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<document ID-CLB-Dataset="4837" ID-DOI="http://dx.doi.org/10.3897/evolsyst.5.63435" ID-GBIF-Dataset="bfba5087-74d4-44e8-b783-94db25e3442a" ID-Pensoft-Pub="2535-0730-2-193" ID-Pensoft-UUID="21773559522D5D9DA4B1A7DA51E4F2A6" ID-ZooBank="984E15D880E04B7DA84F92BB0AD4EA73" ModsDocID="2535-0730-5-2-193" checkinTime="1629846914602" checkinUser="pensoft" docAuthor="Zahiri, Reza, Nazari, Vazrick, Rajaei, Hossein, Wiemers, Martin, Fatahi, Maryam, Seidel, Matthias, Dalsgaard, Thure &amp; Husemann, Martin" docDate="2021" docId="581571755CD957079CB8CEF94505FA85" docLanguage="en" docName="EvolutSyst 5(2): 193-261" docOrigin="Evolutionary Systematics 5 (2)" docPubDate="2021-08-20" docSource="http://dx.doi.org/10.3897/evolsyst.5.63435" docTitle="Melitaea varissima Verity 1914" docType="treatment" docVersion="5" id="21773559522D5D9DA4B1A7DA51E4F2A6" lastPageNumber="193" masterDocId="21773559522D5D9DA4B1A7DA51E4F2A6" masterDocTitle="? An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea" masterLastPageNumber="261" masterPageNumber="193" pageNumber="193" updateTime="1732934557711" updateUser="ExternalLinkService">
<mods:mods id="98EC319CD9742D54D8ECA2A22FC7FFB1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="49325CB9AC932FD69A63A28902011422">? An illustrated catalogue of the type specimens of Lepidoptera housed in the Zoological Museum Hamburg (ZMH): Part II. superfamily Papilionoidea</mods:title>
</mods:titleInfo>
<mods:name id="5BFF0A977DC74B7868755836CB6E1799" type="personal">
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<mods:namePart id="97521C2DDD6F3AA63D869B8AC71A015F">Zahiri, Reza</mods:namePart>
<mods:nameIdentifier id="DB05DB16F5CCE53122F48F3660D5430E" type="ORCID">https://orcid.org/0000-0001-6274-6973</mods:nameIdentifier>
<mods:affiliation id="26ED80EAE6BD259C8445010623020BE8">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany &amp; Canadian Food Inspection Agency, Ottawa Plant Laboratory, Entomology Unit, Bldg. 18, 960 Carling Ave., K 1 A 0 C 6, Ottawa, Ontario, Canada</mods:affiliation>
<mods:nameIdentifier id="3A34A8ECBB58237C45AD9FD03940ADE1" type="email">reza.zahiri@gmail.com</mods:nameIdentifier>
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<mods:namePart id="CECFD6E1B597FD3721E2D78810B3F23F">Nazari, Vazrick</mods:namePart>
<mods:nameIdentifier id="D0D62679C277AEB50556FE7F0DBDC893" type="ORCID">https://orcid.org/0000-0001-9064-8959</mods:nameIdentifier>
<mods:affiliation id="3D6C51F1C0FB57A1FE4B144CDB289499">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
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<mods:roleTerm id="40A26B2A601EA8E663C352D1192280E4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D0C4210127EB5CA970F78B1EC41D86CD">Rajaei, Hossein</mods:namePart>
<mods:nameIdentifier id="DD2BB4E035ADC6F0E23CE37EA117D490" type="ORCID">https://orcid.org/0000-0002-3940-3734</mods:nameIdentifier>
<mods:affiliation id="6FEA77A4C248CCA1EF95CA3544F3C037">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
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<mods:namePart id="6D90BDC4F9517D32BAA7571159E2A08D">Wiemers, Martin</mods:namePart>
<mods:nameIdentifier id="7F39E3AB609ED9EF6624CC77167A3700" type="ORCID">https://orcid.org/0000-0001-5272-3903</mods:nameIdentifier>
<mods:affiliation id="9167CDF52709A29CC410C098D45FB223">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:name id="8CB77D581F5563BE07C3FD2025232911" type="personal">
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<mods:roleTerm id="59988640D310DC801707D39508D7BA68">Author</mods:roleTerm>
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<mods:namePart id="9A9508CE017301AB9B3120E94D099B60">Fatahi, Maryam</mods:namePart>
<mods:affiliation id="BF56F7397060257796B7E5508F60F1FD">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="EA9E7E244DF05AF4F576B5B1BFADB589">Seidel, Matthias</mods:namePart>
<mods:nameIdentifier id="5A6D8AEBE49A6B23C3679E093ABA0E13" type="ORCID">https://orcid.org/0000-0002-4913-8778</mods:nameIdentifier>
<mods:affiliation id="51F7B0FE88A1E4DD64568C9F69563842">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
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<mods:roleTerm id="9739E18FE30E33D3002C9B9E35B6CBA2">Author</mods:roleTerm>
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<mods:namePart id="756BAB7E1549DA704AD46CA44B831918">Dalsgaard, Thure</mods:namePart>
<mods:affiliation id="A8FF41DED8DCB988C85EF9639313CB25">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
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</mods:role>
<mods:namePart id="B673DD3B05CAE9161B1A5EBD39A78EEA">Husemann, Martin</mods:namePart>
<mods:nameIdentifier id="728B0709A9F562E33EEB05595215D4A0" type="ORCID">https://orcid.org/0000-0001-5536-6681</mods:nameIdentifier>
<mods:affiliation id="962EB5384E084AFCF6E6927623F7E675">Leibniz Institute for the Analysis of Biodiversity Change, Martin-Luther-King Platz 3, 20146 Hamburg, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="7925309F1708FCF32DACC088004BC282">text</mods:typeOfResource>
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<mods:titleInfo id="7D90B74CC8BC0664C3C4B8D887823C75">
<mods:title id="3CB6539BD4B0DE8DB9D82C0D3A9286D6">Evolutionary Systematics</mods:title>
</mods:titleInfo>
<mods:part id="097225CC9170FCFB392FC9B305A7C1C3">
<mods:date id="B619732F77388CCBC1D820866CCC82D3">2021</mods:date>
<mods:detail id="61E78A636DDF02F42AA50E085F37C593" type="pubDate">
<mods:number id="64CA38AD726DB1880B0D44B133168ED1">2021-08-20</mods:number>
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<mods:detail id="F72B5FF622BC6A8273DA08CE65FCA9BD" type="volume">
<mods:number id="EC38823511A409C6EBE443155433FD8E">5</mods:number>
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<mods:detail id="5C812D47BFA85E12613441F5665D5EA9" type="issue">
<mods:number id="4DECD07EF52650FBA237C51AEA316A70">2</mods:number>
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<mods:start id="ED19332C2B846359DD8E5870CDF9ADCF">193</mods:start>
<mods:end id="044783E2E5CA8EF183F5CC42B0828F6E">261</mods:end>
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<mods:url id="2A23F917AE010B4831797BEB5007CDD0">http://dx.doi.org/10.3897/evolsyst.5.63435</mods:url>
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<mods:classification id="E44C06412027DB95802DD8C5C8A8B538">journal article</mods:classification>
<mods:identifier id="601D0752777BBB8E0AF91F91CD5634D7" type="DOI">http://dx.doi.org/10.3897/evolsyst.5.63435</mods:identifier>
<mods:identifier id="8AF72E5FE65A95CC00C2DD7854B23825" type="Pensoft-Pub">2535-0730-2-193</mods:identifier>
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<treatment id="581571755CD957079CB8CEF94505FA85" ID-GBIF-Taxon="184941060" LSID="urn:lsid:plazi:treatment:581571755CD957079CB8CEF94505FA85" httpUri="http://treatment.plazi.org/id/581571755CD957079CB8CEF94505FA85" lastPageNumber="193" pageId="0" pageNumber="193" scope_family="Nymphalidae" scope_order="Lepidoptera" scope_subFamily="Nymphalinae" scope_superFamily="Papilionoidea" scope_tribe="Melitaeini">
<subSubSection id="53D5A45B03AAE3B6A35B338ADF5EE96D" pageId="0" pageNumber="193" type="nomenclature">
<paragraph id="96560D7D525CD80F5E3F2E5932001E6C" pageId="0" pageNumber="193">
?111.
<taxonomicName id="53D6F2EEA1A4531F995D18B338EA1421" LSID="58157175-5CD9-5707-9CB8-CEF94505FA85" authority="Verity, 1914" authorityName="Verity" authorityYear="1914" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea varissima" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="varissima">Melitaea varissima Verity, 1914</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="81F6D13D26DFFAEF24A84BB4C251C704" pageId="0" pageNumber="193" type="original combination">
<paragraph id="DCA0A9C2FC0944C0B448079711F4BC0E" pageId="0" pageNumber="193">Original combination.</paragraph>
<paragraph id="CDA04002C4955B8E99A6B3FD5B1C140A" pageId="0" pageNumber="193">
&quot;
<taxonomicName id="051CCB6564E6DDC9D93832C2EA806A71" authorityName="ab. mediofasciata Bubacek" authorityYear="1926" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="genus">
<emphasis id="6D1DF8F8FEFF6B24E520CA4B4424FB5D" italics="true" pageId="0" pageNumber="193">Melitaea</emphasis>
</taxonomicName>
[
<emphasis id="21187373355F88CE992705EFA13D93F7" italics="true" pageId="0" pageNumber="193">parthenie</emphasis>
Bkh]
<taxonomicName id="C508FE390F73017ED716735B1C69F344" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea varia" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="9ACF5F9248EE439D752EE21F4D969F21" italics="true" pageId="0" pageNumber="193">Melitaea varia</emphasis>
</taxonomicName>
<normalizedToken id="39DAF880BB49A69F4110945E7129EF68" originalValue="Meyer-Dür">Meyer-Duer</normalizedToken>
<emphasis id="99D6ECE336E938AA16A3510E1039CCCE" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
nom. nov.&quot; Verity, 1914 (1913) Boll. Soc. Ent. Ital. 45: 210.
</paragraph>
</subSubSection>
<subSubSection id="A2F14461F179DB95F712B7073B3A20BF" pageId="0" pageNumber="193" type="current combination">
<paragraph id="7ACC199AAF0A92D21B400A71D0E9A4C3" pageId="0" pageNumber="193">Current combination.</paragraph>
<paragraph id="227D98E2A6DF3004B98F07A70D540171" pageId="0" pageNumber="193">
<emphasis id="1A6DAD06575BA7691D85C7419CEB7379" bold="true" pageId="0" pageNumber="193">
<taxonomicName id="4A13D24E7ABA1224D069B61D2CD622D3" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea varia subsp. varissima" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="subSpecies" species="varia" subSpecies="varissima">
<emphasis id="CCABFA8BDD50046A9B8EDB8C9CF8208D" italics="true" pageId="0" pageNumber="193">Melitaea varia varissima</emphasis>
</taxonomicName>
Verity, 1914
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="054401AE79A1379CEC2706676684FBD0" pageId="0" pageNumber="193" type="current status">
<paragraph id="AE68E3CFAB7D68AB82A9B08F7BA9D290" pageId="0" pageNumber="193">Current status.</paragraph>
<paragraph id="3B2964C965565FD9D88355A7A6256CF2" pageId="0" pageNumber="193">Valid subspecies.</paragraph>
</subSubSection>
<subSubSection id="D348A7F9BF96300CFE69CE70EE10A8B2" pageId="0" pageNumber="193" type="materials_examined">
<paragraph id="2FC8075F528CD17852DFFC86406BB993" pageId="0" pageNumber="193">Original material.</paragraph>
<paragraph id="48711A1DB4D54EC4AE9DE0A4B674886E" pageId="0" pageNumber="193">
<materialsCitation id="AD1FBD5DC5C525A3E9E7AF9CA9306B8E" collectingDate="1937-06-05" collectingDateMax="1937-06-25" collectingDateMin="1937-06-05" specimenCount="1" typeStatus="Type">
Labelled as
<normalizedToken id="2A2FEA1413CEDDCCE9503250D29A40BB" originalValue="“Type”">
&quot;
<typeStatus id="F7CADB779CF535324E61D0E64C6CDA61">Type</typeStatus>
&quot;
</normalizedToken>
6?? (ZMH 833461-833466) (Fig.
<figureCitation id="E41F9C1162DA88F4E8D249F31DA384E8" captionStart="Plate 16" captionStartId="F16" captionText="Plate 16. 110. Melitaea athalia tenuicula Verity, 1919; 111. Melitaea varia varissima Verity, 1914; 112. Melitaea cinxia riloensis Zuellich, 1936; 113. Melitaea deione ab. mediofasciata Bubacek, 1926; 114. Melitaea diamina ab. johni Stauder, 1922; 115. Melitaea trivia balcanica Zuellich, 1936; 116. Melitaea trivia catapelioides Stauder, 1918; 117. Melitaea trivia ignasiti de Sagarra, 1926. a. Dorsal view, b. ventral view." httpUri="https://binary.pensoft.net/fig/579725" pageId="0" pageNumber="193">111</figureCitation>
). &quot;
<emphasis id="126894837B518E5B6776297FA2F12822" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;
<taxonomicName id="2A4826187425071818F325286F591112" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea varia" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="varia">Melitaea varia</taxonomicName>
<normalizedToken id="2EA7ED19470D60F080CCDF2F2069F1A0" originalValue="Meyer-Dür”">Meyer-Duer&quot;</normalizedToken>
// &quot;Marche / Bolognola /
<collectingDate id="BAA205858C51A0A955E18BCE8E7C2CBB" value="1937-06-25">25.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833461&quot;; &quot;
<emphasis id="6C4887049D0C26746815FC6FFCA1585D" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate id="84B7ECCD7A9DB2946D7EF27883C63831" value="1937-06-20">20.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833462&quot;; &quot;
<emphasis id="27A96E175382F5C78D5677FDF7A24949" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate id="9804C483CE5041E65BE0801801E18868" value="1937-06-14">14.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833462&quot;; &quot;
<emphasis id="055D0DAE3892E97F57E8EC1409032D19" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate id="F04289CCBE7EDD812B96145E4491D384" value="1937-06-14">14.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833463&quot;; &quot;
<emphasis id="47EB85E9E933B414AE4E8BA7F899FB6A" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate id="7B1DCD1D4C35E80DF9C3CD15A677C0C9" value="1937-06-05">5.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833464&quot;; &quot;
<emphasis id="34B01A3E8C76578681B06F16EF1CD39E" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate id="179CD0B2BA6A6EED519CCE8E5B55C654" value="1937-06-22">22.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833465&quot;; &quot;
<emphasis id="2704C88EB7A9D678992A684B6748BB16" italics="true" pageId="0" pageNumber="193">varissima</emphasis>
Vrty&quot; // &quot;Marche / Bolognola /
<collectingDate id="B893960C64231DA6B8956F1639D66835" value="1937-06-25">25.VI.1937</collectingDate>
/ Querci&quot; // [blank label] // &quot;ZMH 833466
</materialsCitation>
&quot;.
</paragraph>
</subSubSection>
<subSubSection id="E63CD48606A8D2A456EACF6F6FBA3802" pageId="0" pageNumber="193" type="original locality">
<paragraph id="06B3A3FCCC3911B148E67E03D3E00FEB" pageId="0" pageNumber="193">Original locality.</paragraph>
<paragraph id="2B5E6D2B2D623FA8F85B567465B47354" pageId="0" pageNumber="193">
Italy: Bologna &quot;Nei Faggeti di Bologna, nella
<normalizedToken id="4987C09B448B9025BE002C363AFEFF59" originalValue="località">localita</normalizedToken>
Costarotara e a Piano Astore dei Monti Sibillini, da 1200 a 1600 m.
<normalizedToken id="F46BFD53C9012B864A3F1BBEF4F25DDA" originalValue="d’altitudine”">d'altitudine&quot;</normalizedToken>
[Bologna, vicinity of Costarotara and Piano Astore of the Sibillini Mountains, 1200-1600 m.].
</paragraph>
</subSubSection>
<subSubSection id="72A2A416921E2C8CF45BA6836D27EA1D" pageId="0" pageNumber="193" type="remarks">
<paragraph id="3667C87594EB30D6850F6B0335606675" pageId="0" pageNumber="193">Remarks.</paragraph>
<paragraph id="8CA031DE81F75ADBCE88FBD2CB48FE9F" pageId="0" pageNumber="193">
<bibRefCitation id="A2C5BC27060C85516B2EEF0A43FCEFC8" author="Verity, R" journalOrPublisher="Entomologist's Record" pageId="0" pageNumber="193" pagination="193 - 201" refId="B97" refString="Verity, R, 1919. Seasonal Polymorphism and Races of Some European Grypocera and Rhopalocera. Entomologist's Record 31: 193 - 201" title="Seasonal Polymorphism and Races of Some European Grypocera and Rhopalocera." volume="31" year="1919">Verity (1919)</bibRefCitation>
proposed this name as a
<normalizedToken id="03CA8928E0943E9F8D1BD8DEE07857F3" originalValue="“race”">&quot;race&quot;</normalizedToken>
of
<taxonomicName id="56BEA94F6A3AEE22D7A4353E5C0D3CD2" class="Insecta" family="Nymphalidae" genus="Melitaea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Melitaea varia" order="Lepidoptera" pageId="0" pageNumber="193" phylum="Arthropoda" rank="species" species="varia">
<emphasis id="23F4C226E46E7D8EAF3412CD4D19B6AA" italics="true" pageId="0" pageNumber="193">Melitaea varia</emphasis>
</taxonomicName>
<normalizedToken id="319C92056258C64CE4D46C8DA1E3A565" originalValue="Meyer-Dür">Meyer-Duer</normalizedToken>
, 1851. According to
<bibRefCitation id="2C8EA57E32B311FD90FF60A2B1282894" author="Kudrna, O" journalOrPublisher="Journal of Research on the Lepidoptera" pageId="0" pageNumber="193" pagination="1 - 105" refId="B56" refString="Kudrna, O, 1983. An annotated catalogue of the butterflies named by Roger Verity. Journal of Research on the Lepidoptera 21 (1): 1 - 105" title="An annotated catalogue of the butterflies named by Roger Verity." volume="21" year="1983">Kudrna (1983</bibRefCitation>
: 84), this name is available as subspecific. The year on the specimen labels (1937) indicate specimens were collected 23 years after description (1914) from the same location the original holotype material was collected. As a result, the black label of the specimens treated as
<normalizedToken id="5A451764183235D4EA1960C24B02CF17" originalValue="“types”">&quot;types&quot;</normalizedToken>
is erroneous. The specimen collected from the same geographical location from which the type specimen was collected is called topotype. According to ICZN (1999), a topotype has no formal standing and is not regulated by the Code. Therefore, these six specimens are erroneously labelled as types, should be annotated as &quot;Not a Type&quot; and are treated here as
<normalizedToken id="0203103EB663130889676D9522F27B83" originalValue="“Non-type”">&quot;Non-type&quot;</normalizedToken>
specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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