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<document id="6F509F04E8DE2A29202226EB41A765C5" ID-DOI="10.11646/zootaxa.5547.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14387105" ID-ZooBank="2089C3EC-8FBB-43E6-A6C5-9E6B6AD512D6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733920242413" checkinUser="plazi" docAuthor="Do Prado, André W., Baptista, Renner Luiz Cerqueira &amp; Framenau, Volker W." docDate="2024" docId="5B2489790D2AB7678ED3FACEFD17FE88" docLanguage="en" docName="zootaxa.5547.1.1.pdf" docOrigin="Zootaxa 5547 (1)" docSource="https://doi.org/10.11646/zootaxa.5547.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Artoria emu Prado, Baptista &amp; Framenau, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="25" masterDocId="A71DF1010D3FB77F8E44FFBBFFCBFF95" masterDocTitle="Taxonomy of the wolf spider genus Artoria in Western Australia (Araneae, Lycosidae, Artoriinae)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="22" updateTime="1733921097182" updateUser="ExternalLinkService">
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<mods:title id="C91E95FCB4A57E15C21A975B2B5E34A1">Taxonomy of the wolf spider genus Artoria in Western Australia (Araneae, Lycosidae, Artoriinae)</mods:title>
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<mods:namePart id="BB5CE2745EC2178CBF6B021C181E9547">Do Prado, André W.</mods:namePart>
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<mods:affiliation id="579B13945382C6CE8E5FC3E4C2DE7A7F">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia &amp; andre. wanderleydoprado @ murdoch. edu. au; https: // orcid. org / 0000 - 0002 - 3536 - 5654</mods:affiliation>
<mods:nameIdentifier id="6D3C9C9504606E43C5D98D809D9E023C" type="email">andre.wanderleydoprado@murdoch.edu.au</mods:nameIdentifier>
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<mods:namePart id="7F29933A557C1C14439C7F7D31DDFE84">Baptista, Renner Luiz Cerqueira</mods:namePart>
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<mods:affiliation id="F496704537B3CF5E59BE8959A848EE40">Departamento de Zoologia, Instituto de Biologia, Universidade do Brasil / Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil &amp; baptistr @ gmail. com; https: // orcid. org / 0000 - 0002 - 2372 - 5034</mods:affiliation>
<mods:nameIdentifier id="6F8CA52B888D5727C53FE97A0592509D" type="email">baptistr@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5C893F35DADB072EFC5187CCF4D06030">Framenau, Volker W.</mods:namePart>
<mods:nameIdentifier id="04A954AF25B2F1976014CAA51C3CB8C1" type="ORCID">0000-0002-7724-3831</mods:nameIdentifier>
<mods:affiliation id="26A9875ACC69AEE288BB47130D10D55E">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Invertebrates, Museum of Nature Hamburg - Zoology, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; volker. framenau @ murdoch. edu. au; https: // orcid. org / 0000 - 0002 - 7724 - 3831 * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="8B66EA59486597231AA725C78D2C4FDA" type="email">volker.framenau@murdoch.edu.au</mods:nameIdentifier>
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<mods:date id="938B374676F774FA6265DF9BD57BB899">2024</mods:date>
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<mods:number id="ADC691FC6BF469FE31115D16DFF2DE42">2024-12-09</mods:number>
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<mods:number id="22238435B573BB3019198C2229C0A260">5547</mods:number>
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<treatment id="5B2489790D2AB7678ED3FACEFD17FE88" ID-DOI="http://doi.org/10.5281/zenodo.14387274" ID-Zenodo-Dep="14387274" LSID="urn:lsid:plazi:treatment:5B2489790D2AB7678ED3FACEFD17FE88" httpUri="http://treatment.plazi.org/id/5B2489790D2AB7678ED3FACEFD17FE88" lastPageId="24" lastPageNumber="25" pageId="21" pageNumber="22">
<subSubSection id="9B976BE40D2AB76A8ED3FACEFE5BFA1A" box="[151,400,1397,1424]" pageId="21" pageNumber="22" type="nomenclature">
<paragraph id="D332386F0D2AB76A8ED3FACEFE5BFA1A" blockId="21.[151,971,1397,1496]" box="[151,400,1397,1424]" pageId="21" pageNumber="22">
<heading id="887A8F030D2AB76A8ED3FACEFE5BFA1A" bold="true" box="[151,400,1397,1424]" fontSize="11" level="2" pageId="21" pageNumber="22" reason="4">
<emphasis id="E1F9E47D0D2AB76A8ED3FACEFE5BFA1A" bold="true" box="[151,400,1397,1424]" pageId="21" pageNumber="22">
<taxonomicName id="148D43EC0D2AB76A8ED3FACEFEE0FA05" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[151,299,1397,1424]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D2AB76A8ED3FACEFEE0FA05" bold="true" box="[151,299,1397,1424]" italics="true" pageId="21" pageNumber="22">Artoria emu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FACA59060D2AB76A8F76FACEFE5BFA1A" box="[306,400,1397,1423]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="9B976BE40D2AB76A8ED3FA22FC00FA4D" pageId="21" pageNumber="22" type="description">
<paragraph id="D332386F0D2AB76A8ED3FA22FECDFA26" blockId="21.[151,971,1397,1496]" box="[151,262,1433,1459]" pageId="21" pageNumber="22">
<figureCitation id="4BB624EA0D2AB76A8ED3FA22FECDFA26" box="[151,262,1433,1459]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="22.[151,250,1704,1728]" captionStartId-1="23.[151,250,1780,1804]" captionTargetBox-0="[151,1425,190,1678]" captionTargetBox-1="[157,1430,746,1747]" captionTargetId-0="figure-20@22.[151,1436,179,1678]" captionTargetId-1="figure-265@23.[151,1436,738,1754]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." captionText-1="FIGURE 10. Distribution of Artoria emu sp. nov., Artoria falcata sp. nov., and Artoria flavimana Simon, 1909." figureDoi-0="http://doi.org/10.5281/zenodo.14387135" figureDoi-1="http://doi.org/10.5281/zenodo.14387139" httpUri-0="https://zenodo.org/record/14387135/files/figure.png" httpUri-1="https://zenodo.org/record/14387139/files/figure.png" pageId="21" pageNumber="22">Figs 910</figureCitation>
</paragraph>
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<uri id="A71C346D0D2AB76A8ED3FA06FC00FA4D" box="[151,971,1469,1496]" pageId="21" pageNumber="22">
urn:lsid:zoobank.org:act:
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</uri>
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D2AB76A8ED3FA44FCF7F910" pageId="21" pageNumber="22" type="materials_examined">
<paragraph id="D332386F0D2AB76A8ED3FA44FF34F9F4" blockId="21.[151,1437,1534,2029]" pageId="21" pageNumber="22">
<emphasis id="E1F9E47D0D2AB76A8ED3FA44FE89F98D" bold="true" box="[151,322,1534,1561]" pageId="21" pageNumber="22">Type material.</emphasis>
<materialsCitation id="63E532320D2AB76A8F02FA44FF34F9F4" collectingDate="2003-08-29" collectingDateMax="2004-10-16" collectingDateMin="2003-08-29" collectionCode="WAM" country="Australia" latitude="-23.266111" location="Pilbara Survey" longLatPrecision="21" longitude="119.13389" pageId="21" pageNumber="22" specimenCode="T98120" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<typeStatus id="0C3686CD0D2AB76A8F02FA44FE65F98C" box="[326,430,1535,1561]" pageId="21" pageNumber="22" type="holotype">Holotype</typeStatus>
male,
<locationDeviation id="C280214D0D2AB76A8FB1FA44FD24F98C" box="[501,751,1535,1561]" location="Giles Point" pageId="21" pageNumber="22">
<quantity id="1475958A0D2AB76A8FB1FA44FD8BF98C" box="[501,576,1535,1561]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="21" pageNumber="22" unit="km" value="3.2">3.2 km</quantity>
SW Giles Point
</locationDeviation>
,
<collectingRegion id="1149F68D0D2AB76A8CB3FA44FC74F98C" box="[759,959,1535,1561]" country="Australia" name="Western Australia" pageId="21" pageNumber="22">Western Australia</collectingRegion>
,
<collectingCountry id="AB9A78FF0D2AB76A8D81FA44FBACF98C" box="[965,1127,1535,1561]" name="Australia" pageId="21" pageNumber="22">AUSTRALIA</collectingCountry>
,
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,
<geoCoordinate id="B6B95EA80D2AB76A8B45FA44FA5CF98C" box="[1281,1431,1535,1561]" degrees="119" direction="east" minutes="08" orientation="longitude" pageId="21" pageNumber="22" precision="15" seconds="02" value="119.13389">119º0802E</geoCoordinate>
,
<date id="A7331EAF0D2AB76A8ED3F998FE9EF9A8" box="[151,341,1571,1597]" pageId="21" pageNumber="22" value="2003-08-29">
<collectingDate id="B777E7470D2AB76A8ED3F998FE9EF9A8" box="[151,341,1571,1597]" pageId="21" pageNumber="22" value="2003-08-29">29 August 2003</collectingDate>
</date>
<date id="A7331EAF0D2AB76A8F20F998FDE7F9A8" box="[356,556,1571,1597]" pageId="21" pageNumber="22" value="2004-10-16">
<collectingDate id="B777E7470D2AB76A8F20F998FDE7F9A8" box="[356,556,1571,1597]" pageId="21" pageNumber="22" value="2004-10-16">16 October 2004</collectingDate>
</date>
,
<collectingMethod id="0ACC40780D2AB76A8C79F998FCB2F9A8" box="[573,889,1571,1597]" pageId="21" pageNumber="22">ethylene glycol pitfall trap</collectingMethod>
, CALM
<location id="D6526EB40D2AB76A8DA3F998FB58F9A8" LSID="urn:lsid:plazi:treatment:5B2489790D2AB7678ED3FACEFD17FE88:D6526EB40D2AB76A8DA3F998FB58F9A8" box="[999,1171,1571,1597]" country="Australia" latitude="-23.266111" longLatPrecision="21" longitude="119.13389" name="Pilbara Survey" pageId="21" pageNumber="22" stateProvince="Western Australia">Pilbara Survey</location>
, site RHNC03 (
<collectionCode id="B59CA0AA0D2AB76A8B12F998FA56F9A8" box="[1366,1437,1571,1597]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="21" pageNumber="22" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D2AB76A8ED3F9FCFF3FF9F4" box="[151,244,1607,1633]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="21" pageNumber="22" type="Museum">T98120</specimenCode>
).
</materialsCitation>
</paragraph>
<paragraph id="D332386F0D2AB76A8E83F9D1FCF7F910" blockId="21.[151,1437,1534,2029]" box="[199,828,1642,1669]" pageId="21" pageNumber="22">
<emphasis id="E1F9E47D0D2AB76A8E83F9D1FE37F911" bold="true" box="[199,508,1642,1668]" pageId="21" pageNumber="22">
<typeStatus id="0C3686CD0D2AB76A8E83F9D1FE3DF911" box="[199,502,1642,1668]" isOtherTypeStatus="true" pageId="21" pageNumber="22">Other material examined</typeStatus>
.
</emphasis>
Only known from
<typeStatus id="0C3686CD0D2AB76A8C96F9D0FCF3F910" box="[722,824,1643,1669]" pageId="21" pageNumber="22" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D2AB76A8E83F934FE22F964" pageId="21" pageNumber="22" type="etymology">
<paragraph id="D332386F0D2AB76A8E83F934FE22F964" blockId="21.[151,1437,1534,2029]" pageId="21" pageNumber="22">
<emphasis id="E1F9E47D0D2AB76A8E83F934FE85F93C" bold="true" box="[199,334,1679,1705]" pageId="21" pageNumber="22">Etymology.</emphasis>
The specific epithet
<taxonomicName id="148D43EC0D2AB76A8C71F934FDB8F93C" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[565,627,1679,1705]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="emu">“emu</taxonomicName>
” is a noun in apposition and refers to the large, flightless Australian bird,
<taxonomicName id="148D43EC0D2AB76A8ED3F908FD4FF958" authority="(Latham, 1790)" baseAuthorityName="Latham" baseAuthorityYear="1790" box="[151,644,1715,1741]" class="Aves" family="Dromaiidae" genus="Dromaius" kingdom="Animalia" order="Casuariiformes" pageId="21" pageNumber="22" phylum="Chordata" rank="species" species="novaehollandiae">
<emphasis id="E1F9E47D0D2AB76A8ED3F908FE01F958" box="[151,458,1715,1741]" italics="true" pageId="21" pageNumber="22">Dromaius novaehollandiae</emphasis>
(
<bibRefCitation id="B71C459E0D2AB76A8F9EF908FDB7F958" author="Latham, J." box="[474,636,1715,1741]" pageId="21" pageNumber="22" refId="ref58672" refString="Latham, J. (1790) Index Ornithologicus, Sive Systema Ornithologiae: Complectens Avium Divisionem in Classes, Ordines, Genera, Species, Ipsarumque Varietates. Vol. 2. Leigh &amp; Sotheby, London, 665 pp. [in Latin] https: // doi. org / 10.5962 / bhl. title. 131313" type="book" year="1790">Latham, 1790</bibRefCitation>
)
</taxonomicName>
, and the resemblance of the tegular apophysis of the males in ventral view to the profile of its head.
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D2AB76A8E83F941FC47F878" pageId="21" pageNumber="22" type="diagnosis">
<paragraph id="D332386F0D2AB76A8E83F941FC47F878" blockId="21.[151,1437,1534,2029]" pageId="21" pageNumber="22">
<emphasis id="E1F9E47D0D2AB76A8E83F941FE89F881" bold="true" box="[199,322,1786,1812]" pageId="21" pageNumber="22">Diagnosis.</emphasis>
The male pedipalp of
<taxonomicName id="148D43EC0D2AB76A8C78F940FD41F881" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[572,650,1787,1813]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D2AB76A8C78F940FD41F881" box="[572,650,1787,1813]" italics="true" pageId="21" pageNumber="22">A. emu</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D2AB76A8CD4F940FD22F880" bold="true" box="[656,745,1787,1813]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="FACA59060D2AB76A8CD4F940FD22F880" box="[656,745,1787,1813]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
resembles that of
<taxonomicName id="148D43EC0D2AB76A8DF1F940FBE9F880" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[949,1058,1787,1813]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="plicata" status="sp. nov.">
<emphasis id="E1F9E47D0D2AB76A8DF1F940FBE9F880" box="[949,1058,1787,1813]" italics="true" pageId="21" pageNumber="22">A. plicata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D2AB76A8A6DF940FB49F880" bold="true" box="[1065,1154,1787,1813]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="FACA59060D2AB76A8A6DF940FB49F880" box="[1065,1154,1787,1813]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
with the distal portion of its tegular apophyses bearing a retrolateral rounded projection pointing basally and by having a poorly sclerotized basoembolic apophyses with rounded tip (
<figureCitation id="4BB624EA0D2AB76A8C28F8F8FD2DF8C8" box="[620,742,1859,1885]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="21" pageNumber="22">Figs 9CE</figureCitation>
,
<figureCitation id="4BB624EA0D2AB76A8CB5F8F8FC88F8C8" box="[753,835,1859,1885]" captionStart="FIGURE 29" captionStartId="55.[151,250,1667,1691]" captionTargetBox="[161,1419,189,1643]" captionTargetId="figure-20@55.[158,1430,179,1643]" captionTargetPageId="55" captionText="FIGURE 29. Artoria plicata sp. nov., male: A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, holotype (WAM T68736). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387201" httpUri="https://zenodo.org/record/14387201/files/figure.png" pageId="21" pageNumber="22">29CE</figureCitation>
). However, in
<taxonomicName id="148D43EC0D2AB76A8DA0F8F8FBF9F8C9" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[996,1074,1859,1885]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D2AB76A8DA0F8F8FBF9F8C9" box="[996,1074,1859,1885]" italics="true" pageId="21" pageNumber="22">A. emu</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D2AB76A8A7CF8F8FB5BF8C8" bold="true" box="[1080,1168,1859,1885]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="FACA59060D2AB76A8A7CF8F8FB5BF8C8" box="[1080,1168,1859,1885]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the tegular apophysis is shorter and narrower, with an evenly rounded distal margin, and the retrolateral projection has robust, elevated and heavily sclerotized distal and proximal margins, which delimit a median latitudinal concavity (
<figureCitation id="4BB624EA0D2AB76A8AF1F830FAE5F830" box="[1205,1326,1931,1957]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="21" pageNumber="22">Fig. 9C, D</figureCitation>
), while in
<taxonomicName id="148D43EC0D2AB76A8ED3F814FECFF85C" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[151,260,1967,1993]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="plicata" status="sp. nov.">
<emphasis id="E1F9E47D0D2AB76A8ED3F814FECFF85C" box="[151,260,1967,1993]" italics="true" pageId="21" pageNumber="22">A. plicata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D2AB76A8F4FF814FEAEF85C" bold="true" box="[267,357,1967,1993]" pageId="21" pageNumber="22">
