170 lines
20 KiB
XML
170 lines
20 KiB
XML
<document id="E25EA59FE508509BEB71520FBCDAD39F" ID="10.11646/zootaxa.4207.1.1" ID-CLB-Dataset="37549" ID-DOI="10.11646/zootaxa.4207.1.1" ID-GBIF-Dataset="cef35771-ff95-4a41-9af6-196dea899a64" ID-ISSN="1175-5326" ID-Zenodo-Dep="200444" ID-ZooBank="F2875582-31E2-496F-AEEF-1D657DD86C33" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1481615616696" checkinUser="plazi" docAuthor="Frolov, Andrey V., Montreuil, Olivier & Akhmetova, Lilia A." docDate="2016" docId="5B4FBD00FF92612EFF0261D0FD9FFF20" docLanguage="en" docName="zootaxa.4207.1.1.pdf" docOrigin="Zootaxa 4207 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Triodontus ankarafantsikae Frolov, Montreuil & Akhmetova, new species" docType="treatment" docVersion="16" lastPageNumber="69" masterDocId="A776C578FFD1616AFF95625CFF8AFFF5" masterDocTitle="Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood" masterLastPageNumber="93" masterPageNumber="1" pageNumber="68" updateTime="1698433759527" updateUser="ExternalLinkService">
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<mods:titleInfo id="7FF53669AEE6328A576D137F3004253C">
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<mods:title id="8CED286B02DB4EE951381E9FAF5749AE">Review of the Madagascan Orphninae (Coleoptera: Scarabaeidae) with a revision of the genus Triodontus Westwood</mods:title>
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<mods:name id="E2D47B46BA7F584A830880752D9ACE25" type="personal">
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<mods:namePart id="DCC8960FC159A2D785807BD4D3AC0093">Frolov, Andrey V.</mods:namePart>
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<mods:namePart id="FC4F2A7D6C926AFD52BF5F41B13F7D09">Montreuil, Olivier</mods:namePart>
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<mods:name id="2DC58C43ED52321101ED4578429565C8" type="personal">
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<mods:namePart id="BF40F012525F6284160E8A3A20FB31BF">Akhmetova, Lilia A.</mods:namePart>
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<mods:typeOfResource id="699D7D4830916F223ACC9592963BC66D">text</mods:typeOfResource>
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<mods:title id="7C6B7F66AA216D7CC95F30154AF7C5E5">Zootaxa</mods:title>
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<mods:part id="C57B7246F7F0472981618CA95C971767">
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<mods:date id="13CA9AFB0DA1204C496E1FBE9948E7FF">2016</mods:date>
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<mods:number id="5964261694C465BD18517F8E649595A3">4207</mods:number>
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<mods:classification id="F7F7B477B9F156D3B74109BDEDA35CCB">journal article</mods:classification>
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<mods:identifier id="CE0B1E660935862FA30DBA2CD0F839B6" type="CLB-Dataset">37549</mods:identifier>
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<mods:identifier id="A0031CD7D51FD26263CB62D68293C74A" type="DOI">10.11646/zootaxa.4207.1.1</mods:identifier>
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<mods:identifier id="12157CD374D15DFAD7EFC03D995BE974" type="ISSN">1175-5326</mods:identifier>
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<treatment id="5B4FBD00FF92612EFF0261D0FD9FFF20" ID-DOI="http://doi.org/10.5281/zenodo.6057688" ID-GBIF-Taxon="125107301" ID-Zenodo-Dep="6057688" LSID="urn:lsid:plazi:treatment:5B4FBD00FF92612EFF0261D0FD9FFF20" httpUri="http://treatment.plazi.org/id/5B4FBD00FF92612EFF0261D0FD9FFF20" lastPageId="68" lastPageNumber="69" pageId="67" pageNumber="68">
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<subSubSection id="9BFC5F9DFF926129FF0261D0FEB0FC32" pageId="67" pageNumber="68" type="nomenclature">
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<paragraph id="D3590C16FF926129FF0261D0FBA9FC53" blockId="67.[151,1059,908,967]" box="[151,1059,908,934]" pageId="67" pageNumber="68">
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<heading id="8811BB7AFF926129FF0261D0FBA9FC53" bold="true" box="[151,1059,908,934]" fontSize="11" level="1" pageId="67" pageNumber="68" reason="1">
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<emphasis id="E192D004FF926129FF0261D0FBA9FC53" bold="true" box="[151,1059,908,934]" pageId="67" pageNumber="68">
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<taxonomicName id="14E67795FF926129FF0261D0FC0DFC53" authority="Frolov, Montreuil & Akhmetova" authorityName="Frolov, Montreuil & Akhmetova" box="[151,903,908,934]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="ankarafantsikae" status="sp. nov.">
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<emphasis id="E192D004FF926129FF0261D0FE62FC53" bold="true" box="[151,488,908,934]" italics="true" pageId="67" pageNumber="68">Triodontus ankarafantsikae</emphasis>
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Frolov, Montreuil & Akhmetova
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</taxonomicName>
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,
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<taxonomicNameLabel id="FAA16D7FFF926129FC0661D0FBA9FC53" box="[915,1059,908,934]" pageId="67" pageNumber="68" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D3590C16FF926129FF0261F2FEB0FC32" blockId="67.[151,1059,908,967]" box="[151,314,942,967]" pageId="67" pageNumber="68">
