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<mods:title id="16E22DEA5C6E5BC8FEF8DCBC87EFD186">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<taxonomicName id="14E67795FF9E6125FF0260E8FDA7FD3B" authority="Paulian, 1977" authorityName="Paulian" authorityYear="1977" box="[151,557,692,718]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E192D004FF9E6125FF0260E8FDA7FD3B" bold="true" box="[151,557,692,718]" pageId="79" pageNumber="80">
<emphasis id="E192D004FF9E6125FF0260E8FEF7FD3B" bold="true" box="[151,381,692,718]" italics="true" pageId="79" pageNumber="80">Triodontus alticola</emphasis>
<bibRefCitation id="B77771E7FF9E6125FE1160E8FDA7FD3B" author="Paulian" box="[388,557,692,718]" pageId="79" pageNumber="80" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
</emphasis>
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</heading>
</paragraph>
<paragraph id="D3590C16FF9E6125FF02608AFEB0FD1A" blockId="79.[151,557,692,751]" box="[151,314,726,751]" pageId="79" pageNumber="80">
(
<figureCitation id="4BDD1093FF9E6125FF0A608AFF73FD1A" box="[159,249,726,751]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="79" pageNumber="80">Figs. 41</figureCitation>
AH)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9E6125FF026142FC84FCA6" pageId="79" pageNumber="80" type="reference_group">
<paragraph id="D3590C16FF9E6125FF026142FDBAFCC1" blockId="79.[151,782,798,851]" box="[151,560,798,820]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="F3F62B38FF9E6125FF026142FDBAFCC1" box="[151,560,798,820]" pageId="79" pageNumber="80">
<taxonomicName id="14E67795FF9E6125FF026142FDA7FCC1" authority="Paulian, 1977: 1215" authorityName="Paulian" authorityPageNumber="1215" authorityYear="1977" box="[151,557,798,820]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E192D004FF9E6125FF026142FED2FCC1" box="[151,344,798,820]" italics="true" pageId="79" pageNumber="80">Triodontus alticola</emphasis>
<treatmentCitation id="52472A07FF9E6125FECB6142FDA7FCC1" author="Paulian" box="[350,557,798,820]" page="1215" pageId="79" pageNumber="80" year="1977">
<bibRefCitation id="B77771E7FF9E6125FECB6142FE67FCC1" author="Paulian" box="[350,493,798,820]" pageId="79" pageNumber="80" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
: 1215
</treatmentCitation>
</taxonomicName>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="D3590C16FF9E6125FF026161FC84FCA6" blockId="79.[151,782,798,851]" box="[151,782,829,851]" pageId="79" pageNumber="80">
<treatmentCitationGroup id="F3F62B38FF9E6125FF026161FC84FCA6" box="[151,782,829,851]" pageId="79" pageNumber="80">
<taxonomicName id="14E67795FF9E6125FF026161FDFBFCA6" authority="Paulian, 1977: 1219" authorityName="Paulian" authorityPageNumber="1219" authorityYear="1977" box="[151,625,829,851]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="maroantsetrae">
<emphasis id="E192D004FF9E6125FF026161FE14FCA6" box="[151,414,829,851]" italics="true" pageId="79" pageNumber="80">Triodontus maroantsetrae</emphasis>
<treatmentCitation id="52472A07FF9E6125FE316161FDFBFCA6" author="Paulian" box="[420,625,829,851]" page="1219" pageId="79" pageNumber="80" year="1977">
<bibRefCitation id="B77771E7FF9E6125FE316161FDB8FCA6" author="Paulian" box="[420,562,829,851]" pageId="79" pageNumber="80" refString="Paulian, R. (1977) Les Orphnidae de Madagascar. Bulletin du Museum National d'histoire naturelle, Paris. Series 3, 411, 1199 - 1223." type="journal article" year="1977">Paulian, 1977</bibRefCitation>
: 1219
</treatmentCitation>
</taxonomicName>
,
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<emphasis id="E192D004FF9E6125FDE96161FC84FCA6" bold="true" box="[636,782,829,851]" pageId="79" pageNumber="80">new synonym</emphasis>
</taxonomicNameLabel>
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</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9E6125FF026123FB42FA65" pageId="79" pageNumber="80" type="materials_examined">
<paragraph id="D3590C16FF9E6125FF026123FA89FC49" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">
<materialsCitation id="638E064BFF9E6125FF026123FA89FC49" ID-GBIF-Occurrence="1416610143" collectingDate="1949-11" collectionCode="TYPE, MNHN" collectorName="Michel" country="Madagascar" determinerName="R. Paulian" location="Toamasina Province" pageId="79" pageNumber="80" specimenCount="3" stateProvince="Toamasina Province" typeStatus="holotype">
<emphasis id="E192D004FF9E6125FF026123FE4EFC6D" bold="true" box="[151,452,895,920]" pageId="79" pageNumber="80">
<typeStatus id="0C5DB2B4FF9E6125FF026123FF58FC6D" box="[151,210,895,920]" pageId="79" pageNumber="80">Type</typeStatus>
material examined.