<taxonomicNameLabel id="FACA59060D2AB76A8F4FF814FEAEF85C" box="[267,357,1967,1993]" pageId="21" pageNumber="22" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the tegular apophysis has a plicated distal margin, and the retrolateral projection is thin, slightly translucent and with a central and rounded concavity (
<figureCitation id="4BB624EA0D2AB76A8CB2F868FCB4F878" box="[758,895,2003,2029]" captionStart="FIGURE 29" captionStartId="55.[151,250,1667,1691]" captionTargetBox="[161,1419,189,1643]" captionTargetId="figure-20@55.[158,1430,179,1643]" captionTargetPageId="55" captionText="FIGURE 29. Artoria plicata sp. nov., male: A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, holotype (WAM T68736). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387201" httpUri="https://zenodo.org/record/14387201/files/figure.png" pageId="21" pageNumber="22">Fig. 29C, D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="87F268E70D29B7698ED3F913FB73F976" ID-DOI="http://doi.org/10.5281/zenodo.14387135" ID-Zenodo-Dep="14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,250,1704,1728]" targetBox="[151,1425,190,1678]" targetPageId="22" targetType="figure">
<paragraph id="D332386F0D29B7698ED3F913FB73F976" blockId="22.[151,1436,1704,1764]" pageId="22" pageNumber="23">
<emphasis id="E1F9E47D0D29B7698ED3F913FED9F955" bold="true" box="[151,274,1704,1728]" pageId="22" pageNumber="23">FIGURE 9.</emphasis>
<taxonomicName id="148D43EC0D29B7698F5CF913FE59F955" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[280,402,1704,1728]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D29B7698F5CF913FE59F955" box="[280,402,1704,1728]" italics="true" pageId="22" pageNumber="23">Artoria emu</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D29B7698FD3F913FE2CF955" bold="true" box="[407,487,1704,1728]" pageId="22" pageNumber="23">
<taxonomicNameLabel id="FACA59060D29B7698FD3F913FE2CF955" box="[407,487,1704,1728]" pageId="22" pageNumber="23" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm.
</paragraph>
</caption>
<subSubSection id="9B976BE40D29B7688E83F88DFD36FD58" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
<paragraph id="D332386F0D29B7698E83F88DFD13F8C5" blockId="22.[151,1437,1846,2017]" box="[199,728,1846,1872]" pageId="22" pageNumber="23">
<emphasis id="E1F9E47D0D29B7698E83F88DFD13F8C5" bold="true" box="[199,728,1846,1872]" pageId="22" pageNumber="23">
Description. Male (
<typeStatus id="0C3686CD0D29B7698FEFF88DFDDEF8C5" box="[427,533,1846,1872]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
, WAM T98120).
</emphasis>
</paragraph>
<paragraph id="D332386F0D29B7688E83F8E1FECEFEF0" blockId="22.[151,1437,1846,2017]" lastBlockId="23.[151,1437,151,717]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<emphasis id="E1F9E47D0D29B7698E83F8E1FE99F8E1" box="[199,338,1882,1908]" italics="true" pageId="22" pageNumber="23">Total length,</emphasis>
4.09.
<emphasis id="E1F9E47D0D29B7698FD3F8E0FDC7F8E0" box="[407,524,1883,1909]" italics="true" pageId="22" pageNumber="23">Carapace,</emphasis>
length 2.29, width 1.36, brown, with a paler median longitudinal band, and lateral darker bands and a well-marked median line from between eyes to near fovea of same colour (
<figureCitation id="4BB624EA0D29B7698A8CF8C5FAE8F80D" box="[1224,1315,1918,1944]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="22" pageNumber="23">Fig. 9A</figureCitation>
).
<emphasis id="E1F9E47D0D29B7698B7CF8C4FA57F80C" box="[1336,1436,1919,1945]" italics="true" pageId="22" pageNumber="23">Sternum,</emphasis>
greyish-yellow suffused with dark grey stains (
<figureCitation id="4BB624EA0D29B7698CECF819FCCAF829" box="[680,769,1954,1980]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="22" pageNumber="23">Fig. 9B</figureCitation>
).
<emphasis id="E1F9E47D0D29B7698D51F818FC9AF829" box="[789,849,1955,1980]" italics="true" pageId="22" pageNumber="23">Eyes,</emphasis>
diameter: AME: 0.10; ALE: 0.11; PME: 0.22; PLE: 0.19, anterior eye row procurved, evenly spaced.
<emphasis id="E1F9E47D0D29B7698C9CF87DFC91F875" box="[728,858,1990,2016]" italics="true" pageId="22" pageNumber="23">Chelicerae,</emphasis>
evenly brown, two retromarginal teeth, subequal, and two promarginal teeth, apical largest.
<emphasis id="E1F9E47D0D28B7688C34FF2CFD07FF24" box="[624,716,151,177]" italics="true" pageId="23" pageNumber="24">Labium,</emphasis>
dark brown, with pale anterior rim (
<figureCitation id="4BB624EA0D28B7688A23FF2CFB0AFF24" box="[1127,1217,151,177]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="23" pageNumber="24">Fig. 9B</figureCitation>
).
<emphasis id="E1F9E47D0D28B7688A91FF2CFADBFF25" box="[1237,1296,151,176]" italics="true" pageId="23" pageNumber="24">Legs,</emphasis>
background colour yellowish-brown, with femur mostly brown, patella, tibia, metatarsus, and tarsus stained in brown (
<figureCitation id="4BB624EA0D28B7688B7AFF00FF67FF6C" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="23" pageNumber="24">Fig. 9A, B</figureCitation>
).
<emphasis id="E1F9E47D0D28B7688E86FF64FE97FF6C" box="[194,348,223,249]" italics="true" pageId="23" pageNumber="24">Opisthosoma,</emphasis>
length 1.93, width 1.58, dorsum with background colour yellowish-brown, mostly covered with numerous black setae and with less numerous pale-yellow setae over the cardiac mark, which is ⅔ as long as the opisthosoma, and a V-shaped area near the posterior margin (
<figureCitation id="4BB624EA0D28B7688D09FE9CFC63FED4" box="[845,936,295,321]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="23" pageNumber="24">Fig. 9A</figureCitation>
). Venter dark yellow, spinnerets dark brown (
<figureCitation id="4BB624EA0D28B7688EDBFEF0FF33FEF0" box="[159,248,331,357]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="23" pageNumber="24">Fig. 9B</figureCitation>
).
</paragraph>
<paragraph id="D332386F0D28B7688E83FED4FDF3FDA8" blockId="23.[151,1437,151,717]" pageId="23" pageNumber="24">
<emphasis id="E1F9E47D0D28B7688E83FED4FEE7FE1C" box="[199,300,367,393]" italics="true" pageId="23" pageNumber="24">Pedipalp</emphasis>
(
<figureCitation id="4BB624EA0D28B7688F78FED4FE7FFE1C" box="[316,436,367,393]" captionStart="FIGURE 9" captionStartId="22.[151,250,1704,1728]" captionTargetBox="[151,1425,190,1678]" captionTargetId="figure-20@22.[151,1436,179,1678]" captionTargetPageId="22" captionText="FIGURE 9. Artoria emu sp. nov., male:A, dorsal habitus; B, ventral habitus; CE, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). AE, (WAM T98120). Scale bars: A, B, 2.0 mm; CE 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387135" httpUri="https://zenodo.org/record/14387135/files/figure.png" pageId="23" pageNumber="24">Fig. 9CE</figureCitation>
), tibia trapezoidal in ventral view, narrower at its base; cymbium free tip ⅓ as long as the whole cymbium in ventral view, with three distal macrosetae; subtegulum inconspicuous in ventral view. Tegular apophysis small, shaped as an emu head profile in ventral view, with a narrow stalk bent retrolaterally. Basoembolic apophysis small, very sclerotized, subrectangular in ventral view. Embolus comma-like, narrow, and flat, gradually tapering to its acute and flexible tip. Terminal apophysis not very sclerotized, semicircular, broad, wider than the embolus, especially at its sharp apex.
</paragraph>
<paragraph id="D332386F0D28B7688E83FDFDFD36FD58" blockId="23.[151,1437,151,717]" pageId="23" pageNumber="24">
<emphasis id="E1F9E47D0D28B7688E83FDFDFD4BFDF5" bold="true" box="[199,640,582,609]" pageId="23" pageNumber="24">Life history and habitat preferences.</emphasis>
An interpretation of the reproductive period based on
<specimenCount id="C58BF3E60D28B7688AAAFDFCFA9FFDF4" box="[1262,1364,583,609]" count="1" pageId="23" pageNumber="24" type="male">one male</specimenCount>
is not possible. In addition, the pitfall trap in which it was caught was exposed for more than a year. The occurrence of this species in the semi-arid Pilbara region is unusual as it is outside the general range of most
<taxonomicName id="148D43EC0D28B7688ADDFD34FA87FD3C" box="[1177,1356,655,681]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="E1F9E47D0D28B7688ADDFD34FB20FD3C" box="[1177,1259,655,681]" italics="true" pageId="23" pageNumber="24">Artoria</emphasis>
species.
</taxonomicName>
It may represent a relictual species in a small mountain range.
</paragraph>
</subSubSection>
<caption id="87F268E70D28B7688ED3F94FFAC7F899" ID-DOI="http://doi.org/10.5281/zenodo.14387139" ID-Zenodo-Dep="14387139" box="[151,1292,1780,1804]" httpUri="https://zenodo.org/record/14387139/files/figure.png" pageId="23" pageNumber="24" startId="23.[151,250,1780,1804]" targetBox="[157,1430,746,1747]" targetPageId="23" targetType="figure">
<paragraph id="D332386F0D28B7688ED3F94FFAC7F899" blockId="23.[151,1292,1780,1804]" box="[151,1292,1780,1804]" pageId="23" pageNumber="24">
<emphasis id="E1F9E47D0D28B7688ED3F94FFEEBF899" bold="true" box="[151,288,1780,1804]" pageId="23" pageNumber="24">FIGURE 10.</emphasis>
Distribution of
<taxonomicName id="148D43EC0D28B7688F86F94EFDF6F899" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[450,573,1781,1804]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D28B7688F86F94EFDF6F899" box="[450,573,1781,1804]" italics="true" pageId="23" pageNumber="24">Artoria emu</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D28B7688C07F94FFD5FF899" bold="true" box="[579,660,1780,1804]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="FACA59060D28B7688C07F94FFD5FF899" box="[579,660,1780,1804]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
,
<taxonomicName id="148D43EC0D28B7688CE5F94EFCF3F899" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[673,824,1780,1804]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="falcata" status="sp. nov.">
<emphasis id="E1F9E47D0D28B7688CE5F94EFCF3F899" box="[673,824,1780,1804]" italics="true" pageId="23" pageNumber="24">Artoria falcata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D28B7688D7BF94FFC44F899" bold="true" box="[831,911,1780,1804]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="FACA59060D28B7688D7BF94FFC44F899" box="[831,911,1780,1804]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="148D43EC0D28B7688D83F94EFAC2F899" authority="Simon, 1909" authorityName="Simon" authorityYear="1909" box="[967,1289,1780,1804]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="flavimana">
<emphasis id="E1F9E47D0D28B7688D83F94EFBB6F899" box="[967,1149,1780,1804]" italics="true" pageId="23" pageNumber="24">Artoria flavimana</emphasis>
<bibRefCitation id="B71C459E0D28B7688AC0F94FFAC2F899" author="Simon, E." box="[1156,1289,1780,1804]" pageId="23" pageNumber="24" pagination="155 - 212" refId="ref59649" refString="Simon, E. (1909) Araneae, 2 me partie. In: Michaelsen, W. &amp; Hartmeyer, R. (Eds.), Die Fauna Sudwest-Australiens. Ergebnisse der Hamburger sudwest-australischen Forschungsreise 1905. Gustav Fischer Verlag, Jena, pp. 155 - 212." type="book chapter" year="1909">Simon, 1909</bibRefCitation>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="9B976BE40D28B7678E83F8E4FC92FF40" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="discussion">
<paragraph id="D332386F0D28B7678E83F8E4FC92FF40" blockId="23.[151,1436,1886,2021]" lastBlockId="24.[151,1436,151,285]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
<emphasis id="E1F9E47D0D28B7688E83F8E4FEF0F8EC" bold="true" box="[199,315,1887,1913]" pageId="23" pageNumber="24">Remarks.</emphasis>
<taxonomicName id="148D43EC0D28B7688F05F8E4FE02F8ED" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[321,457,1887,1913]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D28B7688F05F8E4FE02F8ED" box="[321,457,1887,1913]" italics="true" pageId="23" pageNumber="24">Artoria emu</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D28B7688F8AF8E4FDECF8EC" bold="true" box="[462,551,1887,1913]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="FACA59060D28B7688F8AF8E4FDECF8EC" box="[462,551,1887,1913]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is only known from the
<typeStatus id="0C3686CD0D28B7688D70F8E5FC5DF8ED" box="[820,918,1886,1912]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
male found in the southern Hamersley Ranges in the Pilbara region of
<collectingRegion id="1149F68D0D28B7688FDAF838FDA3F809" box="[414,616,1922,1949]" country="Australia" name="Western Australia" pageId="23" pageNumber="24">Western Australia</collectingRegion>
. This distribution is outside the general Bassian and Torresian occurrence of the genus. No further specimens were found in the comprehensive Pilbara survey in which
<taxonomicName id="148D43EC0D28B7688AA0F81CFAFEF855" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[1252,1333,1959,1985]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="emu" status="sp. nov.">
<emphasis id="E1F9E47D0D28B7688AA0F81CFAFEF855" box="[1252,1333,1959,1985]" italics="true" pageId="23" pageNumber="24">A. emu</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D28B7688B7BF81CFA57F854" bold="true" box="[1343,1436,1959,1985]" pageId="23" pageNumber="24">
<taxonomicNameLabel id="FACA59060D28B7688B7BF81CFA57F854" box="[1343,1436,1959,1985]" pageId="23" pageNumber="24" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
was collected (e.g.,
<bibRefCitation id="B71C459E0D28B7688F3EF870FD93F870" author="Durrant, B. J. &amp; Harvey, M. S. &amp; Framenau, V. W. &amp; Ott, R. &amp; Waldock, J. M." box="[378,600,1994,2021]" pageId="23" pageNumber="24" pagination="185 - 204" refId="ref57811" refString="Durrant, B. J., Harvey, M. S., Framenau, V. W., Ott, R. &amp; Waldock, J. M. (2010) Patterns in the composition of ground-dwelling spider communities in the Pilbara bioregion, Western Australia. Records of the Western Australian Museum, Supplement, 78, 185 - 204. https: // doi. org / 10.18195 / issn. 0313 - 122 x. 78 (1). 2010.185 - 204" type="journal article" year="2010">
Durrant
<emphasis id="E1F9E47D0D28B7688F9DF870FDD8F871" box="[473,531,1994,2020]" italics="true" pageId="23" pageNumber="24">et al.</emphasis>
2010
</bibRefCitation>
). As this single record is not from a conservation reserve, we propose the Priority 1 (P1) conservation status for this species (“Poorly known species—known from few localities, none on conservation land”) (WA Biodiversity Conservation Act 2016).
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D27B7678E83FF65FD17FE88" pageId="24" pageNumber="25" type="distribution">
<paragraph id="D332386F0D27B7678E83FF65FD17FE88" blockId="24.[151,1436,151,285]" pageId="24" pageNumber="25">
<emphasis id="E1F9E47D0D27B7678E83FF65FEAAFF6D" bold="true" box="[199,353,222,248]" pageId="24" pageNumber="25">Distribution.</emphasis>
Sole male known from Giles Point, in a semi-arid area of the Hamersley Ranges in the Pilbara region in north-western
<collectingRegion id="1149F68D0D27B7678FE0FEB8FDA7FE88" box="[420,620,259,285]" country="Australia" name="Western Australia" pageId="24" pageNumber="25">Western Australia</collectingRegion>
(
<figureCitation id="4BB624EA0D27B7678C3FFEB8FD05FE88" box="[635,718,259,285]" captionStart="FIGURE 10" captionStartId="23.[151,250,1780,1804]" captionTargetBox="[157,1430,746,1747]" captionTargetId="figure-265@23.[151,1436,738,1754]" captionTargetPageId="23" captionText="FIGURE 10. Distribution of Artoria emu sp. nov., Artoria falcata sp. nov., and Artoria flavimana Simon, 1909." figureDoi="http://doi.org/10.5281/zenodo.14387139" httpUri="https://zenodo.org/record/14387139/files/figure.png" pageId="24" pageNumber="25">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>