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(
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<figureCitation id="4BDD1093FF926129FF0A61F2FF70FC32" box="[159,250,942,967]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Figs. 34</figureCitation>
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A–G)
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFF926129FF0261AAFAE9FBC6" pageId="67" pageNumber="68" type="materials_examined">
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<paragraph id="D3590C16FF926129FF0261AAFAE9FBC6" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">
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<materialsCitation id="638E064BFF926129FF0261AAFAD5FBC6" ID-GBIF-Occurrence="1416610097" collectionCode="MUSEUM, PARIS, OUEST, MNHN" country="Madagascar" location="MADAGASCAR" pageId="67" pageNumber="68" specimenCount="3" typeStatus="holotype">
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<emphasis id="E192D004FF926129FF0261AAFECDFBFA" bold="true" box="[151,327,1014,1039]" pageId="67" pageNumber="68">
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<typeStatus id="0C5DB2B4FF926129FF0261AAFF58FBFA" box="[151,210,1014,1039]" pageId="67" pageNumber="68">Type</typeStatus>
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material.
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</emphasis>
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<typeStatus id="0C5DB2B4FF926129FEC561AAFE32FBFA" box="[336,440,1014,1039]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="4BDD1093FF926129FE5F61AAFDA3FBFA" box="[458,553,1014,1039]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Figs. 34</figureCitation>
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A–F), male, “ANKARAFANTSIKA AMPIJOROA 1.10-11-1973/
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<collectionCode id="B5F794D3FF926129FAB461ABFF6EFBC6" pageId="67" pageNumber="68">MUSEUM PARIS</collectionCode>
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<collectingCountry id="ABF14C86FF926129FF7E6647FE38FBC6" box="[235,434,1051,1075]" name="Madagascar" pageId="67" pageNumber="68">MADAGASCAR</collectingCountry>
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<collectionCode id="B5F794D3FF926129FE2C6647FD99FBC6" box="[441,531,1051,1075]" pageId="67" pageNumber="68">OUEST</collectionCode>
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/
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<taxonomicName id="14E67795FF926129FDBC6646FCD4FBC6" box="[553,862,1050,1075]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="ankarafantsikae">
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<emphasis id="E192D004FF926129FDBC6646FCD4FBC6" box="[553,862,1050,1075]" italics="true" pageId="67" pageNumber="68">Triodontus ankarafantsikae</emphasis>
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</taxonomicName>
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<typeStatus id="0C5DB2B4FF926129FCF16647FB80FBC6" box="[868,1034,1051,1075]" pageId="67" pageNumber="68" type="holotype">HOLOTYPUS</typeStatus>
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Frolov et. al. 2010” (
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<collectionCode id="B5F794D3FF926129FB6E6647FADDFBC6" box="[1275,1367,1051,1075]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="67" pageNumber="68">MNHN</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFF926129FF526662FE77FB36" pageId="67" pageNumber="68" type="diagnosis">
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<paragraph id="D3590C16FF926129FF526662FE77FB36" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">
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<emphasis id="E192D004FF926129FF526662FECBFBA2" bold="true" box="[199,321,1086,1111]" pageId="67" pageNumber="68">Diagnosis.</emphasis>
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<taxonomicName id="14E67795FF926129FEDD6662FDF4FBA2" box="[328,638,1086,1111]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="ankarafantsikae">
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<emphasis id="E192D004FF926129FEDD6662FDF4FBA2" box="[328,638,1086,1111]" italics="true" pageId="67" pageNumber="68">Triodontus ankarafantsikae</emphasis>
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</taxonomicName>
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is similar to
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<taxonomicName id="14E67795FF926129FC856663FC02FBA2" box="[784,904,1086,1111]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lemoulti" status="sp. nov.">
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<emphasis id="E192D004FF926129FC856663FC02FBA2" box="[784,904,1086,1111]" italics="true" pageId="67" pageNumber="68">T. lemoulti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="FAA16D7FFF926129FC1A6663FB9CFBA2" box="[911,1046,1086,1111]" pageId="67" pageNumber="68" rank="species">new species</taxonomicNameLabel>
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in having the abdominal sternite 8 with a more or less developed tubercule medially, but differs from it in having the tubercule rounded apically (as opposed to being concave in
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<taxonomicName id="14E67795FF926129FE7266DBFDEAFB6A" box="[487,608,1158,1183]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="67" pageNumber="68" phylum="Arthropoda" rank="species" species="lemoulti">
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<emphasis id="E192D004FF926129FE7266DBFDEAFB6A" box="[487,608,1158,1183]" italics="true" pageId="67" pageNumber="68">T. lemoulti</emphasis>
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</taxonomicName>
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) and lateral lobes of parameres with keel-shaped processes on proximal parts (in dorsal view,
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<figureCitation id="4BDD1093FF926129FE1D66F6FE50FB36" box="[392,474,1194,1219]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Fig. 34</figureCitation>
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D).
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</paragraph>
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</subSubSection>
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<subSubSection id="9BFC5F9DFF926129FF526692FE28F866" pageId="67" pageNumber="68" type="description">
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<paragraph id="D3590C16FF926129FF526692FBFAFAFE" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">
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<emphasis id="E192D004FF926129FF526692FDAEFB12" bold="true" box="[199,548,1230,1255]" pageId="67" pageNumber="68">
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Description.
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<typeStatus id="0C5DB2B4FF926129FEF16692FE59FB12" box="[356,467,1230,1255]" pageId="67" pageNumber="68" type="holotype">Holotype</typeStatus>
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, male.
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</emphasis>
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Body length
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<quantity id="141EA1F3FF926129FD5D6692FCBEFB13" box="[712,820,1230,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="67" pageNumber="68" unit="mm" value="10.6">10.6 mm</quantity>
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. Body strongly shiny (
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<figureCitation id="4BDD1093FF926129FBD76692FB29FB12" box="[1090,1187,1230,1255]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Figs. 34</figureCitation>
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A–B). Color of head, pronotum and elytra dark brown; legs, antennae, and underside of the body light brown.
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</paragraph>
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<paragraph id="D3590C16FF926129FF52674AFB8EFA4A" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">Frontoclypeus convex anteriorly, rounded laterally, anterior margin slightly crenulate and setose in dorsal view. Eyes relatively small (diameter slightly smaller than the distance between eye and gula in ventral view), incompletely divided by canthus into small dorsal and large ventral parts. Frontoclypeus with small tubercles mediad of each eye and with a long horn in center of frontoclypeus; horn is longer than width of head, acutely rounded apically, somewhat curved caudally, slightly rugose on posterior side.</paragraph>
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<paragraph id="D3590C16FF926129FF526797FBE4F9DF" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">Pronotum convex, almost smooth, feebly excavated anteriorly, with a medial bulge divided into 2 separated tubercles and 2 tubercles laterad of the bulge. Anterior margin with wide, smooth border, feebly sinuate medially. Posterior margin with fine border, not crenulate, punctate with small, narrow punctures.</paragraph>
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<paragraph id="D3590C16FF926129FF52646AFC57F9BA" blockId="67.[151,1437,1014,2011]" box="[199,989,1590,1615]" pageId="67" pageNumber="68">Scutellum rounded apically, its visible part about 1/15 length of elytra.</paragraph>
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<paragraph id="D3590C16FF926129FF526406FD35F92B" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">Elytra convex, with distinct humeral humps. Maximum width approximately at the middle. Elytra with 10 feebly visible striae and with oblique line from base of 6th stria to approximately middle of 8th stria. Stria with small, indistinct punctures. Intervals with minute punctures, almost smooth. Base of elytra with an irregular row of coarse punctures each bearing small, yellow seta.</paragraph>
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<paragraph id="D3590C16FF926129FF5264B6FAA8F8F6" blockId="67.[151,1437,1014,2011]" box="[199,1314,1770,1795]" pageId="67" pageNumber="68">
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Abdominal sternite 8 with a developed tubercule medially, tubercule is rounded apically (
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<figureCitation id="4BDD1093FF926129FB2764B6FA8EF8F6" box="[1202,1284,1770,1795]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Fig. 34</figureCitation>
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F).
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</paragraph>
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<paragraph id="D3590C16FF926129FF526553FD0EF89A" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">
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Parameres have outer lobes with keel-shaped processes on proximal parts (in dorsal view,
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<figureCitation id="4BDD1093FF926129FB516552FA9CF8D2" box="[1220,1302,1806,1831]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Fig. 34</figureCitation>
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D) and long feebly sclerotized apices. Internal sac of the aedeagus with 1 strongly sclerotized somewhat pentagonal sclerite, and 2 additional similar sclerites (
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<figureCitation id="4BDD1093FF926129FD87650AFDEEF89A" box="[530,612,1878,1903]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Fig. 34</figureCitation>
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E).
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</paragraph>
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<paragraph id="D3590C16FF926129FF526526FE28F866" blockId="67.[151,1437,1014,2011]" box="[199,418,1914,1939]" pageId="67" pageNumber="68">
|
||
<emphasis id="E192D004FF926129FF526526FEAFF866" bold="true" box="[199,293,1914,1939]" pageId="67" pageNumber="68">Female.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFF926129FF5265C2FE9FF82E" pageId="67" pageNumber="68" type="distribution">
|
||
<paragraph id="D3590C16FF926129FF5265C2FE9FF82E" blockId="67.[151,1437,1014,2011]" pageId="67" pageNumber="68">
|
||
<emphasis id="E192D004FF926129FF5265C2FEEAF842" bold="true" box="[199,352,1950,1975]" pageId="67" pageNumber="68">Distribution.</emphasis>
|
||
The species is known from one locality in western
|
||
<collectingCountry id="ABF14C86FF926129FC2B65C2FBCDF842" box="[958,1095,1950,1975]" name="Madagascar" pageId="67" pageNumber="68">Madagascar</collectingCountry>
|
||
, from dry tropical forest area (
|
||
<figureCitation id="4BDD1093FF926129FF0A659EFF78F82E" box="[159,242,1986,2011]" captionStart="FIGURE 34" captionStartId="68.[151,250,1902,1924]" captionTargetBox="[208,1391,268,1880]" captionTargetId="figure@68.[195,1392,267,1881]" captionTargetPageId="68" captionText="FIGURE 34. Triodontus ankarafantsikae, holotype male. Habitus in dorsal (A) and dorsolateral view (B), labels (C), aedeagus in lateral view and parameres in dorsal and ventral view (D), internal sac of aedeagus (E), abdomen (F), distributional records (G)." httpUri="https://zenodo.org/record/200477/files/figure.png" pageId="67" pageNumber="68">Fig. 34</figureCitation>
|
||
G).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9BFC5F9DFF95612EFF5262CBFD9FFF20" pageId="68" pageNumber="69" type="etymology">
|
||
<paragraph id="D3590C16FF95612EFF5262CBFD9FFF20" blockId="68.[151,1436,151,213]" pageId="68" pageNumber="69">
|
||
<emphasis id="E192D004FF95612EFF5262CBFEC4FF45" bold="true" box="[199,334,151,176]" pageId="68" pageNumber="69">Etymology.</emphasis>
|
||
The species name is derived from the name of the
|
||
<typeStatus id="0C5DB2B4FF95612EFC1362C4FC3CFF45" box="[902,950,152,176]" pageId="68" pageNumber="69">type</typeStatus>
|
||
locality, Ankarafantsika. This name should be treated as a noun in apposition.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |