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<document id="91A430D01442AA185284D805ECA2592D" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier &amp; Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FF9A6127FF02655EFDD4FAF8" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triodontus viettei Frolov, Montreuil &amp; Akhmetova, new species" docType="treatment" docVersion="16" lastPageNumber="78" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="76" updateTime="1698433759527" updateUser="ExternalLinkService">
<mods:mods id="837BF335BEE12F09EE7E6879218B3315" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="3DADA4ABDC8BB63CCD8E6220FD63D27A">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
</mods:titleInfo>
<mods:name id="FF071C41E2715311ADF95891E07EAB09" type="personal">
<mods:role id="4D932545C036CEB6E5B7F09187AB07B8">
<mods:roleTerm id="FA5F79E219065F84CA53B0607077D668">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4F402EAAC7FADE38319A787B6E63FDF3">Frolov, Andrey V.</mods:namePart>
</mods:name>
<mods:name id="D3929CE433CD348551D5223C0BEFC1EE" type="personal">
<mods:role id="A053980AB988B93ADA959E236C69BD39">
<mods:roleTerm id="96E125FC4B9987CA29A5E74F29A1A49A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="AF81A295B765FDFCD056BE29B59D31F1">Montreuil, Olivier</mods:namePart>
</mods:name>
<mods:name id="28E9E4FA0AE0E813537C9B168FE1AC15" type="personal">
<mods:role id="9630BC2181E8D4FB48A3BD6130404478">
<mods:roleTerm id="44E47F5CD967E570F35DD5D42342B127">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="392E7EFEE47E2A224DB18F95768FFECF">Akhmetova, Lilia A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0A3DD853C63A4E3BF6F9253A3E0093C9">text</mods:typeOfResource>
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<mods:titleInfo id="5AACD9111FCEE96B91C9B476F3D215DD">
<mods:title id="31D10E358B38AB3E46FC68C62312016F">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="CE3A85F6321BA1FDFF515FEB37B3FB3D">
<mods:date id="5F59CF7AAB99A1317A1F7773745DA78A">2016</mods:date>
<mods:detail id="0BF234448DA16881F80BBE766FF6F185" type="volume">
<mods:number id="4C9FE697AAA1124EBF8C64A5849173DE">4207</mods:number>
</mods:detail>
<mods:detail id="B6DAEA4D333E90D2659A476EDCE317B1" type="issue">
<mods:number id="51345BE51C7934B3BC400FAE230D41EA">1</mods:number>
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<mods:extent id="86C9C2B0AF69FA21EF9711DD923C388C" unit="page">
<mods:start id="0F23162C9AF8A20A2EE2029E68965B6E">1</mods:start>
<mods:end id="2EE20BA1DA1DA2147B829717B9BD7B4A">93</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="A99B2921F89BBCEDB12558FA4B60433C">journal article</mods:classification>
<mods:identifier id="390C4A7CF66F26934B87DF72069B6CE0" type="CLB-Dataset">37549</mods:identifier>
<mods:identifier id="ECD52BFB3CC7A88CEC4655D439FAB677" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
<mods:identifier id="8EB5566F398ABAC15D98A9305D8EED84" type="GBIF-Dataset">cef35771-ff95-4a41-9af6-196dea899a64</mods:identifier>
<mods:identifier id="37303F494F456738B0D33BD643407453" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="212BEE9F32F0B8D763C4E6950FE5912C" type="Zenodo-Dep">200444</mods:identifier>
<mods:identifier id="3CBBE354C63FFAB77D88D55CCAF8EB0E" type="ZooBank">F2875582-31E2-496F-AEEF-1D657DD86C33</mods:identifier>
</mods:mods>
<treatment id="5B4FBD00FF9A6127FF02655EFDD4FAF8" ID-DOI="http://doi.org/10.5281/zenodo.6057697" ID-GBIF-Taxon="125107297" ID-Zenodo-Dep="6057697" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9A6127FF02655EFDD4FAF8" httpUri="http://treatment.plazi.org/id/5B4FBD00FF9A6127FF02655EFDD4FAF8" lastPageId="77" lastPageNumber="78" pageId="75" pageNumber="76">
<subSubSection id="9BFC5F9DFF9A6121FF02655EFEB0F8C8" pageId="75" pageNumber="76" type="nomenclature">
<paragraph id="D3590C16FF9A6121FF02655EFC2CF8E9" blockId="75.[151,934,1794,1853]" box="[151,934,1794,1820]" pageId="75" pageNumber="76">
<heading id="8811BB7AFF9A6121FF02655EFC2CF8E9" bold="true" box="[151,934,1794,1820]" fontSize="11" level="1" pageId="75" pageNumber="76" reason="1">
<emphasis id="E192D004FF9A6121FF02655EFC2CF8E9" bold="true" box="[151,934,1794,1820]" pageId="75" pageNumber="76">
<taxonomicName id="14E67795FF9A6121FF02655EFC83F8E9" authority="Frolov, Montreuil &amp; Akhmetova" authorityName="Frolov, Montreuil &amp; Akhmetova" box="[151,777,1794,1820]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="viettei" status="sp. nov.">
<emphasis id="E192D004FF9A6121FF02655EFEE1F8E9" bold="true" box="[151,363,1794,1820]" italics="true" pageId="75" pageNumber="76">Triodontus viettei</emphasis>
Frolov, Montreuil &amp; Akhmetova
</taxonomicName>
,
<taxonomicNameLabel id="FAA16D7FFF9A6121FC83655EFC2CF8E9" box="[790,934,1794,1820]" pageId="75" pageNumber="76" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="D3590C16FF9A6121FF026578FEB0F8C8" blockId="75.[151,934,1794,1853]" box="[151,314,1828,1853]" pageId="75" pageNumber="76">
(
<figureCitation id="4BDD1093FF9A6121FF0A6578FF70F8C8" box="[159,250,1828,1853]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="75" pageNumber="76">Figs. 39</figureCitation>
A–H)
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9A6126FF026530FD1AFF21" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="materials_examined">
<paragraph id="D3590C16FF9A6121FF026530FB89F85C" blockId="75.[151,1436,1900,2033]" pageId="75" pageNumber="76">
<materialsCitation id="638E064BFF9A6121FF026530FC75F85C" ID-GBIF-Occurrence="1416610106" collectionCode="MNHN" country="Madagascar" county="Perrot Freres" location="Alaotra" municipality="Trimestre" pageId="75" pageNumber="76" specimenCount="3" typeStatus="holotype">
<emphasis id="E192D004FF9A6121FF026530FECDF870" bold="true" box="[151,327,1900,1925]" pageId="75" pageNumber="76">
<typeStatus id="0C5DB2B4FF9A6121FF026530FF58F870" box="[151,210,1900,1925]" pageId="75" pageNumber="76">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0C5DB2B4FF9A6121FEC56530FE32F870" box="[336,440,1900,1925]" pageId="75" pageNumber="76" type="holotype">Holotype</typeStatus>
(
<figureCitation id="4BDD1093FF9A6121FE5F6530FDA2F870" box="[458,552,1900,1925]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="75" pageNumber="76">Figs. 39</figureCitation>
A–C, E–G), male, “
<collectingCountry id="ABF14C86FF9A6121FC9E6530FC1FF870" box="[779,917,1900,1925]" name="Madagascar" pageId="75" pageNumber="76">Madagascar</collectingCountry>
Antsianaka et lac
<location id="D6395ACDFF9A6121FBFB6530FB48F870" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9A6127FF02655EFDD4FAF8:D6395ACDFF9A6121FBFB6530FB48F870" box="[1134,1218,1900,1925]" country="Madagascar" county="Perrot Freres" municipality="Trimestre" name="Alaotra" pageId="75" pageNumber="76">Alaotra</location>
2e
<collectingMunicipality id="333D966CFF9A6121FB7A6530FAD1F870" box="[1263,1371,1900,1925]" pageId="75" pageNumber="76">Trimestre</collectingMunicipality>
1889
<collectingCounty id="3A38749AFF9A6121FF0265CDFEA2F85D" box="[151,296,1937,1960]" pageId="75" pageNumber="76">Perrot Freres</collectingCounty>
/
<typeStatus id="0C5DB2B4FF9A6121FEAB65CDFE6EF85C" box="[318,484,1937,1961]" pageId="75" pageNumber="76" type="holotype">HOLOTYPUS</typeStatus>
<taxonomicName id="14E67795FF9A6121FE7E65CCFC08F85C" authority="Frolov et al. 2010" authorityName="Frolov et al." authorityYear="2010" box="[491,898,1936,1961]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="E192D004FF9A6121FE7E65CCFD3AF85C" box="[491,688,1936,1961]" italics="true" pageId="75" pageNumber="76">Triodontus viettei</emphasis>
Frolov
<emphasis id="E192D004FF9A6121FC9C65CEFCCBF85C" box="[777,833,1936,1961]" italics="true" pageId="75" pageNumber="76">et al.</emphasis>
2010
</taxonomicName>
” (
<collectionCode id="B5F794D3FF9A6121FC0E65CDFC7DF85C" box="[923,1015,1937,1961]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="75" pageNumber="76">MNHN</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="D3590C16FF9A6126FF5265E9FD1AFF21" blockId="75.[151,1436,1900,2033]" lastBlockId="76.[151,1436,151,213]" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
Paratypes: one male, “Tananarive Lamberton” (MNHN);
<materialsCitation id="638E064BFF9A6126FCF865E9FCF6FF45" ID-GBIF-Occurrence="1416610148" collectingDate="1963-12-09" collectionCode="MNHN" collectorName="du Tsaratanana &amp; Haut Sambirano &amp; de la Besanetribe &amp; P. Viette" country="Madagascar" elevation="1200" lastPageId="76" lastPageNumber="77" location="Upper Sambirano River" municipality="Besanetribe Valley" pageId="75" pageNumber="76" specimenCount="1">
one male and two females, “
<collectingCountry id="ABF14C86FF9A6121FB5065E8FAC5F838" box="[1221,1359,1972,1997]" name="Madagascar" pageId="75" pageNumber="76">Madagascar</collectingCountry>
Nord, contreforts
<collectorName id="7E1369C0FF9A6121FE8C6584FE4DF805" box="[281,455,2008,2033]" pageId="75" pageNumber="76">du Tsaratanana</collectorName>
,
<collectorName id="7E1369C0FF9A6121FE416585FD05F804" box="[468,655,2008,2033]" pageId="75" pageNumber="76">Haut Sambirano</collectorName>
,
<elevation id="58CBEB25FF9A6121FD086585FD7CF805" box="[669,758,2009,2033]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="75" pageNumber="76" unit="m" value="1200.0">
<quantity id="141EA1F3FF9A6121FD086585FD7CF805" box="[669,758,2009,2033]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="75" pageNumber="76" unit="m" value="1200.0">1200 m</quantity>
</elevation>
, vallee
<collectorName id="7E1369C0FF9A6121FCDA6584FB9EF804" box="[847,1044,2008,2033]" pageId="75" pageNumber="76">de la Besanetribe</collectorName>
[
<location id="D6395ACDFF9A6121FBB16585FAB2F804" LSID="urn:lsid:plazi:treatment:5B4FBD00FF9A6127FF02655EFDD4FAF8:D6395ACDFF9A6121FBB16585FAB2F804" box="[1060,1336,2008,2033]" country="Madagascar" municipality="Besanetribe Valley" name="Upper Sambirano River" pageId="75" pageNumber="76">Upper Sambirano River</location>
,
<elevation id="58CBEB25FF9A6126FAD06585FF46FF45" lastPageId="76" lastPageNumber="77" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="75" pageNumber="76" unit="m" value="1200.0">
<quantity id="141EA1F3FF9A6121FAD06585FA16F805" box="[1349,1436,2009,2033]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="75" pageNumber="76" unit="m" value="1200.0">1200 m</quantity>
a.s.l.
</elevation>
,
<collectingMunicipality id="333D966CFF9D6126FF7662CBFE4EFF45" box="[227,452,151,176]" pageId="76" pageNumber="77">Besanetribe Valley</collectingMunicipality>
], 5/
<collectingDate id="B71CD33EFF9D6126FE6062CBFDFDFF45" box="[501,631,151,176]" pageId="76" pageNumber="77" value="1963-12-09">
<date id="A7582AD6FF9D6126FE6062CBFDFDFF45" box="[501,631,151,176]" pageId="76" pageNumber="77" value="1963-12-09">9-XII-1963</date>
</collectingDate>
,
<collectorName id="7E1369C0FF9D6126FD1962C4FD7FFF45" box="[652,757,151,176]" pageId="76" pageNumber="77">P. Viette</collectorName>
” (
<collectionCode id="B5F794D3FF9D6126FC8D62C4FCFEFF45" box="[792,884,152,176]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="76" pageNumber="77">MNHN</collectionCode>
)
</materialsCitation>
; two males, “MADAGASCAR Reg. Perinet [Andasibe] G. Olsoufieff col. 193” (MNHN).
</paragraph>
</subSubSection>
<caption id="87995C9EFF9D6126FF0265CEFD56F810" httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="76" pageNumber="77" targetBox="[209,1408,277,1900]" targetPageId="76">
<paragraph id="D3590C16FF9D6126FF0265CEFD56F810" blockId="76.[151,1436,1938,2021]" pageId="76" pageNumber="77">
<emphasis id="E192D004FF9D6126FF0265CEFEAFF85D" bold="true" box="[151,293,1938,1960]" pageId="76" pageNumber="77">FIGURE 39.</emphasis>
<taxonomicName id="14E67795FF9D6126FEA565CEFE62F85D" box="[304,488,1938,1960]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="76" pageNumber="77" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="E192D004FF9D6126FEA565CEFE62F85D" box="[304,488,1938,1960]" italics="true" pageId="76" pageNumber="77">Triodontus viettei</emphasis>
</taxonomicName>
, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H).
</paragraph>
</caption>
<subSubSection id="9BFC5F9DFF9C6127FF5262CBFAA6FF21" pageId="77" pageNumber="78" type="diagnosis">
<paragraph id="D3590C16FF9C6127FF5262CBFAA6FF21" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF5262CBFECBFF45" bold="true" box="[199,321,151,176]" pageId="77" pageNumber="78">Diagnosis.</emphasis>
<taxonomicName id="14E67795FF9C6127FEDD62CBFD84FF45" box="[328,526,151,176]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="E192D004FF9C6127FEDD62CBFD84FF45" box="[328,526,151,176]" italics="true" pageId="77" pageNumber="78">Triodontus viettei</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="14E67795FF9C6127FD7562C5FC04FF45" box="[736,910,151,176]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9C6127FD7562C5FC04FF45" box="[736,910,151,176]" italics="true" pageId="77" pageNumber="78">T. hildebrandtii</emphasis>
</taxonomicName>
but differs from it in having outer lobes of the parameres with slit-shaped, lateral notches, as opposed to having wide lateral notches in
<taxonomicName id="14E67795FF9C6127FBED62E1FAAFFF20" box="[1144,1317,188,213]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="hildebrandtii">
<emphasis id="E192D004FF9C6127FBED62E1FAAFFF20" box="[1144,1317,188,213]" italics="true" pageId="77" pageNumber="78">T. hildebrandtii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6127FF526283FE83FB55" pageId="77" pageNumber="78" type="description">
<paragraph id="D3590C16FF9C6127FF526283FD40FEE9" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF526283FDA2FF0D" bold="true" box="[199,552,223,248]" pageId="77" pageNumber="78">
Description.
<typeStatus id="0C5DB2B4FF9C6127FEF36283FE5FFF0D" box="[358,469,223,248]" pageId="77" pageNumber="78" type="holotype">Holotype</typeStatus>
, male.
</emphasis>
Body length
<quantity id="141EA1F3FF9C6127FD416283FCC8FF02" box="[724,834,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="1.45" pageId="77" pageNumber="78" unit="mm" value="14.5">14.5 mm</quantity>
. Color uniformly dark brown, legs, antennae, and underside of the body slightly paler (
<figureCitation id="4BDD1093FF9C6127FDA16358FD0FFEE9" box="[564,645,260,285]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="77" pageNumber="78">Fig. 39</figureCitation>
A–B).
</paragraph>
<paragraph id="D3590C16FF9C6127FF52637BFB8EFE25" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">Frontoclypeus slightly convex anteriorly, rounded laterally, anterior margin crenulate and setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small dorsal and large ventral parts. Frontoclypeus with tubercles mediad of each eye and with long horn in the center of frontoclypeus. The horn is longer than width of the head, acutely rounded apically, somewhat curved caudally, slightly rugose on posterior side.</paragraph>
<paragraph id="D3590C16FF9C6127FF526381FC26FD5D" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">Pronotum with a bulge medially, with large excavations at each side of bulge, and with 2 ridge-shaped tubercles laterally of each excavation. The median bulge is rounded and somewhat depressed on the top, with 2 distinct tubercles and a depression, receiving frontoclypeal horn. Lateral margins with wide border appearing somewhat crenulate (in dorsal view), with a row of brown setae. Anterior margin with wide, smooth border. Posterior margin with fine border; not crenulate; punctate with small, narrow, longitudinal punctures. Surface of most pronotum almost smooth, with minute, feebly visible punctation.</paragraph>
<paragraph id="D3590C16FF9C6127FF5260E8FC5CFD38" blockId="77.[151,1437,151,1293]" box="[199,982,692,717]" pageId="77" pageNumber="78">Scutellum rounded apically, visible part is about 1/13 length of elytra.</paragraph>
<paragraph id="D3590C16FF9C6127FF52608BFD59FCA9" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">Elytra convex, with distinct humeral humps. Maximum width approximately at basal 1/3. Elytra with 10 feebly visible striae on disc and with oblique line from base of 6th stria to approximately middle of 8th stria. Stria with small punctures. Intervals with sparse, minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures each bearing a small, yellow seta.</paragraph>
<paragraph id="D3590C16FF9C6127FF52613BFDFFFC51" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
Sternite 8 medially as wide as sternites 2–5 combined, without tubercle or concavity in the middle, with apical border deeply sinuate medially (
<figureCitation id="4BDD1093FF9C6127FD9561D0FDDAFC51" box="[512,592,908,933]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="77" pageNumber="78">Fig. 39</figureCitation>
G).
</paragraph>
<paragraph id="D3590C16FF9C6127FF5261ECFA04FC19" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
Parameres with outer lobes being relatively wide (in dorsal view) and having slit-shaped, lateral notches (
<figureCitation id="4BDD1093FF9C6127FAFB61F3FF3FFC19" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="77" pageNumber="78">Fig. 39</figureCitation>
E). Internal sac of the aedeagus with 1 strongly sclerotized sclerite and 2 additional similar sclerites (
<figureCitation id="4BDD1093FF9C6127FA886188FAE5FC19" box="[1309,1391,980,1005]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="77" pageNumber="78">Fig. 39</figureCitation>
F).
</paragraph>
<paragraph id="D3590C16FF9C6127FF5261ABFDC8FBAD" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF5261ABFEAFFBE5" bold="true" box="[199,293,1015,1040]" pageId="77" pageNumber="78">Female.</emphasis>
Females differ from males in having apical spur on protibiae; absence of the head and pronotum armature; densely punctate dorsal side of the head; and in longer abdominal sternites including 6th sternite, which is evenly rounded apically (
<figureCitation id="4BDD1093FF9C6127FE586663FD95FBAD" box="[461,543,1087,1112]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="77" pageNumber="78">Fig. 39</figureCitation>
D).
</paragraph>
<paragraph id="D3590C16FF9C6127FF526638FE83FB55" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF526638FEB6FB88" bold="true" box="[199,316,1124,1149]" pageId="77" pageNumber="78">Variation.</emphasis>
Body length of the examined paratypes varied from 13.5–14.0 mm (males) and from
<quantity id="141EA1F3FF9C6127FB696638FA17FB89" box="[1276,1437,1124,1148]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.19" metricValueMin="1.17" pageId="77" pageNumber="78" unit="mm" value="11.8" valueMax="11.9" valueMin="11.7">11.7–11.9 mm</quantity>
(females).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6127FF5266F0FA19FB31" box="[199,1427,1196,1221]" pageId="77" pageNumber="78" type="distribution">
<paragraph id="D3590C16FF9C6127FF5266F0FA19FB31" blockId="77.[151,1437,151,1293]" box="[199,1427,1196,1221]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF5266F0FEEAFB30" bold="true" box="[199,352,1196,1221]" pageId="77" pageNumber="78">Distribution.</emphasis>
<taxonomicName id="14E67795FF9C6127FEF266F0FDA6FB31" box="[359,556,1196,1221]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="species" species="viettei">
<emphasis id="E192D004FF9C6127FEF266F0FDA6FB31" box="[359,556,1196,1221]" italics="true" pageId="77" pageNumber="78">Triodontus viettei</emphasis>
</taxonomicName>
is known from four localities in eastern and northern
<collectingCountry id="ABF14C86FF9C6127FB1266F0FA84FB30" box="[1159,1294,1196,1221]" name="Madagascar" pageId="77" pageNumber="78">Madagascar</collectingCountry>
(
<figureCitation id="4BDD1093FF9C6127FA8B66F0FAFAFB31" box="[1310,1392,1196,1221]" captionStart="FIGURE 39" captionStartId="76.[151,250,1938,1960]" captionTargetBox="[209,1408,277,1900]" captionTargetId="figure@76.[169,1417,265,1934]" captionTargetPageId="76" captionText="FIGURE 39. Triodontus viettei, holotype male (A, B, C, E, F, G), paratype female (D). Habitus in dorsal (A, D) and dorsolateral view (B), labels of holotype (C), aedeagus in lateral view and parameres in dorsal and ventral view (E), internal sac of aedeagus (F), abdomen (G), distributional records (H)." httpUri="https://zenodo.org/record/200482/files/figure.png" pageId="77" pageNumber="78">Fig. 39</figureCitation>
H).
</paragraph>
</subSubSection>
<subSubSection id="9BFC5F9DFF9C6127FF526693FDD4FAF8" pageId="77" pageNumber="78" type="etymology">
<paragraph id="D3590C16FF9C6127FF526693FDD4FAF8" blockId="77.[151,1437,151,1293]" pageId="77" pageNumber="78">
<emphasis id="E192D004FF9C6127FF526693FEC4FB1D" bold="true" box="[199,334,1231,1256]" pageId="77" pageNumber="78">Etymology.</emphasis>
The new species is dedicated to the memory of Pierre Viette, French entomologist, specialist of
<taxonomicName id="14E67795FF9C6127FF0266A8FE20FAF8" authority="(Lepidoptera)" baseAuthorityName="Lepidoptera" box="[151,426,1268,1293]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="77" pageNumber="78" phylum="Arthropoda" rank="family">Noctuidae (Lepidoptera)</taxonomicName>
of
<collectingCountry id="ABF14C86FF9C6127FE4566A8FDD0FAF8" box="[464,602,1268,1293]" name="Madagascar" pageId="77" pageNumber="78">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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