFDC8" box="[865,1008,548,573]" class="Insecta" family="Orphnidae" genus="Orphnus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="coquerelii">
|
||
<emphasis id="E192D004FFDD6166FCF46079FC7AFDC8" box="[865,1008,548,573]" italics="true" pageId="12" pageNumber="13">P. coquerelii</emphasis>
|
||
</taxonomicName>
|
||
, is quite common in the collections, which may indicate a secondary adaptation to the developing in the crop fields (similar to
|
||
<taxonomicName id="14E67795FFDD6166FB1D601BFAE0FD95" box="[1160,1386,583,608]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="nitidulus">
|
||
<emphasis id="E192D004FFDD6166FB1D601BFAE0FD95" box="[1160,1386,583,608]" italics="true" pageId="12" pageNumber="13">Triodontus nitidulus</emphasis>
|
||
</taxonomicName>
|
||
, see below). Nesting behavior and preimaginal stages are unknown.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |