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<document id="6FBC6AEDA9F331BEDB6040A40E61177B" ENCODING="UTF-8" ID-GBIF-Dataset="d5fd4c73-69cc-419e-ae2e-811f29efd1bc" ModsDocID="z00791p001" checkinTime="1246965505869" checkinUser="thomas" docAuthor="Victor G. Springer &amp; Gerald R. Allen" docDate="2004" docId="58FA14A1CBE7B5745A7186182860937D" docLanguage="en" docName="2004_Springer_Allen_gg1.xml" docOrigin="Zootaxa 791" docSource="http://www.zoobank.org/urn:lsid:zoobank.org:pub:87531ECD-559C-4BFB-98B8-29828E58A3A6" docTitle="Ecsenius shirleyae Springer &amp; Allen, 2004, new species" docType="treatment" docUuid="DE3BA438-05EF-49DC-A1C8-3BC6A77D7F00" docUuidSource="ZooBank" docVersion="15" lastPageNumber="9" masterDocId="EF1D1312A59C6883FD90C25C51F795A2" masterDocTitle="Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius." masterLastPageNumber="12" masterPageNumber="1" pageNumber="8" updateTime="1701314078103" updateUser="plazi">
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<mods:title id="AD9C79839DED481F1CDBC9ECC0607E9C">Ecsenius caeruliventris and E. shirleyae, two new species of blenniid fishes from Indonesia, and new distribution records for other species of Ecsenius.</mods:title>
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<mods:namePart id="FB33BAE43CA626A8DEAD08461B385F99">Victor G. Springer</mods:namePart>
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<mods:namePart id="1D453FD2AF8BCB644F3C36BE1990AEDC">Gerald R. Allen</mods:namePart>
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<mods:date id="C19F1C432C275A61769F25763AA27B41">2004</mods:date>
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<treatment id="58FA14A1CBE7B5745A7186182860937D" ID-DOI="http://doi.org/10.5281/zenodo.6271248" ID-GBIF-Taxon="100122825" ID-Zenodo-Dep="6271248" LSID="urn:lsid:zoobank.org:act:DE3BA438-05EF-49DC-A1C8-3BC6A77D7F00" httpUri="http://treatment.plazi.org/id/58FA14A1CBE7B5745A7186182860937D" lastPageNumber="9" pageNumber="8">
<subSubSection id="C0342BB4B17F3EC902D14B9ECA05CD40" type="nomenclature">
<paragraph id="DDF4D8EF52FA5E4244298700341D6444" pageNumber="8">
<taxonomicName id="2DDE183D4F163FA5CC759A3306B0FE0F" ID-CoL="38P23" LSID-ZBK="urn:lsid:zoobank.org:act:DE3BA438-05EF-49DC-A1C8-3BC6A77D7F00" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius shirleyae Springer &amp; Allen 2004:8" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="shirleyae">Ecsenius shirleyae</taxonomicName>
,
<taxonomicNameLabel id="552CFDC9A2AC63066B299FF99FEF9772" rank="species">new species</taxonomicNameLabel>
</paragraph>
<paragraph id="2AF3E45C6C1BFF53FD18F86A22DBBDDE" pageNumber="8">Shirleys blenny</paragraph>
</subSubSection>
<subSubSection id="A0E1334FA26A61B853414B93B4210C13" type="description">
<paragraph id="9169C919151926D4B9A2AF25E224671A" pageNumber="8">Figures 1d, 2b, 3</paragraph>
</subSubSection>
<subSubSection id="B86B576449AC6ABD02EDDC3C6D43B89C" type="reference_group">
<paragraph id="5B463DC30BE84CE13E3FA23BE5502385" pageNumber="8">
<taxonomicName id="6568AFEFC5BF64B77BC003B8D135F002" LSID-ZBK="urn:lsid:zoobank.org:act:76CFCEFC-20FB-4FD7-8B70-ACEF9019DF09" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius bandanus Springer 1971:38" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="bandanus">Ecsenius bandanus</taxonomicName>
(not Springer), in part: Springer, 1988:106, fig. 57, pl.13, fig. 5; Kuiter &amp; Tonozuka, 2001:606 (Maumere, Flores); Allen &amp; Adrin, 2003:54 &amp; 18,
<materialsCitation id="AA6E2AFEBBA374329E0E7528F7B114F1" ID-GBIF-Occurrence="923906145" country="Indonesia" stateProvince="Flores">
fig. 65 (Maumere,
<collectingRegion id="3F7899BCD27F05FF19FFEC27BA5E7B76">Flores</collectingRegion>
)
</materialsCitation>
; Myers, 1999:224 &amp; pl. 146K.
</paragraph>
</subSubSection>
<subSubSection id="DCC931B3FA02D756F94ED3C6D9B7CDF0" type="nomenclature">
<paragraph id="DDE4CA104C8B656AAFCE9F9F912B3BD7" pageNumber="8">
<materialsCitation id="B05F570C1550324B6188B38C1A1A1C84" ID-GBIF-Occurrence="923906149" country="Indonesia" stateProvince="Pulau Seribu">
<typeStatus id="EEEBDDEB338A8EA45077EEE263516AA1">Holotype</typeStatus>
:
<collectionCode id="ED1FCF39CE693C704DF2C892FA3714CF" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
211996, male, 28.4 mm SL, off NW side of Pulau Tikus, Pulau Pari group,
<collectingRegion id="02F673D70F8C9C380AA3C437374AA061">Pulau Seribu</collectingRegion>
,
<collectingCountry id="00264A471389E0ECA6479F8659FEB11B">Indonesia</collectingCountry>
(ca.
<geoCoordinate id="9B7AA981119E15CD898E4EF56B811C12" direction="south" orientation="latitude" value="-5.8569446">05°5125&quot; S</geoCoordinate>
, ca.
<geoCoordinate id="396BAE168717B790B7B370301FFEFFA9" direction="east" orientation="longitude" value="106.57083">106°3415&quot; E</geoCoordinate>
), to ca. 5 m, rotenone, V.G. Springer, M.F. Gomon, &amp; Sukarno,
<date id="D41A563ED51FD58ACAD5412B51E45890" day="05" month="04" value="1974-04-05" year="1974">5 April 1974</date>
; VGS 74-34.
</materialsCitation>
</paragraph>
<paragraph id="E4973CC66BA11F292B1C74A81E71DC2F" pageNumber="8">
<materialsCitation id="E955604B0704D160F5477129AC6AFF01" ID-GBIF-Occurrence="923906154" country="Indonesia" stateProvince="Pulau Seribu">
<typeStatus id="6841629C45B7C48CFA09459F070D5223">Paratypes</typeStatus>
:
<collectionCode id="06FB59E55A90274533D44E393F1D92B2" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
379409 (15:15.6-33.5)
</materialsCitation>
and
<materialsCitation id="F8BD83FE910563A96F7338F95A305A53" ID-GBIF-Occurrence="923906155" country="Indonesia" stateProvince="Pulau Seribu">
<collectionCode id="BD63DA10D82FF89AD80BC8A860298FBD" collectionName="Australia, Western Australia, Perth, Western Australian Museum">WAM</collectionCode>
P32602.001 (2: 22.2-29.1), collected with holotype
</materialsCitation>
.
<materialsCitation id="9BA30266A1504297419406D29CFE26B9" ID-GBIF-Occurrence="923906143" country="Indonesia" stateProvince="Celebes">
<collectionCode id="F631C028A2A1E339648D36EE75900F21" collectionName="Australia, New South Wales, Sydney, Australian Museum">AMS</collectionCode>
I.18490001 (10:17.3-28.6), Tallabassi Bay, just off NW shore of Big Damalawa Islet, Kabaena Island,
<collectingRegion id="6ECEB1DDA93D33B70AA7672AF465155C">Celebes</collectingRegion>
,
<collectingCountry id="FC5CC6D0A0FC2B397B0C0831D5270A9A">Indonesia</collectingCountry>
(ca.
<geoCoordinate id="A62CE493E2F77E1CF9884F45259F1C2B" direction="south" orientation="latitude" value="-5.288889">05°1720&quot; S</geoCoordinate>
, ca.
<geoCoordinate id="9A238B72345C73606BD436F15C534EC1" direction="east" orientation="longitude" value="122.066666">122°04 E</geoCoordinate>
), 4-8 m, rotenone, V.G. Springer, M.F. Gomon, &amp; Indonesians,
<date id="5C76C09964122700370147E6947A9124" day="24" month="02" value="1974-02-24" year="1974">24 February 1974</date>
, VGS 74-1
</materialsCitation>
.
<materialsCitation id="860ACFB8FC89531949DDE602426C8F63" ID-GBIF-Occurrence="923906151" country="Indonesia" stateProvince="Celebes">
<collectionCode id="7CFCBBB572531B21F607322BD03AD013" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
211916 (14:22.5-29.2, includes one specimen now lost), Tallabassi Bay, just off NE tip of Big Damalawa Islet, Kabaena Island,
<collectingRegion id="68A56AE951E4DD954159C3D706311779">Celebes</collectingRegion>
,
<collectingCountry id="5D4336D319B21AC028C1B4D7DF60D6CD">Indonesia</collectingCountry>
(ca.
<geoCoordinate id="C98B26D2D42092BA75B5E376DC672F5B" direction="south" orientation="latitude" value="-5.288889">05°1720&quot; S</geoCoordinate>
, ca.
<geoCoordinate id="DA0113299F1BA2EE75A7B3E5CD8772D3" direction="east" orientation="longitude" value="122.066666">122°04 E</geoCoordinate>
), 2-15 m, rotenone, V G Springer, M. F. Gomon, &amp; Indonesians,
<date id="9F179E10897E52F31A75429964009DD4" day="25" month="02" value="1974-02-25" year="1974">25 February 1974</date>
, VGS 74-2
</materialsCitation>
.
<materialsCitation id="44431BF0581410E20E782897389D237F" ID-GBIF-Occurrence="923906157" country="Indonesia" stateProvince="Pulau Seribu">
<collectionCode id="1F6E913E7549A79CAA9440547EEA6D89" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
211993 (18:13.3-29.2), off W side of Pulau Ajer,
<collectingRegion id="D24CAF085A4475E20E57225C8A01FDCA">Pulau Seribu</collectingRegion>
,
<collectingCountry id="927D1CBEFE7A58108439A386EC81F34A">Indonesia</collectingCountry>
(ca.
<geoCoordinate id="FAA88D4137B4D7055D07B3D31379D28A" direction="south" orientation="latitude" value="-5.7666664">05°46 S</geoCoordinate>
, ca.
<geoCoordinate id="2BCED226FDBD85F5B11679CB61EBEC90" direction="east" orientation="longitude" value="106.583336">106°35 E</geoCoordinate>
), to ca.19 m, rotenone, V.G. Springer, M.F. Gomon, &amp; Sukarno,
<date id="32540BF7E3944299C294C8C6E9BB7BD8" day="04" month="04" value="1974-04-04" year="1974">4 April 1974</date>
; VGS 74-32
</materialsCitation>
.
<materialsCitation id="8DD8E00E55F1786CC139FBD56BC214DE" ID-GBIF-Occurrence="923906164" country="Indonesia" stateProvince="Java">
<collectionCode id="CEDEBC52343B9DC83156C95700640A1D" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]">USNM</collectionCode>
211979 (11:23.7-31.2), off SW coast Karimundjawa Island,
<collectingCountry id="0ED45C4F6123157293531F139CC98FED">Indonesia</collectingCountry>
, ca.
<geoCoordinate id="CE5F17D608DC3E9B9802D6CE1989413F" direction="south" orientation="latitude" value="-5.875">05°5230” S</geoCoordinate>
,
<geoCoordinate id="05CEC6A81EDB7A8A2F3CE39C25F356C0" direction="east" orientation="longitude" value="110.42777">110°2540” E</geoCoordinate>
, ca. 1-12 m, rotenone, V.G. Springer &amp; Indonesians,
<date id="1CF9489425974A689656591FE3073271" day="29" month="03" value="1974-03-29" year="1974">29 March 1974</date>
, VGS 74-28
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="AED6FB7926230622317CBA5C8BFE43C3" type="materials_examined">
<paragraph id="BB837E5E1C617F8D74154AD478BD4B60" pageNumber="8">
Additional specimen:
<materialsCitation id="DC0EC161DDDDE5292E17057012FE035D" ID-GBIF-Occurrence="923906160" country="Indonesia" stateProvince="Flores">
<collectionCode id="86306288FB6A1C16A7D85B0E63681837" collectionName="Australia, New South Wales, Sydney, Australian Museum">AMS</collectionCode>
I.34501010 (1:15.6), off Wailiti, 11 km NW Sao Wisata Resort,
<collectingRegion id="CD0DA4236BE833FD1F1620C365CE93AB">Flores</collectingRegion>
,
<collectingCountry id="4EC1250D316D80A38BCD6901637BA0D6">Indonesia</collectingCountry>
,
<geoCoordinate id="9871F15082240DB89B4ABBEBA606EB1A" direction="south" orientation="latitude" value="-8.571667">08°34.3 S</geoCoordinate>
,
<geoCoordinate id="9F1C2D7BAC4B01A3652513EFF017EBC2" direction="east" orientation="longitude" value="122.195">122°11.7 E</geoCoordinate>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="825C089A925EA78B4F346563309A9E01" type="diagnosis">
<paragraph id="CFAA9A1F28C285A8EFE733FF29F077B1" pageNumber="8">
Diagnosis. A species of the
<taxonomicName id="8881D9C77C280C7931E00237571EA659" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman &amp; Schultz 1952:519" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
group of
<taxonomicName id="54ABAD83F42C41F00957B90AA63D5CC9" LSID-ZBK="urn:lsid:zoobank.org:act:B3F80107-BABB-495D-BFB7-721DCA10C4F8" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius McCulloch 1923:121" order="Perciformes" pageNumber="8" phylum="Chordata" rank="genus">Ecsenius</taxonomicName>
with the following combination of characters: Body without alternating dark and pale stripes or distinct dark spots; head with dark postorbital stripe extending across head onto body anteriorly. In life, dark postorbital stripe bordered ventrally by bright white stripe; other bright markings on head white or pale yellowish; belly bluish.
</paragraph>
</subSubSection>
<subSubSection id="B44C51D511B40480CF495F8B15A689A5" type="description">
<paragraph id="FBF07EDB16E2A31A4EA04796249E07A9" lastPageNumber="9" pageNumber="8">
Description (only differentiating characters or characters not otherwise mentioned in the introduction, identification key, and Table 1 are discussed). Preserved specimens (Figure1d) are most notable for the black stripe extending across the head posteriorly from the postorbital margin at about the three oclock position. The stripe may extend for a variable, usually short, distance, onto the body anteriorly. On each side of the ventral surface of the head of the male holotype there are two pairs of dark spots similar to those of
<taxonomicName id="BC1327EEDDC5A4F9B757139B1540A440" LSID-ZBK="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius caeruliventris Springer &amp; Allen 2004:3" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="caeruliventris">E. caeruliventris</taxonomicName>
, although the halo is not evident (presence or absence of the spots varies in other males). The body is not so darkly dusky anteriorly as in
<taxonomicName id="5920A2D8F2B75E3FDC8E027AB497396F" LSID-ZBK="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius caeruliventris Springer &amp; Allen 2004:3" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="caeruliventris">E. caeruliventris</taxonomicName>
. The fins are similar to those of
<taxonomicName id="D8DC439847702E6E62962E769DE95811" LSID-ZBK="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius caeruliventris Springer &amp; Allen 2004:3" order="Perciformes" pageNumber="8" phylum="Chordata" rank="species" species="caeruliventris">E. caeruliventris</taxonomicName>
, except that an immaculate stripe-like basal area in the segmented-ray portion of the dorsal fin is not evident. Females are generally more pale than males, and the dark spots on the ventral head surface are absent or replaced by one or two pairs of larger, pale spots (Figure 1c) in the positions occupied by the dark spots of the male.
</paragraph>
<paragraph id="2A4EB9C113D585ADC1EBAC3A2E3F3F6F" pageNumber="9">
In life (Figure 2b), the most obvious color markings are the bluish abdominal area, a horizontal pair of white stripes extending through the orbit dorsal and ventral to the pupil, a pair of small white spots in the interorbital region anteriorly, a fine, white mid-predorsal stripe (all of these white stripes may be very faintly tinged with yellow), the black stripe extending posteriorly from the orbit margined ventrally by a bright white stripe, which continues well out onto the body. The head and body are brownish dorsal to the black stripe, and may show indications of a few broad, slightly darker bands anteriorly; posteriorly the body is similar to
<taxonomicName id="6D10BC69B96A0DEA4428B6F49D151F23" LSID-ZBK="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius caeruliventris Springer &amp; Allen 2004:3" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="caeruliventris">E. caeruliventris</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="FD21D1ADBE8C94C55FE85682EF4CE051" type="discussion">
<paragraph id="481F3ECA364F935A18A72CB3B93EF33D" pageNumber="9">
Comparisons. Within the
<taxonomicName id="B0FE6DC32A1970D64787569FE51BDE25" LSID-ZBK="urn:lsid:zoobank.org:act:669CE437-D461-4DF0-AF57-F94B3740CCAE" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius prooculis Chapman &amp; Schultz 1952:519" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="prooculis">Prooculis</taxonomicName>
group,
<taxonomicName id="072D55A9D9EDDCE5F2168F0FF2688A80" LSID-ZBK="urn:lsid:zoobank.org:act:DE3BA438-05EF-49DC-A1C8-3BC6A77D7F00" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius shirleyae Springer &amp; Allen 2004:8" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="shirleyae">Ecsenius shirleyae</taxonomicName>
appears to be most similar to
<taxonomicName id="5633B269E86FDEF41402FEFC63BC8D82" LSID-ZBK="urn:lsid:zoobank.org:act:C6554AD6-8A16-41DD-B379-168B0416A3FB" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius bimaculatus Springer 1971:38" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="bimaculatus">E. bimaculatus Springer</taxonomicName>
, from which it differs mainly in lacking the two black abdominal spots and in that some individuals have a bluish abdomen. It is similar to
<taxonomicName id="12BD1938C792774480C4691E4FAFE900" LSID-ZBK="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius caeruliventris Springer &amp; Allen 2004:3" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="caeruliventris">E. caeruliventris</taxonomicName>
and
<taxonomicName id="EA20368AFB05EED4C23B9D9749FB019E" LSID-ZBK="urn:lsid:zoobank.org:act:76CFCEFC-20FB-4FD7-8B70-ACEF9019DF09" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius bandanus Springer 1971:38" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="bandanus">E. bandanus</taxonomicName>
in having a bluish abdomen (at least in some specimens), in having broad, faintly dusky bands on the body, and in lacking distinct dark spots or stripes on the entire body. It differs from both species in having a dark postorbital stripe that frequently extends well out onto the body and in having the stripe margined below by a white stripe that also extends well out onto the body. It further differs from
<taxonomicName id="B955C2A5054E331F193E62A819D2C75A" LSID-ZBK="urn:lsid:zoobank.org:act:6515A914-D516-4C35-959F-8310DAE39262" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius caeruliventris Springer &amp; Allen 2004:3" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="caeruliventris">E. caeruliventris</taxonomicName>
in that the blue of the abdomen is not as strongly manifested, and from
<taxonomicName id="705FF9C6149BDAA705223F80F2694D66" LSID-ZBK="urn:lsid:zoobank.org:act:76CFCEFC-20FB-4FD7-8B70-ACEF9019DF09" family="Blenniidae" genus="Ecsenius" higherTaxonomySource="CoL" kingdom="Animalia" lsidName-ZBK="Ecsenius Ecsenius bandanus Springer 1971:38" order="Perciformes" pageNumber="9" phylum="Chordata" rank="species" species="bandanus">E. bandanus</taxonomicName>
in that the pale markings on the head and eye are bright white (or very faintly tinged with yellow) rather than bright yellow, and that the head portion of the dark post-orbital stripe is generally less deep (depth measurement is difficult because margins of the stripe are not sharply demarcated, but the difference is usually apparent when specimens of equal size are compared).
</paragraph>
</subSubSection>
<subSubSection id="E72FA4D5C445AF63F731D62022B74E54" type="etymology">
<paragraph id="E8F4FFB38808DBF1171DB9507167599C" pageNumber="9">Etymology: This species is named for my (VGS) wife, Shirley, in recognition of her unstinting support for my research for the past almost 40 years.</paragraph>
</subSubSection>
</treatment>
</document>