</emphasis>
<taxonomicName id="14E67795FF9E6125FE586123FD2FFC6D" box="[461,677,895,920]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E192D004FF9E6125FE586123FD2FFC6D" box="[461,677,895,920]" italics="true" pageId="79" pageNumber="80">Triodontus alticola</emphasis>
</taxonomicName>
.
<typeStatus id="0C5DB2B4FF9E6125FD206123FC97FC6D" box="[693,797,895,920]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FF9E6125FCBA6123FC04FC6D" box="[815,910,895,920]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="79" pageNumber="80">Figs. 41</figureCitation>
C, D), male, “
<collectingCountry id="ABF14C86FF9E6125FBBA6123FB33FC6D" box="[1071,1209,895,920]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ambohitsitondrona [
<collectingRegion id="1122C2F4FF9E6125FF0861F8FE0DFC48" box="[157,391,932,957]" country="Madagascar" name="Toamasina" pageId="79" pageNumber="80">Toamasina Province</collectingRegion>
]
<collectingDate id="B71CD33EFF9E6125FE0161F8FE78FC49" box="[404,498,932,956]" pageId="79" pageNumber="80" value="1949-11">
<date id="A7582AD6FF9E6125FE0161F8FE78FC49" box="[404,498,932,956]" pageId="79" pageNumber="80" value="1949-11">XI-1949</date>
</collectingDate>
<collectorName id="7E1369C0FF9E6125FE6F61F8FDC3FC48" box="[506,585,932,957]" pageId="79" pageNumber="80">Michel</collectorName>
/
<taxonomicName id="14E67795FF9E6125FDCA61F8FCBFFC48" box="[607,821,932,957]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="alticola" status="sp. nov.">
<emphasis id="E192D004FF9E6125FDCA61F8FCBFFC48" box="[607,821,932,957]" italics="true" pageId="79" pageNumber="80">Triodontus alticola</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAA16D7FFF9E6125FCA961F9FCF2FC49" box="[828,888,933,956]" pageId="79" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="2685E0D2FF9E6125FCEA61F9FC79FC48" box="[895,1011,932,957]" pageId="79" pageNumber="80">R. Paulian</determinerName>
det. /
<collectionCode id="B5F794D3FF9E6125FBAC61F8FB0BFC49" box="[1081,1153,932,956]" pageId="79" pageNumber="80">
<typeStatus id="0C5DB2B4FF9E6125FBAC61F8FB0BFC49" box="[1081,1153,932,956]" pageId="79" pageNumber="80">TYPE</typeStatus>
</collectionCode>
” (
<collectionCode id="B5F794D3FF9E6125FB0F61F8FB7CFC49" box="[1178,1270,932,956]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D3590C16FF9E6125FF526194FA0FFBDD" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">
<materialsCitation id="638E064BFF9E6125FF526194FC7BFC15" ID-GBIF-Occurrence="1416610150" box="[199,1009,967,992]" collectionCode="MNHN" pageId="79" pageNumber="80" specimenCount="1" typeStatus="paratype">
<typeStatus id="0C5DB2B4FF9E6125FF526194FEB3FC15" box="[199,313,968,992]" pageId="79" pageNumber="80" type="paratype">Paratypes</typeStatus>
: two females with the same locality and date label (
<collectionCode id="B5F794D3FF9E6125FC186194FC63FC15" box="[909,1001,968,992]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FC6B6194FD54FBF1" ID-GBIF-Occurrence="1416610139" collectingDate="1952-12" collectionCode="MNHN" pageId="79" pageNumber="80" specimenCount="1" typeStatus="paratype">
two males and two females with the same locality label but the date “
<collectingDate id="B71CD33EFF9E6125FD9A61B0FDD5FBF0" box="[527,607,1004,1029]" pageId="79" pageNumber="80" value="1952-12">
<date id="A7582AD6FF9E6125FD9A61B0FDD5FBF0" box="[527,607,1004,1029]" pageId="79" pageNumber="80" value="1952-12">XII-52</date>
</collectingDate>
” (
<collectionCode id="B5F794D3FF9E6125FDEF61B0FD5CFBF1" box="[634,726,1004,1028]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FD7961B1FE1CFBDD" ID-GBIF-Occurrence="1416610105" collectionCode="MNHN" country="Madagascar" location="Vadon" pageId="79" pageNumber="80" specimenCount="1" stateProvince="Antsiranana Province" typeStatus="paratype">
one male, “
<collectingCountry id="ABF14C86FF9E6125FCF861B0FC7DFBF0" box="[877,1015,1004,1029]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Sambava [
<collectingRegion id="1122C2F4FF9E6125FBE061B0FAFAFBF0" box="[1141,1392,1004,1029]" country="Madagascar" name="Antsiranana" pageId="79" pageNumber="80">Antsiranana Province</collectingRegion>
] XI [...]
<location id="D6395ACDFF9E6125FF506653FE9EFBDD" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9E613BFF0260E8FD71FCCD:D6395ACDFF9E6125FF506653FE9EFBDD" box="[197,276,1039,1064]" country="Madagascar" name="Vadon" pageId="79" pageNumber="80" stateProvince="Antsiranana Province">Vadon</location>
!” (
<collectionCode id="B5F794D3FF9E6125FEA4664CFE07FBDD" box="[305,397,1040,1064]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FE34664CFA0BFBDD" ID-GBIF-Occurrence="1416610104" box="[417,1409,1039,1064]" collectingDate="1950-01" collectionCode="MNHN" country="Madagascar" location="Vadon" pageId="79" pageNumber="80" specimenCount="1" stateProvince="Toamasina Province" typeStatus="paratype">
one male, “
<collectingCountry id="ABF14C86FF9E6125FD8B6653FD22FBDD" box="[542,680,1039,1064]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ambodivoangy [
<collectingRegion id="1122C2F4FF9E6125FCFE6653FBDFFBDD" box="[875,1109,1039,1064]" country="Madagascar" name="Toamasina" pageId="79" pageNumber="80">Toamasina Province</collectingRegion>
]
<date id="A7582AD6FF9E6125FBF46653FB23FBDD" box="[1121,1193,1039,1064]" pageId="79" pageNumber="80" value="1950-01">
<collectingDate id="B71CD33EFF9E6125FBF46653FB23FBDD" box="[1121,1193,1039,1064]" pageId="79" pageNumber="80" value="1950-01">I.1950</collectingDate>
</date>
<location id="D6395ACDFF9E6125FB246653FA8BFBDD" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9E613BFF0260E8FD71FCCD:D6395ACDFF9E6125FB246653FA8BFBDD" box="[1201,1281,1039,1064]" country="Madagascar" name="Vadon" pageId="79" pageNumber="80" stateProvince="Toamasina Province">Vadon</location>
!” (
<collectionCode id="B5F794D3FF9E6125FA88664CFAF3FBDD" box="[1309,1401,1040,1064]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D3590C16FF9E6125FF526668FA18FB85" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">
<materialsCitation id="638E064BFF9E6125FF526668FA04FB85" ID-GBIF-Occurrence="1416610145" collectingDate="1935-12" collectionCode="MUSEUM, PARIS, TYPE, MNHN" collectorName="J. Vadon &amp; E. Lebis" country="Madagascar" determinerName="R. Paulian" location="Vadon Antakotako" pageId="79" pageNumber="80" specimenCode="PARIS 1938" specimenCount="2" typeStatus="holotype">
<taxonomicName id="14E67795FF9E6125FF526668FE7BFBB9" box="[199,497,1076,1101]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="maroantsetrae">
<emphasis id="E192D004FF9E6125FF526668FE7BFBB9" box="[199,497,1076,1101]" italics="true" pageId="79" pageNumber="80">Triodontus maroantsetrae</emphasis>
</taxonomicName>
.
<typeStatus id="0C5DB2B4FF9E6125FD926668FDF8FBB8" box="[519,626,1076,1101]" pageId="79" pageNumber="80" type="holotype">Holotype</typeStatus>
, male, “
<collectingCountry id="ABF14C86FF9E6125FD4E6668FCEFFBB8" box="[731,869,1076,1101]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ron Maroantsetra
<date id="A7582AD6FF9E6125FBC16668FB14FBB8" box="[1108,1182,1076,1101]" pageId="79" pageNumber="80" value="1935-12">
<collectingDate id="B71CD33EFF9E6125FBC16668FB14FBB8" box="[1108,1182,1076,1101]" pageId="79" pageNumber="80" value="1935-12">XII.35</collectingDate>
</date>
<location id="D6395ACDFF9E6125FB386668FA0CFBB8" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9E613BFF0260E8FD71FCCD:D6395ACDFF9E6125FB386668FA0CFBB8" box="[1197,1414,1076,1101]" country="Madagascar" name="Vadon Antakotako" pageId="79" pageNumber="80">Vadon Antakotako</location>
/
<collectionCode id="B5F794D3FF9E6125FF026604FE98FB85" box="[151,274,1112,1136]" pageId="79" pageNumber="80">MUSEUM</collectionCode>
<specimenCode id="8340A46DFF9E6125FE8C6604FE2EFB85" box="[281,420,1112,1136]" collectionCode="PARIS" pageId="79" pageNumber="80">
<collectionCode id="B5F794D3FF9E6125FE8C6604FEECFB85" box="[281,358,1112,1136]" pageId="79" pageNumber="80">PARIS</collectionCode>
1938
</specimenCode>
<collectorName id="7E1369C0FF9E6125FE396604FD86FB85" box="[428,524,1111,1136]" pageId="79" pageNumber="80">J. Vadon</collectorName>
et
<collectorName id="7E1369C0FF9E6125FDBB6604FD01FB85" box="[558,651,1111,1136]" pageId="79" pageNumber="80">E. Lebis</collectorName>
/
<collectionCode id="B5F794D3FF9E6125FD346604FD6DFB9A" box="[673,743,1112,1135]" pageId="79" pageNumber="80">
<typeStatus id="0C5DB2B4FF9E6125FD346604FD6DFB9A" box="[673,743,1112,1135]" pageId="79" pageNumber="80">TYPE</typeStatus>
</collectionCode>
/
<taxonomicName id="14E67795FF9E6125FD68660BFCFDFB85" box="[765,887,1111,1136]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="genus">Triodontus</taxonomicName>
<taxonomicName id="14E67795FF9E6125FCEB6605FBAAFB85" box="[894,1056,1113,1136]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="maroantsetrae" status="sp. nov.">
<emphasis id="E192D004FF9E6125FCEB6605FBAAFB85" box="[894,1056,1113,1136]" italics="true" pageId="79" pageNumber="80">maroantsetrae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FAA16D7FFF9E6125FBB26604FBE9FB9A" box="[1063,1123,1112,1135]" pageId="79" pageNumber="80" rank="species">n. sp.</taxonomicNameLabel>
<determinerName id="2685E0D2FF9E6125FBFF6604FB55FB85" box="[1130,1247,1111,1136]" pageId="79" pageNumber="80">R. Paulian</determinerName>
det.” (
<collectionCode id="B5F794D3FF9E6125FABF6604FA0CFB85" box="[1322,1414,1112,1136]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D3590C16FF9E6125FF526620FB42FA65" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">
<materialsCitation id="638E064BFF9E6125FF526620FC85FB4D" ID-GBIF-Occurrence="1416610141" collectingDate="1948-02" collectionCode="MNHN" collectorName="Michel" country="Madagascar" latitude="-15.5830555" location="Madagascar Ambohitsitondrona" longLatPrecision="21" longitude="50.016666" pageId="79" pageNumber="80" specimenCount="1" stateProvince="Toamasina">
<emphasis id="E192D004FF9E6125FF526620FC28FB60" bold="true" box="[199,930,1148,1173]" pageId="79" pageNumber="80">
Additional material examined.
<collectingCountry id="ABF14C86FF9E6125FDAC6620FC81FB60" box="[569,779,1148,1173]" name="Madagascar" pageId="79" pageNumber="80">MADAGASCAR</collectingCountry>
:
<collectingRegion id="1122C2F4FF9E6125FC826620FC17FB60" box="[791,925,1148,1173]" country="Madagascar" name="Toamasina" pageId="79" pageNumber="80">Toamasina</collectingRegion>
:
</emphasis>
two males, “
<location id="D6395ACDFF9E6125FBA76620FA16FB60" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9E613BFF0260E8FD71FCCD:D6395ACDFF9E6125FBA76620FA16FB60" box="[1074,1436,1148,1173]" country="Madagascar" latitude="-15.5830555" longLatPrecision="21" longitude="50.016666" name="Madagascar Ambohitsitondrona" pageId="79" pageNumber="80" stateProvince="Toamasina">Madagascar Ambohitsitondrona</location>
<collectingDate id="B71CD33EFF9E6125FF0266FCFF61FB4D" box="[151,235,1184,1208]" pageId="79" pageNumber="80" value="1948-02">
<date id="A7582AD6FF9E6125FF0266FCFF61FB4D" box="[151,235,1184,1208]" pageId="79" pageNumber="80" value="1948-02">II-1948</date>
</collectingDate>
<collectorName id="7E1369C0FF9E6125FF6E66C3FEC0FB4D" box="[251,330,1183,1208]" pageId="79" pageNumber="80">Michel</collectorName>
[
<geoCoordinate id="B6D26AD1FF9E6125FEF466C3FE6DFB4D" box="[353,487,1183,1208]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-15.5830555">15°3459″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FE6E66C3FD0BFB4D" box="[507,641,1183,1208]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="50.016666">50°0100″E</geoCoordinate>
]” (
<collectionCode id="B5F794D3FF9E6125FD3E66FCFC8DFB4D" box="[683,775,1184,1208]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FCB666FCFDD8FB29" ID-GBIF-Occurrence="1416610144" collectingDate="1946-01" collectionCode="MNHN" country="Madagascar" latitude="-15.296945" location="Madagascar" longLatPrecision="21" longitude="49.613052" pageId="79" pageNumber="80" specimenCount="1">
one male, “
<collectingCountry id="ABF14C86FF9E6125FC2666C3FBB7FB4D" box="[947,1085,1183,1208]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ambodivoangy [
<geoCoordinate id="B6D26AD1FF9E6125FA8766C3FA1DFB4D" box="[1298,1431,1183,1208]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-15.296945">15°1749″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FF026698FE96FB28" box="[151,284,1220,1245]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="49.613052">49°3647″E</geoCoordinate>
]
<collectingDate id="B71CD33EFF9E6125FEB96698FEFEFB28" box="[300,372,1220,1245]" pageId="79" pageNumber="80" value="1946-01">
<date id="A7582AD6FF9E6125FEB96698FEFEFB28" box="[300,372,1220,1245]" pageId="79" pageNumber="80" value="1946-01">I.1946</date>
</collectingDate>
Vadon!” (
<collectionCode id="B5F794D3FF9E6125FE7A6698FDC1FB29" box="[495,587,1220,1244]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
; one male, same label but
<date id="A7582AD6FF9E6125FC1B6698FC7EFB29" box="[910,1012,1220,1244]" pageId="79" pageNumber="80" value="1947-12">XII.1947</date>
(MNHN);
<materialsCitation id="638E064BFF9E6125FBED6699FCFFFAF5" ID-GBIF-Occurrence="1416610134" collectingDate="1946-02" collectionCode="MNHN" country="Madagascar" latitude="-15.316667" location="Madagascar" longLatPrecision="21" longitude="49.8" pageId="79" pageNumber="80" specimenCount="1">
one female, “
<collectingCountry id="ABF14C86FF9E6125FA876698FA16FB28" box="[1298,1436,1220,1245]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Antakotako [
<geoCoordinate id="B6D26AD1FF9E6125FEBD66BBFE27FAF5" box="[296,429,1255,1280]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-15.316667">15°1900″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FE2C66BBFDB4FAF5" box="[441,574,1255,1280]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="49.8">49°4800″E</geoCoordinate>
]
<collectingDate id="B71CD33EFF9E6125FDD966BBFD17FAF5" box="[588,669,1255,1280]" pageId="79" pageNumber="80" value="1946-02">
<date id="A7582AD6FF9E6125FDD966BBFD17FAF5" box="[588,669,1255,1280]" pageId="79" pageNumber="80" value="1946-02">II.1946</date>
</collectingDate>
Vadon!” (
<collectionCode id="B5F794D3FF9E6125FC8466B4FCE7FAF5" box="[785,877,1256,1280]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FC1566B4FA9AFAD1" ID-GBIF-Occurrence="1416610128" collectionCode="MNHN" collectorName="N. Antanambe Vadon" country="Madagascar" latitude="-16.433054" location="Mananara District" longLatPrecision="21" longitude="49.85" pageId="79" pageNumber="80" specimenCount="1">
one male, “
<collectingCountry id="ABF14C86FF9E6125FC6B66BBFB02FAF5" box="[1022,1160,1255,1280]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Est distr. Mananara -
<collectorName id="7E1369C0FF9E6125FA1466B4FEE2FAD0" pageId="79" pageNumber="80">N. Antanambe Vadon</collectorName>
et Peyrieras [
<location id="D6395ACDFF9E6125FD916751FD5CFAD0" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9E613BFF0260E8FD71FCCD:D6395ACDFF9E6125FD916751FD5CFAD0" box="[516,726,1292,1317]" country="Madagascar" latitude="-16.433054" longLatPrecision="21" longitude="49.85" name="Mananara District" pageId="79" pageNumber="80">Mananara District</location>
, Antanambe,
<geoCoordinate id="B6D26AD1FF9E6125FCE16750FC7DFAD0" box="[884,1015,1292,1317]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-16.433054">16°2559″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FB916750FB01FAD0" box="[1028,1163,1292,1317]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="49.85">49°5100″E</geoCoordinate>
]” (
<collectionCode id="B5F794D3FF9E6125FB396750FA82FAD1" box="[1196,1288,1292,1316]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FA886750FA92FABD" ID-GBIF-Occurrence="1416610140" collectingDate="1946-01" collectionCode="MNHN" collectorName="Vadon" country="Madagascar" latitude="-15.5830555" location="Madagascar" longLatPrecision="21" longitude="50.016666" pageId="79" pageNumber="80" specimenCount="1">
four males, one female, “
<collectingCountry id="ABF14C86FF9E6125FEA36773FE4AFABD" box="[310,448,1327,1352]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ambohitsitondrona
<collectingDate id="B71CD33EFF9E6125FD216773FD74FABD" box="[692,766,1327,1352]" pageId="79" pageNumber="80" value="1946-01">
<date id="A7582AD6FF9E6125FD216773FD74FABD" box="[692,766,1327,1352]" pageId="79" pageNumber="80" value="1946-01">I-1946</date>
</collectingDate>
<collectorName id="7E1369C0FF9E6125FC996773FCD0FABD" box="[780,858,1327,1352]" pageId="79" pageNumber="80">Vadon</collectorName>
! [
<geoCoordinate id="B6D26AD1FF9E6125FCE46773FC7CFABD" box="[881,1014,1327,1352]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-15.5830555">15°3459″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FB9D6773FB07FABD" box="[1032,1165,1327,1352]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="50.016666">50°0100″E</geoCoordinate>
]” (
<collectionCode id="B5F794D3FF9E6125FB21676CFA9AFABD" box="[1204,1296,1328,1352]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FABC676CFB7DFA99" ID-GBIF-Occurrence="1416610126" collectingDate="1948-01-25" collectionCode="MNHN" country="Madagascar" latitude="-15.5830555" location="Madagascar" longLatPrecision="21" longitude="50.016666" pageId="79" pageNumber="80" specimenCount="1">
one male, “
