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<document id="98A37E6FA55DC1D0AF9355C612F1D55B" ID-DOI="10.11646/zootaxa.5547.1.1" ID-ISSN="1175-5334" ID-Zenodo-Dep="14387105" ID-ZooBank="2089C3EC-8FBB-43E6-A6C5-9E6B6AD512D6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1733920242413" checkinUser="plazi" docAuthor="Do Prado, André W., Baptista, Renner Luiz Cerqueira &amp; Framenau, Volker W." docDate="2024" docId="5B2489790D10B74D8ED3F94CFF0FF90C" docLanguage="en" docName="zootaxa.5547.1.1.pdf" docOrigin="Zootaxa 5547 (1)" docSource="https://doi.org/10.11646/zootaxa.5547.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.19:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="19" docTitle="Artoria pileata Prado, Baptista &amp; Framenau, 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="51" masterDocId="A71DF1010D3FB77F8E44FFBBFFCBFF95" masterDocTitle="Taxonomy of the wolf spider genus Artoria in Western Australia (Araneae, Lycosidae, Artoriinae)" masterLastPageNumber="81" masterPageNumber="1" pageNumber="48" updateTime="1733921097182" updateUser="ExternalLinkService">
<mods:mods id="6BEF0AAE1A30FA48F723898B92F7ACB4" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="25C221D341CA0BAE24EB127F347D100A">
<mods:title id="DE2E7B1A0504367628A5E5E2FA3D3F06">Taxonomy of the wolf spider genus Artoria in Western Australia (Araneae, Lycosidae, Artoriinae)</mods:title>
</mods:titleInfo>
<mods:name id="C3BFA7D61C2DFEA9EEE0FA35A301ADF6" type="personal">
<mods:role id="FD1B075302FCB2AFA763393C64963DD6">
<mods:roleTerm id="14D264F62463E35C077CD0482358329E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="779B0B6D32C03C833E559290C8FE4CD8">Do Prado, André W.</mods:namePart>
<mods:nameIdentifier id="9000AE8ACA8D1D8EAACD07637E49AB02" type="ORCID">0000-0002-3536-5654</mods:nameIdentifier>
<mods:affiliation id="BD825FA782AD0DBF09B27430A473578F">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia &amp; andre. wanderleydoprado @ murdoch. edu. au; https: // orcid. org / 0000 - 0002 - 3536 - 5654</mods:affiliation>
<mods:nameIdentifier id="40C09ABDFAD52C192257709F6C6B70A1" type="email">andre.wanderleydoprado@murdoch.edu.au</mods:nameIdentifier>
</mods:name>
<mods:name id="39305C904DDD0717EED25953D347A6C0" type="personal">
<mods:role id="FC6B1C848AD4DCB0DC7243F99C7E682E">
<mods:roleTerm id="621F77147D351F4B0838EDC29BC26EA2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B274FF9199440A84A507B74419B8D7F">Baptista, Renner Luiz Cerqueira</mods:namePart>
<mods:nameIdentifier id="CD61356B59952FD87AE2C8A5E7178ED6" type="ORCID">0000-0002-2372-5034</mods:nameIdentifier>
<mods:affiliation id="793DD8222F24ABCD99C171D8C62EA5CA">Departamento de Zoologia, Instituto de Biologia, Universidade do Brasil / Universidade Federal do Rio de Janeiro, Rio de Janeiro, Brazil &amp; baptistr @ gmail. com; https: // orcid. org / 0000 - 0002 - 2372 - 5034</mods:affiliation>
<mods:nameIdentifier id="A11A7DEC7C14882669BCBB7BE5AF7D22" type="email">baptistr@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="C5EF6F0FA8C046309FA7F61A80D23120" type="personal">
<mods:role id="0FE280C2894DF043FC53062E9BCA93EE">
<mods:roleTerm id="64B4096BBE8FFA03881C446687F5F42E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C12DBD89A4CE0418ADB4015213723D88">Framenau, Volker W.</mods:namePart>
<mods:nameIdentifier id="65F72E602FECC670215295CC0157B340" type="ORCID">0000-0002-7724-3831</mods:nameIdentifier>
<mods:affiliation id="4B91E5CEC98A102558830F8430BDB52A">Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Terrestrial Zoology, Western Australian Museum, Locked Bag 49, Welshpool DC, Western Australia 6986, Australia &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; Department of Invertebrates, Museum of Nature Hamburg - Zoology, Leibnitz Institute for the Analysis of Biodiversity Change (LIB), Martin-Luther-King-Platz 3, 20146 Hamburg, Germany &amp; Harry Butler Institute, Murdoch University, 90 South St, Murdoch, Western Australia 6150, Australia &amp; volker. framenau @ murdoch. edu. au; https: // orcid. org / 0000 - 0002 - 7724 - 3831 * Corresponding author</mods:affiliation>
<mods:nameIdentifier id="5C3B64EF6351C07FD62137928A5AF78B" type="email">volker.framenau@murdoch.edu.au</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="26149D6F746BD8CBB548E6EF656E56C2">text</mods:typeOfResource>
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<mods:title id="6A938FACFE094E34E2730FD8DEABFCBC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="59F0E16FBFE7A5C4B96455ED1948F945">
<mods:date id="E316E855AAF7793CEDCF16D14557DBDD">2024</mods:date>
<mods:detail id="A43A2B1EA43652A3BAA777324B512676" type="pubDate">
<mods:number id="92FD1BECDDF4D4850F84833EC0D15F19">2024-12-09</mods:number>
</mods:detail>
<mods:detail id="13B9A0CA79B615500DBB011BCD036D1E" type="volume">
<mods:number id="96B90B08FA9E2FFFB3B956B2345FB4BF">5547</mods:number>
</mods:detail>
<mods:detail id="05FB03EACC28FC54C207CD3A10FF9468" type="issue">
<mods:number id="39639F2B1E26350144628B574E313F4B">1</mods:number>
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<mods:start id="5D0A69D65446254D3DBC3CBDC9815E8E">1</mods:start>
<mods:end id="914867C0D49B578C670F9EF4C7377865">81</mods:end>
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<mods:location id="E78660F627C2900B9A7CE04AB1810B19">
<mods:url id="E67D6981993707EAA91BEFDC73934A7C">https://doi.org/10.11646/zootaxa.5547.1.1</mods:url>
</mods:location>
<mods:classification id="38B2D9BA9B8285CDECB5BE51821F5CE2">journal article</mods:classification>
<mods:identifier id="348728B97D69EF3F060828041F464B99" type="DOI">10.11646/zootaxa.5547.1.1</mods:identifier>
<mods:identifier id="EF2322CDD93A1495B86F1CCD43A2563E" type="ISSN">1175-5334</mods:identifier>
<mods:identifier id="5D3D10BA2E5C9C022ED8C6692629E6C0" type="Zenodo-Dep">14387105</mods:identifier>
<mods:identifier id="804D5EF322AE14E4E4F4CCBFFE138C0F" type="ZooBank">2089C3EC-8FBB-43E6-A6C5-9E6B6AD512D6</mods:identifier>
</mods:mods>
<treatment id="5B2489790D10B74D8ED3F94CFF0FF90C" LSID="urn:lsid:plazi:treatment:5B2489790D10B74D8ED3F94CFF0FF90C" httpUri="http://treatment.plazi.org/id/5B2489790D10B74D8ED3F94CFF0FF90C" lastPageId="50" lastPageNumber="51" pageId="47" pageNumber="48">
<subSubSection id="9B976BE40D10B7508ED3F94CFE66F887" box="[151,429,1783,1810]" pageId="47" pageNumber="48" type="nomenclature">
<paragraph id="D332386F0D10B7508ED3F94CFE66F887" blockId="47.[151,978,1783,1882]" box="[151,429,1783,1810]" pageId="47" pageNumber="48">
<heading id="887A8F030D10B7508ED3F94CFE66F887" bold="true" box="[151,429,1783,1810]" fontSize="11" level="2" pageId="47" pageNumber="48" reason="4">
<emphasis id="E1F9E47D0D10B7508ED3F94CFE66F887" bold="true" box="[151,429,1783,1810]" pageId="47" pageNumber="48">
<taxonomicName id="148D43EC0D10B7508ED3F94CFE83F887" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[151,328,1783,1810]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="pileata" status="sp. nov.">
<emphasis id="E1F9E47D0D10B7508ED3F94CFE83F887" bold="true" box="[151,328,1783,1810]" italics="true" pageId="47" pageNumber="48">Artoria pileata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="FACA59060D10B7508F0BF943FE66F887" box="[335,429,1784,1810]" pageId="47" pageNumber="48" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D10B7508ED3F8A0FC19F8CF" pageId="47" pageNumber="48" type="description">
<paragraph id="D332386F0D10B7508ED3F8A0FEDEF8A0" blockId="47.[151,978,1783,1882]" box="[151,277,1819,1845]" pageId="47" pageNumber="48">
<figureCitation id="4BB624EA0D10B7508ED3F8A0FEDEF8A0" box="[151,277,1819,1845]" captionStart-0="FIGURE 24" captionStart-1="FIGURE 25" captionStart-2="FIGURE 26" captionStartId-0="48.[151,250,1779,1803]" captionStartId-1="49.[151,250,1618,1642]" captionStartId-2="51.[151,250,1414,1438]" captionTargetBox-0="[157,1425,190,1755]" captionTargetBox-1="[151,1409,189,1593]" captionTargetBox-2="[160,1428,383,1381]" captionTargetId-0="figure-20@48.[151,1436,179,1755]" captionTargetId-1="figure-20@49.[151,1436,179,1593]" captionTargetId-2="figure-101@51.[151,1436,378,1389]" captionTargetPageId-0="48" captionTargetPageId-1="49" captionTargetPageId-2="51" captionText-0="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." captionText-1="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." captionText-2="FIGURE 26. Distribution of Artoria pileata sp. nov., Artoria pinnata sp. nov., Artoria plicata sp. nov., Artoria retorta sp. nov., and Artoria schizocoides Framenau &amp; Hebets, 2007." figureDoi-0="http://doi.org/10.5281/zenodo.14387183" figureDoi-1="http://doi.org/10.5281/zenodo.14387187" figureDoi-2="http://doi.org/10.5281/zenodo.14387191" httpUri-0="https://zenodo.org/record/14387183/files/figure.png" httpUri-1="https://zenodo.org/record/14387187/files/figure.png" httpUri-2="https://zenodo.org/record/14387191/files/figure.png" pageId="47" pageNumber="48">Figs 24–26</figureCitation>
</paragraph>
<paragraph id="D332386F0D10B7508ED3F884FC19F8CF" blockId="47.[151,978,1783,1882]" box="[151,978,1855,1882]" pageId="47" pageNumber="48">
<uri id="A71C346D0D10B7508ED3F884FC19F8CF" box="[151,978,1855,1882]" pageId="47" pageNumber="48">
urn:lsid:zoobank.org:act:
<uuid id="A72B02BA0D10B7508FE2F884FC19F8CF" box="[422,978,1855,1882]" pageId="47" pageNumber="48">6EBDD579-8BDE-40F4-ACFF-63C6340494A8</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D10B74E8ED3F833FDE0F8FD" lastPageId="49" lastPageNumber="50" pageId="47" pageNumber="48" type="materials_examined">
<paragraph id="D332386F0D10B7508ED3F833FD72F850" blockId="47.[151,1437,1927,1990]" pageId="47" pageNumber="48">
<emphasis id="E1F9E47D0D10B7508ED3F833FE8DF834" bold="true" box="[151,326,1927,1954]" pageId="47" pageNumber="48">Type material.</emphasis>
<materialsCitation id="63E532320D10B7508F0AF83CFD72F850" collectingDate="1971-10-13" collectionCode="WAM" collectorName="R. J. McKay" country="Australia" latitude="-16.666666" location="Old Lissadell Station" longLatPrecision="1281" longitude="138.55" pageId="47" pageNumber="48" specimenCode="T162498" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<typeStatus id="0C3686CD0D10B7508F0AF83CFE7DF834" box="[334,438,1927,1953]" pageId="47" pageNumber="48" type="holotype">Holotype</typeStatus>
male,
<location id="D6526EB40D10B7508C40F83CFD3EF834" LSID="urn:lsid:plazi:treatment:5B2489790D10B74D8ED3F94CFF0FF90C:D6526EB40D10B7508C40F83CFD3EF834" box="[516,757,1927,1953]" country="Australia" latitude="-16.666666" longLatPrecision="1281" longitude="138.55" name="Old Lissadell Station" pageId="47" pageNumber="48" stateProvince="Western Australia">Old Lissadell Station</location>
, near,
<collectingRegion id="1149F68D0D10B7508D04F833FBC0F834" box="[832,1035,1927,1954]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">Western Australia</collectingRegion>
,
<collectingCountry id="AB9A78FF0D10B7508A52F833FB73F837" box="[1046,1208,1928,1954]" name="Australia" pageId="47" pageNumber="48">AUSTRALIA</collectingCountry>
,
<geoCoordinate id="B6B95EA80D10B7508A87F83CFAD4F834" box="[1219,1311,1927,1953]" degrees="16" direction="south" minutes="40" orientation="latitude" pageId="47" pageNumber="48" precision="925" value="-16.666666">16º40’S</geoCoordinate>
,
<geoCoordinate id="B6B95EA80D10B7508B68F833FA5CF837" box="[1324,1431,1928,1954]" degrees="138" direction="east" minutes="33" orientation="longitude" pageId="47" pageNumber="48" precision="925" value="138.55">138º33’E</geoCoordinate>
,
<date id="A7331EAF0D10B7508ED3F817FE9EF853" box="[151,341,1963,1990]" pageId="47" pageNumber="48" value="1971-10-13">
<collectingDate id="B777E7470D10B7508ED3F817FE9EF853" box="[151,341,1963,1990]" pageId="47" pageNumber="48" value="1971-10-13">13 October 1971</collectingDate>
</date>
,
<collectorName id="7E785DB90D10B7508F25F817FE23F853" box="[353,488,1964,1990]" pageId="47" pageNumber="48">R.J. McKay</collectorName>
(
<collectionCode id="B59CA0AA0D10B7508FB2F817FDF6F853" box="[502,573,1964,1990]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="47" pageNumber="48" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D10B7508C07F810FD66F850" box="[579,685,1963,1989]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="47" pageNumber="48" type="Museum">T162498</specimenCode>
).
</materialsCitation>
</paragraph>
<caption id="87F268E70D0FB74F8ED3F948FE9FF8C6" ID-DOI="http://doi.org/10.5281/zenodo.14387183" ID-Zenodo-Dep="14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="48" pageNumber="49" startId="48.[151,250,1779,1803]" targetBox="[157,1425,190,1755]" targetPageId="48" targetType="figure">
<paragraph id="D332386F0D0FB74F8ED3F948FE9FF8C6" blockId="48.[151,1436,1779,1875]" pageId="48" pageNumber="49">
<emphasis id="E1F9E47D0D0FB74F8ED3F948FEE8F89E" bold="true" box="[151,291,1779,1803]" pageId="48" pageNumber="49">FIGURE 24.</emphasis>
<taxonomicName id="148D43EC0D0FB74F8F68F94FFE0DF89E" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[300,454,1779,1803]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="pileata" status="sp. nov.">
<emphasis id="E1F9E47D0D0FB74F8F68F94FFE0DF89E" box="[300,454,1779,1803]" italics="true" pageId="48" pageNumber="49">Artoria pileata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D0FB74F8F94F948FDE8F89E" bold="true" box="[464,547,1779,1803]" pageId="48" pageNumber="49">
<taxonomicNameLabel id="FACA59060D0FB74F8F94F948FDE8F89E" box="[464,547,1779,1803]" pageId="48" pageNumber="49" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm.
</paragraph>
</caption>
<caption id="87F268E70D0EB74E8ED3F9E9FB8FF91B" ID-DOI="http://doi.org/10.5281/zenodo.14387187" ID-Zenodo-Dep="14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="49" pageNumber="50" startId="49.[151,250,1618,1642]" targetBox="[151,1409,189,1593]" targetPageId="49" targetType="figure">
<paragraph id="D332386F0D0EB74E8ED3F9E9FB8FF91B" blockId="49.[151,1436,1618,1679]" pageId="49" pageNumber="50">
<emphasis id="E1F9E47D0D0EB74E8ED3F9E9FEE9F9FF" bold="true" box="[151,290,1618,1642]" pageId="49" pageNumber="50">FIGURE 25.</emphasis>
<taxonomicName id="148D43EC0D0EB74E8F6EF9E8FE09F9FE" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[298,450,1619,1643]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pileata" status="sp. nov.">
<emphasis id="E1F9E47D0D0EB74E8F6EF9E8FE09F9FE" box="[298,450,1619,1643]" italics="true" pageId="49" pageNumber="50">Artoria pileata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D0EB74E8F8FF9E8FDD6F9FE" bold="true" box="[459,541,1619,1643]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="FACA59060D0EB74E8F8FF9E8FDD6F9FE" box="[459,541,1619,1643]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm.
</paragraph>
</caption>
<paragraph id="D332386F0D0EB74E8E83F95BFDE0F8FD" blockId="49.[151,1437,1760,2039]" pageId="49" pageNumber="50">
<emphasis id="E1F9E47D0D0EB74E8E83F95BFAB5F96F" bold="true" box="[199,1406,1760,1787]" pageId="49" pageNumber="50">
Other material examined (
<specimenCount id="C58BF3E60D0EB74E8C43F95BFDAFF96E" box="[519,612,1760,1787]" count="6" pageId="49" pageNumber="50" type="male">6 males</specimenCount>
,
<specimenCount id="C58BF3E60D0EB74E8C35F95AFD3AF96F" box="[625,753,1760,1787]" count="14" pageId="49" pageNumber="50" type="female">14 females</specimenCount>
,
<specimenCount id="C58BF3E60D0EB74E8CBAF95AFC8AF96F" box="[766,833,1760,1787]" count="2" pageId="49" pageNumber="50" type="generic">2 juv.</specimenCount>
in 8 records).
<materialsCitation id="63E532320D0EB74E8DABF95BFAB5F96F" box="[1007,1406,1760,1787]" country="Australia" location="Western Australia" pageId="49" pageNumber="50" specimenCount="1" stateProvince="Western Australia">
<collectingCountry id="AB9A78FF0D0EB74E8DABF95BFB53F96F" box="[1007,1176,1760,1786]" name="Australia" pageId="49" pageNumber="50">AUSTRALIA</collectingCountry>
:
<collectingRegion id="1149F68D0D0EB74E8AE0F95AFAB5F96F" box="[1188,1406,1760,1787]" country="Australia" name="Western Australia" pageId="49" pageNumber="50">Western Australia</collectingRegion>
</materialsCitation>
</emphasis>
:
<materialsCitation id="63E532320D0EB74E8BCAF95AFBF8F88A" collectionCode="WAM" country="Australia" latitude="-16.666666" location="Old Lissadell Station" longLatPrecision="1281" longitude="138.55" pageId="49" pageNumber="50" specimenCode="T86691" specimenCount="10" specimenCount-female="8" specimenCount-male="2" stateProvince="Western Australia">
<specimenCount id="C58BF3E60D0EB74E8BCAF95AFF10F88A" count="2" pageId="49" pageNumber="50" type="male">2 males</specimenCount>
,
<specimenCount id="C58BF3E60D0EB74E8EACF8BEFE93F88A" box="[232,344,1797,1823]" count="8" pageId="49" pageNumber="50" type="female">8 females</specimenCount>
,
<location id="D6526EB40D0EB74E8F20F8BEFD9EF88A" LSID="urn:lsid:plazi:treatment:5B2489790D10B74D8ED3F94CFF0FF90C:D6526EB40D0EB74E8F20F8BEFD9EF88A" box="[356,597,1797,1823]" country="Australia" latitude="-16.666666" longLatPrecision="1281" longitude="138.55" name="Old Lissadell Station" pageId="49" pageNumber="50" stateProvince="Western Australia">Old Lissadell Station</location>
, near,
<geoCoordinate id="B6B95EA80D0EB74E8CE4F8BEFD30F88A" box="[672,763,1797,1823]" degrees="16" direction="south" minutes="40" orientation="latitude" pageId="49" pageNumber="50" precision="925" value="-16.666666">16º40’S</geoCoordinate>
,
<geoCoordinate id="B6B95EA80D0EB74E8D4CF8BEFCB9F88A" box="[776,882,1797,1823]" degrees="138" direction="east" minutes="33" orientation="longitude" pageId="49" pageNumber="50" precision="925" value="138.55">138º33’E</geoCoordinate>
(
<collectionCode id="B59CA0AA0D0EB74E8DC5F8BEFC03F88A" box="[897,968,1797,1823]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D0EB74E8D8BF8BEFBE0F88A" box="[975,1067,1797,1823]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">T86691</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="63E532320D0EB74E8A7BF8BEFEC2F8D6" collectionCode="WAM" country="Australia" latitude="-16.666666" location="Old Lissadell Station" longLatPrecision="1281" longitude="138.55" pageId="49" pageNumber="50" specimenCode="T162496" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount id="C58BF3E60D0EB74E8A7BF8BEFB6FF88A" box="[1087,1188,1797,1823]" count="1" pageId="49" pageNumber="50" type="female">1 female</specimenCount>
, same locality (
<collectionCode id="B59CA0AA0D0EB74E8B12F8BEFA56F88A" box="[1366,1437,1797,1823]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D0EB74E8ED3F892FECAF8D6" box="[151,257,1833,1859]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">T162496</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="63E532320D0EB74E8F51F892FD2DF8D6" box="[277,742,1833,1859]" collectionCode="WAM" country="Australia" latitude="-16.666666" location="Old Lissadell Station" longLatPrecision="1281" longitude="138.55" pageId="49" pageNumber="50" specimenCode="T162497" specimenCount="1" specimenCount-female="1" stateProvince="Western Australia">
<specimenCount id="C58BF3E60D0EB74E8F51F892FEB2F8D6" box="[277,377,1833,1859]" count="1" pageId="49" pageNumber="50" type="female">1 female</specimenCount>
, same locality (
<collectionCode id="B59CA0AA0D0EB74E8C63F892FDA5F8D6" box="[551,622,1833,1859]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D0EB74E8C30F892FD15F8D6" box="[628,734,1833,1859]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">T162497</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="63E532320D0EB74E8CB6F892FB6BF8D6" box="[754,1184,1833,1859]" collectionCode="WAM" country="Australia" latitude="-16.666666" location="Old Lissadell Station" longLatPrecision="1281" longitude="138.55" pageId="49" pageNumber="50" specimenCode="T86690" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount id="C58BF3E60D0EB74E8CB6F892FC8BF8D6" box="[754,832,1833,1859]" count="1" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
, same locality (
<collectionCode id="B59CA0AA0D0EB74E8DAAF892FBFEF8D6" box="[1006,1077,1833,1859]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D0EB74E8A78F892FB5CF8D6" box="[1084,1175,1833,1859]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">T86690</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="63E532320D0EB74E8AEFF892FDECF8FD" collectionCode="WAM" country="Australia" latitude="-16.416666" location="Walcott Inlet" longLatPrecision="1282" longitude="124.916664" pageId="49" pageNumber="50" specimenCode="T64059" specimenCount="1" specimenCount-male="1" stateProvince="Western Australia">
<specimenCount id="C58BF3E60D0EB74E8AEFF892FB32F8D6" box="[1195,1273,1833,1859]" count="1" pageId="49" pageNumber="50" type="male">1 male</specimenCount>
,
<location id="D6526EB40D0EB74E8B40F892FA53F8D6" LSID="urn:lsid:plazi:treatment:5B2489790D10B74D8ED3F94CFF0FF90C:D6526EB40D0EB74E8B40F892FA53F8D6" box="[1284,1432,1833,1859]" country="Australia" latitude="-16.416666" longLatPrecision="1282" longitude="124.916664" name="Walcott Inlet" pageId="49" pageNumber="50" stateProvince="Western Australia">Walcott Inlet</location>
,
<geoCoordinate id="B6B95EA80D0EB74E8ED3F8F6FF38F8FD" box="[151,243,1869,1896]" degrees="16" direction="south" minutes="25" orientation="latitude" pageId="49" pageNumber="50" precision="925" value="-16.416666">16º25’S</geoCoordinate>
,
<geoCoordinate id="B6B95EA80D0EB74E8EBBF8F6FEA3F8FD" box="[255,360,1869,1896]" degrees="124" direction="east" minutes="55" orientation="longitude" pageId="49" pageNumber="50" precision="925" value="124.916664">124º55’E</geoCoordinate>
(
<collectionCode id="B59CA0AA0D0EB74E8F32F8F6FE76F8F2" box="[374,445,1869,1895]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">WAM</collectionCode>
<specimenCode id="832B90140D0EB74E8F87F8F6FDD4F8FD" box="[451,543,1869,1896]" collectionCode="WAM" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="49" pageNumber="50" type="Museum">T64059</specimenCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D0EB74E8E83F8CAFF3CF846" pageId="49" pageNumber="50" type="etymology">
<paragraph id="D332386F0D0EB74E8E83F8CAFF3CF846" blockId="49.[151,1437,1760,2039]" pageId="49" pageNumber="50">
<emphasis id="E1F9E47D0D0EB74E8E83F8CAFE85F81E" bold="true" box="[199,334,1905,1931]" pageId="49" pageNumber="50">Etymology.</emphasis>
The specific epithet
<taxonomicName id="148D43EC0D0EB74E8C74F8CAFD40F81E" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[560,651,1905,1931]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pileata">
“
<emphasis id="E1F9E47D0D0EB74E8C79F8CAFD40F81E" box="[573,651,1905,1931]" italics="true" pageId="49" pageNumber="50">pileata</emphasis>
</taxonomicName>
” is a Latin adjective meaning “with a cap”, referring to the conspicuous and wide posterior projection of the anterior margin of the atrium of the female’s epigyne, that resembles the flap of a cap.
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D0EB74D8E83F867FAD7FE38" lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="diagnosis">
<paragraph id="D332386F0D0EB74E8E83F867FA57F862" blockId="49.[151,1437,1760,2039]" box="[199,1436,2012,2039]" pageId="49" pageNumber="50">
<emphasis id="E1F9E47D0D0EB74E8E83F867FE89F863" bold="true" box="[199,322,2012,2038]" pageId="49" pageNumber="50">Diagnosis.</emphasis>
The pedipalps of male
<taxonomicName id="148D43EC0D0EB74E8C0EF866FD39F862" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[586,754,2013,2039]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="pileata" status="sp. nov.">
<emphasis id="E1F9E47D0D0EB74E8C0EF866FD39F862" box="[586,754,2013,2039]" italics="true" pageId="49" pageNumber="50">Artoria pileata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D0EB74E8CBDF866FC98F862" bold="true" box="[761,851,2013,2039]" pageId="49" pageNumber="50">
<taxonomicNameLabel id="FACA59060D0EB74E8CBDF866FC98F862" box="[761,851,2013,2039]" pageId="49" pageNumber="50" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are similar to those of
<taxonomicName id="148D43EC0D0EB74E8A1CF866FA57F862" authority="Framenau, 2002" authorityName="Framenau" authorityYear="2002" box="[1112,1436,2013,2039]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="albopedipalpis">
<emphasis id="E1F9E47D0D0EB74E8A1CF866FAD6F862" box="[1112,1309,2013,2039]" italics="true" pageId="49" pageNumber="50">A. albopedipalpis</emphasis>
<bibRefCitation id="B71C459E0D0EB74E8B61F866FA57F862" author="Framenau, V. W." box="[1317,1436,2013,2039]" pageId="49" pageNumber="50" pagination="209 - 235" refId="ref57945" refString="Framenau, V. W. (2002) Review of the wolf spider genus Artoria Thorell (Araneae: Lycosidae). Invertebrate Systematics, 16, 209 - 235. https: // doi. org / 10.1071 / IT 01028" type="journal article" year="2002">Framenau,</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="D332386F0D0DB74D8ED3FF2CFAD7FE38" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
2002, a species that does not occur in
<collectingRegion id="1149F68D0D0DB74D8C05FF2CFCC7FF24" box="[577,780,151,177]" country="Australia" name="Western Australia" pageId="50" pageNumber="51">Western Australia</collectingRegion>
, in having a sinuous tegular apophysis with an acute small projection on its prolateral margin, with a distal inner concavity, and a curved apex. However, in
<taxonomicName id="148D43EC0D0DB74D8B45FF00FABAFF40" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[1281,1393,187,213]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pileata" status="sp. nov.">
<emphasis id="E1F9E47D0D0DB74D8B45FF00FABAFF40" box="[1281,1393,187,213]" italics="true" pageId="50" pageNumber="51">A. pileata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D0DB74D8B3FFF00FF03FF6C" bold="true" pageId="50" pageNumber="51">
<taxonomicNameLabel id="FACA59060D0DB74D8B3FFF00FF03FF6C" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
the tegular apophysis is narrower, and more sclerotized, and the ventral margin of its distal part is thicker, less projected ventrally, appearing truncated in retrolateral view (
<figureCitation id="4BB624EA0D0DB74D8D08FEB8FC1CFE88" box="[844,983,259,285]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24C, D</figureCitation>
), while in
<taxonomicName id="148D43EC0D0DB74D8A17FEB8FAD1FE88" authorityName="Framenau" authorityYear="2002" box="[1107,1306,259,285]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="albopedipalpis">
<emphasis id="E1F9E47D0D0DB74D8A17FEB8FAD1FE88" box="[1107,1306,259,285]" italics="true" pageId="50" pageNumber="51">A. albopedipalpis</emphasis>
</taxonomicName>
the tegular apophysis is wider, more sclerotized, with the ventral margin of its distal part thinner, very projected ventrally as an acute tip (
<bibRefCitation id="B71C459E0D0DB74D8F6BFEF0FE2CFEF0" author="Framenau, V. W." box="[303,487,331,357]" pageId="50" pageNumber="51" pagination="209 - 235" refId="ref57945" refString="Framenau, V. W. (2002) Review of the wolf spider genus Artoria Thorell (Araneae: Lycosidae). Invertebrate Systematics, 16, 209 - 235. https: // doi. org / 10.1071 / IT 01028" type="journal article" year="2002">Framenau 2002</bibRefCitation>
; fig. 1B). Females of
<taxonomicName id="148D43EC0D0DB74D8CA3FEF0FC9CFEF0" authorityName="Do Prado &amp; Baptista &amp; Framenau" authorityYear="2024" box="[743,855,331,357]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="pileata" status="sp. nov.">
<emphasis id="E1F9E47D0D0DB74D8CA3FEF0FC9CFEF0" box="[743,855,331,357]" italics="true" pageId="50" pageNumber="51">A. pileata</emphasis>
</taxonomicName>
<emphasis id="E1F9E47D0D0DB74D8D25FEF0FC76FEF0" bold="true" box="[865,957,331,357]" pageId="50" pageNumber="51">
<taxonomicNameLabel id="FACA59060D0DB74D8D25FEF0FC76FEF0" box="[865,957,331,357]" pageId="50" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
are distinguishable among those of other described
<taxonomicName id="148D43EC0D0DB74D8F4FFED4FE96FE1C" authorityName="Thorell" authorityYear="1877" box="[267,349,367,393]" class="Arachnida" family="Lycosidae" genus="Artoria" kingdom="Animalia" order="Araneae" pageId="50" pageNumber="51" phylum="Arthropoda" rank="genus">
<emphasis id="E1F9E47D0D0DB74D8F4FFED4FE96FE1C" box="[267,349,367,393]" italics="true" pageId="50" pageNumber="51">Artoria</emphasis>
</taxonomicName>
by their epigyne with its atrium restricted to the posterior third of the epigyne, covered by the rounded expansion of its anterior margin, and delimited posteriorly by large lateral lobes (
<figureCitation id="4BB624EA0D0DB74D8AC2FE28FAC4FE38" box="[1158,1295,402,429]" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25C, D</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D0DB74D8E83FE0DFBECF9C4" pageId="50" pageNumber="51" type="description">
<paragraph id="D332386F0D0DB74D8E83FE0DFD2DFE45" blockId="50.[151,1437,151,1689]" box="[199,742,438,465]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FE0DFD2DFE45" bold="true" box="[199,742,438,465]" pageId="50" pageNumber="51">
Description. Male (
<typeStatus id="0C3686CD0D0DB74D8FEFFE0DFDDEFE45" box="[427,533,438,464]" pageId="50" pageNumber="51" type="holotype">holotype</typeStatus>
, WAM T162498).
</emphasis>
</paragraph>
<paragraph id="D332386F0D0DB74D8E83FE60FB4AFD58" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FE60FE92FE60" box="[199,345,475,501]" italics="true" pageId="50" pageNumber="51">Total length,</emphasis>
4.29.
<emphasis id="E1F9E47D0D0DB74D8FEFFE60FDEBFE60" box="[427,544,475,501]" italics="true" pageId="50" pageNumber="51">Carapace,</emphasis>
length 2.68, width 1.84, dark brown, with paler median longitudinal band (
<figureCitation id="4BB624EA0D0DB74D8EDBFE44FEC1FD8C" box="[159,266,511,537]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24A</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8F5AFE44FE49FD8C" box="[286,386,511,537]" italics="true" pageId="50" pageNumber="51">Sternum,</emphasis>
orange-brown (
<figureCitation id="4BB624EA0D0DB74D8C7CFE44FD6AFD8C" box="[568,673,511,537]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24B</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8CF1FE44FD3AFD8D" box="[693,753,511,536]" italics="true" pageId="50" pageNumber="51">Eyes,</emphasis>
diameter of AME: 0.08; ALE: 0.08; PME: 0.25; PLE: 0.18, anterior eye row slightly procurved, evenly spaced.
<emphasis id="E1F9E47D0D0DB74D8CA1FD98FCACFDA8" box="[741,871,547,573]" italics="true" pageId="50" pageNumber="51">Chelicerae,</emphasis>
evenly dark brown, three retromarginal and three promarginal teeth, median largest.
<emphasis id="E1F9E47D0D0DB74D8C5FFDFCFDB3FDF4" box="[539,632,583,609]" italics="true" pageId="50" pageNumber="51">Labium,</emphasis>
dark brown, with pale anterior rim (
<figureCitation id="4BB624EA0D0DB74D8A49FDFCFBBFFDF4" box="[1037,1140,583,609]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24B</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8AC3FDFCFB08FDF5" box="[1159,1219,583,608]" italics="true" pageId="50" pageNumber="51">Legs,</emphasis>
background colour reddish-brown, mostly dark brown stained, tibiae darker (
<figureCitation id="4BB624EA0D0DB74D8D7CFDD0FC0CFD10" box="[824,967,619,645]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24A, B</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8DA4FDD0FBB0FD10" box="[992,1147,619,645]" italics="true" pageId="50" pageNumber="51">Opisthosoma,</emphasis>
length 1.65, width 1.30. Dorsum mostly dark grey except for the pale-brown cardiac mark as long as half of the opisthosoma (Fig. 224A). Venter mostly pale brown scattered with dark grey spots, spinnerets dark grey (
<figureCitation id="4BB624EA0D0DB74D8A49FD08FBBFFD58" box="[1037,1140,691,717]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24B</figureCitation>
).
</paragraph>
<paragraph id="D332386F0D0DB74D8E83FD6CFC78FC78" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FD6CFEE7FD64" box="[199,300,727,753]" italics="true" pageId="50" pageNumber="51">Pedipalp</emphasis>
(
<figureCitation id="4BB624EA0D0DB74D8F7BFD6CFE03FD64" box="[319,456,727,753]" captionStart="FIGURE 24" captionStartId="48.[151,250,1779,1803]" captionTargetBox="[157,1425,190,1755]" captionTargetId="figure-20@48.[151,1436,179,1755]" captionTargetPageId="48" captionText="FIGURE 24. Artoria pileata sp. nov., male: A, dorsal habitus; B, ventral habitus; C–E, left pedipalp (C, ventral view; D, retrolateral view; E, embolic division, ventral view). A–D, holotype (WAM T162498); E, (WAM T64059). Scale bars: A, B, 2.0 mm; C–E 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387183" httpUri="https://zenodo.org/record/14387183/files/figure.png" pageId="50" pageNumber="51">Fig. 24C–E</figureCitation>
), Tibia rectangular in ventral view, cymbium free tip almost as longer as half of the cymbium in ventral view; subtegulum heavily sclerotized and very slim in ventral view. Tegular apophysis a subrectangular stalk in ventral view, inclined retrolaterally, that slightly broads distally, and forms an inner concavity at its distal half, while its truncated, thick, and heavily sclerotized apex curves distally. Tegular apophysis has a sinuous prolateral margin with a median digitiform and poorly sclerotized small projection. Basoembolic apophysis wider than long, poorly sclerotized, small, and rounded. Embolus comma-like, tapering, almost filiform through most of its extension, ending in an acute and flexible tip. Terminal apophysis sclerotized, very broad, much wider than the embolus, with its basal portion bearing a concavity as a gutter.
</paragraph>
<paragraph id="D332386F0D0DB74D8E83FC4CFE38FB85" blockId="50.[151,1437,151,1689]" box="[199,499,1014,1041]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FC4CFE38FB85" bold="true" box="[199,499,1014,1041]" pageId="50" pageNumber="51">Female (WAM T162496).</emphasis>
</paragraph>
<paragraph id="D332386F0D0DB74D8E83FBA0FBF3FB7C" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FBA0FE9FFBA0" box="[199,340,1051,1077]" italics="true" pageId="50" pageNumber="51">Total length,</emphasis>
5.08.
<emphasis id="E1F9E47D0D0DB74D8FDFFBA0FDDBFBA0" box="[411,528,1051,1077]" italics="true" pageId="50" pageNumber="51">Carapace,</emphasis>
length 2.46, width 1.76.
<emphasis id="E1F9E47D0D0DB74D8D6FFBA0FC52FBA0" box="[811,921,1051,1077]" italics="true" pageId="50" pageNumber="51">Carapace</emphasis>
and
<emphasis id="E1F9E47D0D0DB74D8D94FBA0FBE1FBA1" box="[976,1066,1051,1076]" italics="true" pageId="50" pageNumber="51">sternum</emphasis>
colouration like male (
<figureCitation id="4BB624EA0D0DB74D8B74FBA0FF67FBCC" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25A, B</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8E80FB84FF34FBCD" box="[196,255,1087,1112]" italics="true" pageId="50" pageNumber="51">Eyes,</emphasis>
diameter of AME 0.09, ALE 0.09, PME 0.25, PLE 0.20, anterior eye row like male.
<emphasis id="E1F9E47D0D0DB74D8A9BFB84FAAAFBCC" box="[1247,1377,1087,1113]" italics="true" pageId="50" pageNumber="51">Chelicerae,</emphasis>
dark brown (as the carapace), with irregular dark grey streaks, teeth pattern as in male.
<emphasis id="E1F9E47D0D0DB74D8A0CFBD8FB6FFBE8" box="[1096,1188,1123,1149]" italics="true" pageId="50" pageNumber="51">Labium,</emphasis>
like male (
<figureCitation id="4BB624EA0D0DB74D8B63FBD8FA5BFBE8" box="[1319,1424,1122,1149]" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25B</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8ED3FB3CFF18FB35" box="[151,211,1159,1184]" italics="true" pageId="50" pageNumber="51">Legs,</emphasis>
like male (
<figureCitation id="4BB624EA0D0DB74D8F10FB3CFE15FB34" box="[340,478,1158,1185]" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25A, B</figureCitation>
).
<emphasis id="E1F9E47D0D0DB74D8FB0FB3CFD44FB34" box="[500,655,1159,1185]" italics="true" pageId="50" pageNumber="51">Opisthosoma,</emphasis>
length 2.48, width 1.71. Dorsum similar to male, but with four pairs of pale brown dots in two longitudinal lines throughout the middle of the opisthosoma following the sides of the cardiac mark; venter similar to male, but with pale brown spinnerets (
<figureCitation id="4BB624EA0D0DB74D8DE7FB74FBE0FB7C" box="[931,1067,1230,1257]" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25A, B</figureCitation>
).
</paragraph>
<paragraph id="D332386F0D0DB74D8E83FB48FB93FA70" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FB48FEE1FA99" box="[199,298,1267,1292]" italics="true" pageId="50" pageNumber="51">Epigyne,</emphasis>
wider than long; atrium small, placed in posterior half of epigynal area, mostly covered by the extension of its anterior margin, which is slightly rounded, almost reaches posterior margin of epigyne and has a contrastive black line which follows to the inner margins of lateral lobes. Median septum totally covered by the extension of anterior margin of the epigyne. Lateral lobes occupying the posterior fourth of epigynal area, projected mesally, almost parallel to epigastric furrow and touching each other (
<figureCitation id="4BB624EA0D0DB74D8D13FA38FC0BFA08" box="[855,960,1410,1437]" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25C</figureCitation>
). Spermathecal heads globular, about one diameter apart; spermathecal stalks about as long as one spermathecal diameter, attached posteriorly and following ventrally straight to the copulatory openings in the central area of epigyne (
<figureCitation id="4BB624EA0D0DB74D8DA7FA70FB80FA70" box="[995,1099,1482,1509]" captionStart="FIGURE 25" captionStartId="49.[151,250,1618,1642]" captionTargetBox="[151,1409,189,1593]" captionTargetId="figure-20@49.[151,1436,179,1593]" captionTargetPageId="49" captionText="FIGURE 25. Artoria pileata sp. nov., female: A, dorsal habitus; B, ventral habitus; C, epigyne; D, internal genitalia, dorsal view. A–C, (WAM T162496); D, (WAM T162497). Scale bars: A, B, 2.0 mm; C, D, 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.14387187" httpUri="https://zenodo.org/record/14387187/files/figure.png" pageId="50" pageNumber="51">Fig. 25D</figureCitation>
).
</paragraph>
<paragraph id="D332386F0D0DB74D8E83FA55FBDCF99C" blockId="50.[151,1437,151,1689]" box="[199,1047,1518,1545]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83FA55FEF4F99D" bold="true" box="[199,319,1518,1544]" pageId="50" pageNumber="51">Variation.</emphasis>
Body length: Males (n=5): 4.29–4.39; Females (n=5): 4.19–5.08
</paragraph>
<paragraph id="D332386F0D0DB74D8E83F9A9FBECF9C4" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83F9A9FDB7F9B9" bold="true" box="[199,636,1554,1581]" pageId="50" pageNumber="51">Life history and habitat preferences.</emphasis>
Males and females were collected in June and October, in the northern Australian dry season. There is no habitat information with any of the specimens.
</paragraph>
</subSubSection>
<subSubSection id="9B976BE40D0DB74D8E83F9E1FF0FF90C" pageId="50" pageNumber="51" type="distribution">
<paragraph id="D332386F0D0DB74D8E83F9E1FF0FF90C" blockId="50.[151,1437,151,1689]" pageId="50" pageNumber="51">
<emphasis id="E1F9E47D0D0DB74D8E83F9E1FEAAF9E1" bold="true" box="[199,353,1626,1652]" pageId="50" pageNumber="51">Distribution.</emphasis>
Only known from two locations in the Kimberley region in northern
<collectingRegion id="1149F68D0D0DB74D8AD6F9E0FA96F9E0" box="[1170,1373,1627,1653]" country="Australia" name="Western Australia" pageId="50" pageNumber="51">Western Australia</collectingRegion>
(
<figureCitation id="4BB624EA0D0DB74D8B2BF9E0FF7CF90C" captionStart="FIGURE 26" captionStartId="51.[151,250,1414,1438]" captionTargetBox="[160,1428,383,1381]" captionTargetId="figure-101@51.[151,1436,378,1389]" captionTargetPageId="51" captionText="FIGURE 26. Distribution of Artoria pileata sp. nov., Artoria pinnata sp. nov., Artoria plicata sp. nov., Artoria retorta sp. nov., and Artoria schizocoides Framenau &amp; Hebets, 2007." figureDoi="http://doi.org/10.5281/zenodo.14387191" httpUri="https://zenodo.org/record/14387191/files/figure.png" pageId="50" pageNumber="51">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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