<collectingCountry id="ABF14C86FF9E6125FF346708FEA1FA98" box="[161,299,1364,1389]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ambohitsitondrona
<collectingDate id="B71CD33EFF9E6125FD846708FD0BFA98" box="[529,641,1364,1389]" pageId="79" pageNumber="80" value="1948-01-25">
<date id="A7582AD6FF9E6125FD846708FD0BFA98" box="[529,641,1364,1389]" pageId="79" pageNumber="80" value="1948-01-25">25-I-1948</date>
</collectingDate>
Michel [
<geoCoordinate id="B6D26AD1FF9E6125FD716708FCE3FA98" box="[740,873,1364,1389]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-15.5830555">15°3459″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FCE16708FC73FA98" box="[884,1017,1364,1389]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="50.016666">50°0100″E</geoCoordinate>
,
<collectingDate id="B71CD33EFF9E6125FB966708FBF8FA98" box="[1027,1138,1364,1389]" pageId="79" pageNumber="80" value="1948-01-25">
<date id="A7582AD6FF9E6125FB966708FBF8FA98" box="[1027,1138,1364,1389]" pageId="79" pageNumber="80" value="1948-01-25">25.I.1948</date>
</collectingDate>
]” (
<collectionCode id="B5F794D3FF9E6125FB066708FB65FA99" box="[1171,1263,1364,1388]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="638E064BFF9E6125FA946709FB4EFA65" ID-GBIF-Occurrence="1416610154" collectingDate="1947-10" collectionCode="MNHN" collectorName="Vadon" country="Madagascar" latitude="-15.5830555" location="Madagascar" longLatPrecision="21" longitude="50.016666" pageId="79" pageNumber="80" specimenCount="1">
one male, one female, “
<collectingCountry id="ABF14C86FF9E6125FF6E672BFE0FFA65" box="[251,389,1399,1424]" name="Madagascar" pageId="79" pageNumber="80">Madagascar</collectingCountry>
Ambohitsitondrona
<collectingDate id="B71CD33EFF9E6125FDF86724FD49FA65" box="[621,707,1400,1424]" pageId="79" pageNumber="80" value="1947-10">
<date id="A7582AD6FF9E6125FDF86724FD49FA65" box="[621,707,1400,1424]" pageId="79" pageNumber="80" value="1947-10">X-1947</date>
</collectingDate>
<collectorName id="7E1369C0FF9E6125FD5C672BFC9DFA65" box="[713,791,1399,1424]" pageId="79" pageNumber="80">Vadon</collectorName>
! [
<geoCoordinate id="B6D26AD1FF9E6125FCBD672BFC27FA65" box="[808,941,1399,1424]" direction="south" orientation="latitude" pageId="79" pageNumber="80" precision="15" value="-15.5830555">15°3459″S</geoCoordinate>
,
<geoCoordinate id="B6D26AD1FF9E6125FC2C672BFBCAFA65" box="[953,1088,1399,1424]" direction="east" orientation="longitude" pageId="79" pageNumber="80" precision="15" value="50.016666">50°0100″E</geoCoordinate>
]” (
<collectionCode id="B5F794D3FF9E6125FBCA6724FB31FA65" box="[1119,1211,1400,1424]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="79" pageNumber="80">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9E6125FF5267C0FB40FA09" pageId="79" pageNumber="80" type="diagnosis">
<paragraph id="D3590C16FF9E6125FF5267C0FB40FA09" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">
<emphasis id="E192D004FF9E6125FF5267C0FECBFA40" bold="true" box="[199,321,1436,1461]" pageId="79" pageNumber="80">Diagnosis.</emphasis>
<taxonomicName id="14E67795FF9E6125FEDC67C0FDAAFA40" box="[329,544,1436,1461]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="79" pageNumber="80" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E192D004FF9E6125FEDC67C0FDAAFA40" box="[329,544,1436,1461]" italics="true" pageId="79" pageNumber="80">Triodontus alticola</emphasis>
</taxonomicName>
can be separated from other species of the genus by the characteristic shape of the parameres having the outer lobes long, curved downward and inward; with more-or-less separated, strongly sclerotized lateral parts and membranous, overlapping medial parts; and without lateral notches.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9E613BFF52645BFB0AFDC8" lastPageId="81" lastPageNumber="82" pageId="79" pageNumber="80" type="description">
<paragraph id="D3590C16FF9E6125FF52645BFE50F9B1" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">
<emphasis id="E192D004FF9E6125FF52645BFE29F9D5" bold="true" box="[199,419,1543,1568]" pageId="79" pageNumber="80">Description. Male.</emphasis>
Color of head, pronotum, and elytra dark brown, legs, antennae, and underside of the body slightly paler (
<figureCitation id="4BDD1093FF9E6125FEAC6470FE1FF9B1" box="[313,405,1580,1605]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="79" pageNumber="80">Figs. 41</figureCitation>
A, C).
</paragraph>
<paragraph id="D3590C16FF9E6125FF526413FBD5F90D" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">Frontoclypeus slightly convex anteriorly, obtusely rounded laterally, anterior margin setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small, dorsal and large, ventral parts. Frontoclypeus with tubercles mediad of each eye and with a horn in center. The horn is longer than width of the head, acutely rounded apically, somewhat curved caudally, with slightly flattened anterior side and slightly rugose posterior side.</paragraph>
<paragraph id="D3590C16FF9E6125FF526559FB46F859" blockId="79.[151,1437,895,2001]" pageId="79" pageNumber="80">Pronotum with a bulge medially in center, with excavations each side of bulge, and with 2 ridge-shaped tubercles laterally of each excavation. The median bulge with 2 distinct tubercles and somewhat excavated between the tubercles. Lateral margins with wide border bearing a row of brown setae. Anterior margin with wide, smooth border, almost not sinuate medially. Posterior margin with fine border, not crenulate, punctate with a few narrow, longitudinal punctures. Surface of most pronotum punctate with minute, feebly visible puctures.</paragraph>
<paragraph id="D3590C16FF9E6125FF5265E4FC5CF824" blockId="79.[151,1437,895,2001]" box="[199,982,1976,2001]" pageId="79" pageNumber="80">Scutellum rounded apically, visible part is about 1/15 length of elytra.</paragraph>
<caption id="87995C9EFF81613AFF02653DFE15F841" httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="80" pageNumber="81" targetBox="[193,1375,246,1865]" targetPageId="80">
<paragraph id="D3590C16FF81613AFF02653DFE15F841" blockId="80.[151,1436,1889,1972]" pageId="80" pageNumber="81">
<emphasis id="E192D004FF81613AFF02653DFE95F882" bold="true" box="[151,287,1889,1911]" pageId="80" pageNumber="81">FIGURE 41.</emphasis>
<taxonomicName id="14E67795FF81613AFEB0653DFE6CF882" box="[293,486,1889,1911]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="80" pageNumber="81" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E192D004FF81613AFEB0653DFE6CF882" box="[293,486,1889,1911]" italics="true" pageId="80" pageNumber="81">Triodontus alticola</emphasis>
</taxonomicName>
, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H).
</paragraph>
</caption>
<paragraph id="D3590C16FF80613BFF5262CBFC85FF02" blockId="81.[151,1437,151,824]" pageId="81" pageNumber="82">Elytra convex, with distinct humeral humps. Maximum width approximately at middle. Elytra with 10 striae; stria 1 (sutural) distinct, other feebly marked with small punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures.</paragraph>
<paragraph id="D3590C16FF80613BFF526358FAA7FEE9" blockId="81.[151,1437,151,824]" box="[199,1325,260,285]" pageId="81" pageNumber="82">
Abdominal sternite 8 medially deeply sinuate, without tubercles or concavities in middle (
<figureCitation id="4BDD1093FF80613BFB2D6358FA81FEE9" box="[1208,1291,260,285]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="81" pageNumber="82">Fig. 41</figureCitation>
G).
</paragraph>
<paragraph id="D3590C16FF80613BFF526374FD76FE7D" blockId="81.[151,1437,151,824]" pageId="81" pageNumber="82">
Parameres (
<figureCitation id="4BDD1093FF80613BFEC5637BFE2DFEB5" box="[336,423,295,320]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="81" pageNumber="82">Fig. 41</figureCitation>
E) with outer lobes long, curved downward and inward; with more-or-less separated, strongly sclerotized lateral parts and membranous, overlapping medial parts; and without lateral notches. Internal sac of aedeagus with a hoof-shaped sclerite (
<figureCitation id="4BDD1093FF80613BFD1E6333FD54FE7D" box="[651,734,367,392]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="81" pageNumber="82">Fig. 41</figureCitation>
F).
</paragraph>
<paragraph id="D3590C16FF80613BFF5263C8FE20FE00" blockId="81.[151,1437,151,824]" pageId="81" pageNumber="82">
<emphasis id="E192D004FF80613BFF5263C8FEAFFE58" bold="true" box="[199,293,404,429]" pageId="81" pageNumber="82">Female.</emphasis>
Females (
<figureCitation id="4BDD1093FF80613BFE0D63C8FE60FE59" box="[408,490,404,429]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="81" pageNumber="82">Fig. 41</figureCitation>
B) differ from males in having a protibial apical spur; absence of head and pronotum armature including tubercles mediad of each eye; and longer abdominal sternites including 6th sternite, which is evenly rounded apically.
</paragraph>
<paragraph id="D3590C16FF80613BFF5263A3FB0AFDC8" blockId="81.[151,1437,151,824]" pageId="81" pageNumber="82">
<emphasis id="E192D004FF80613BFF5263A3FEB6FDED" bold="true" box="[199,316,511,536]" pageId="81" pageNumber="82">Variation.</emphasis>
The pronotal and head armature of males is less developed in some specimens. Body length of examined specimens varied from 8.812.0 mm (males) and from 9.011.0 mm (females).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF80613BFF52601BFE39FD71" pageId="81" pageNumber="82" type="distribution">
<paragraph id="D3590C16FF80613BFF52601BFE39FD71" blockId="81.[151,1437,151,824]" pageId="81" pageNumber="82">
<emphasis id="E192D004FF80613BFF52601BFEEAFD95" bold="true" box="[199,352,583,608]" pageId="81" pageNumber="82">Distribution.</emphasis>
This species is known from a few localities in the northeastern part of the island, mostly north of Antongil Bay (
<figureCitation id="4BDD1093FF80613BFEAB6030FE05FD71" box="[318,399,620,645]" captionStart="FIGURE 41" captionStartId="80.[151,250,1889,1911]" captionTargetBox="[193,1375,246,1865]" captionTargetId="figure@80.[176,1411,193,1868]" captionTargetPageId="80" captionText="FIGURE 41. Triodontus alticola, holotype male (C, D, E, F), male (A), female (B). Habitus in dorsal view (A, B, C), aedeagus in lateral view and parameres in dorsal and ventral view (E), labels of holotype (D), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200484/files/figure.png" pageId="81" pageNumber="82">Fig. 41</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF80613BFF5260D3FD71FCCD" pageId="81" pageNumber="82" type="discussion">
<paragraph id="D3590C16FF80613BFF5260D3FD71FCCD" blockId="81.[151,1437,151,824]" pageId="81" pageNumber="82">
<emphasis id="E192D004FF80613BFF5260D3FEB1FD5D" bold="true" box="[199,315,655,680]" pageId="81" pageNumber="82">Remarks.</emphasis>
<taxonomicName id="14E67795FF80613BFED460D3FDE9FD5D" box="[321,611,655,680]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="maroantsetrae">
<emphasis id="E192D004FF80613BFED460D3FDE9FD5D" box="[321,611,655,680]" italics="true" pageId="81" pageNumber="82">Triodontus maroantsetrae</emphasis>
</taxonomicName>
was described from one male specimen from Antakotako. Comparison of this specimen with the
<typeStatus id="0C5DB2B4FF80613BFE0F60E9FE40FD38" box="[410,458,693,717]" pageId="81" pageNumber="82">type</typeStatus>
series of
<taxonomicName id="14E67795FF80613BFDA260E9FD23FD38" box="[567,681,692,717]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="alticola">
<emphasis id="E192D004FF80613BFDA260E9FD23FD38" box="[567,681,692,717]" italics="true" pageId="81" pageNumber="82">T. alticola</emphasis>
</taxonomicName>
has shown that they have similar shape of the parameres. The only difference is the feebly developed frontoclypeal horn and almost absent ridges and tubercles on the pronotum in the
<typeStatus id="0C5DB2B4FF80613BFF0260A0FF73FCE0" box="[151,249,764,789]" pageId="81" pageNumber="82" type="holotype">holotype</typeStatus>
of
<taxonomicName id="14E67795FF80613BFEB560A1FE54FCE1" box="[288,478,765,788]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="81" pageNumber="82" phylum="Arthropoda" rank="species" species="maroantsetrae">
<emphasis id="E192D004FF80613BFEB560A1FE54FCE1" box="[288,478,765,788]" italics="true" pageId="81" pageNumber="82">T. maroantsetrae</emphasis>
</taxonomicName>
. We believe all these specimens fall within the limits of allometric variability of one species, therefore the new synonymy is here proposed.
</paragraph>
</subSubSection>
</treatment>
</document>