157 lines
18 KiB
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157 lines
18 KiB
XML
<document id="21E3A530EF7DA7F08C333B89A647BE41" ID-DOI="http://dx.doi.org/10.3897/zookeys.542.5975" ID-GBIF-Dataset="b53aa92a-b9ac-43bd-a9b9-f468d62f0bdc" ID-PMC="PMC4714360" ID-Pensoft-Pub="1313-2970-542-1" ID-PubMed="26798285" ID-ZooBank="026407877355425BAB10BF1674510F12" ModsDocAuthor="" ModsDocDate="2015" ModsDocID="1313-2970-542-1" ModsDocOrigin="ZooKeys 542" ModsDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" checkinTime="1451243652205" checkinUser="pensoft" docAuthor="Bistroem, Olof, Nilsson, Anders N. & Bergsten, Johannes" docDate="2015" docId="57C8EC226994231FE3B5AC9036C2E50E" docLanguage="en" docName="ZooKeys 542: 1-379" docOrigin="ZooKeys 542" docSource="http://dx.doi.org/10.3897/zookeys.542.5975" docTitle="Laccophilus pulcher Bilardo & Rocchi 2004" docType="treatment" docVersion="5" lastPageNumber="153" masterDocId="E3118410B56DFF839B32FF89FFCCFFB0" masterDocTitle="Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)" masterLastPageNumber="379" masterPageNumber="1" pageNumber="152" updateTime="1701314941532" updateUser="plazi">
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<mods:title id="695271E08B78AA2348DE2468765FAA0C">Taxonomic revision of Afrotropical Laccophilus Leach, 1815 (Coleoptera, Dytiscidae)</mods:title>
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<mods:namePart id="A353F98F5149310A65FB44EBF88D9F55">Bistroem, Olof</mods:namePart>
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<mods:namePart id="A42498358B417D2E5984AF194A6CCD82">Nilsson, Anders N.</mods:namePart>
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<mods:namePart id="F0BFA1C23C24DBE6DCFA707BB5E9F43C">Bergsten, Johannes</mods:namePart>
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<mods:typeOfResource id="B9A063309391BB0EF52457FE2444F16D">text</mods:typeOfResource>
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<mods:date id="C32732843F23A4A398DD9592417829D5">2015</mods:date>
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<mods:number id="BEC1068D83405823BC405FFBBECD3302">542</mods:number>
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<mods:classification id="5B5380270F2056707013E41872FCA5BB">journal article</mods:classification>
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<mods:identifier id="0932621501639B6553347EB7BC3C930A" type="DOI">http://dx.doi.org/10.3897/zookeys.542.5975</mods:identifier>
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<treatment id="57C8EC226994231FE3B5AC9036C2E50E" ID-GBIF-Taxon="127862789" LSID="urn:lsid:plazi:treatment:57C8EC226994231FE3B5AC9036C2E50E" httpUri="http://treatment.plazi.org/id/57C8EC226994231FE3B5AC9036C2E50E" lastPageId="152" lastPageNumber="153" pageId="151" pageNumber="152">
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<subSubSection id="13B97B53CAC5F5AA637226DBF8FD4802" pageId="151" pageNumber="152" type="multiple">
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<paragraph id="6D9C655C6C4A853FA71E419811E280AE" pageId="151" pageNumber="152">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="A26C10A493EE55D4CC7F25776EDFA7D7" pageId="151" pageNumber="152" type="nomenclature">
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<paragraph id="C1E08644ED7C3D01EF7E9EF7011C1C32" pageId="151" pageNumber="152">
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<taxonomicName id="7FADEBDA8F4A62881F54EFCA7D6A4D3A" ID-CoL="6NRNN" authority="Bilardo & Rocchi, 2004" authorityName="Bilardo & Rocchi" authorityYear="2004" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher Bilardo & Rocchi, 2004</taxonomicName>
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Figs 118-119, 307-308, 457, 562
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</paragraph>
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</subSubSection>
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<subSubSection id="2A01F13D91594945441FF55AD14120CB" pageId="151" pageNumber="152" type="reference_group">
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<paragraph id="C6C56510655DBA704AB60940B69BC717" pageId="151" pageNumber="152">
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<taxonomicName id="B76264B14424A82D1C130B534331B1AF" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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<bibRefCitation id="246A64A1B0B12DB695B1A1335A2E9135" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="283 - 299" title="Dytiscidae (Coleoptera) del Gabon. (Parte quarta)." volume="145" year="2004">Bilardo and Rocchi 2004</bibRefCitation>
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: 286, 290, 291 (original description, faunistics);
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<bibRefCitation id="15ACFD9AF755B29979539025FDFE1833" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="195 - 238" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (6 eme partie). Parc National des Plateaux Bateke (missions 2005 et 2006)." volume="149" year="2008">Bilardo and Rocchi 2008</bibRefCitation>
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: 211, 234 (faunistics, biology);
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<bibRefCitation id="D1CBFEEF1D053B84482CB990DEE0B624" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="255" pageNumber="256" pagination="131 - 155" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (9 eme partie). Parc National des Plateaux Bateke (missions 2010 et 2012)." volume="154" year="2013">Bilardo and Rocchi 2013</bibRefCitation>
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: 141 (faunistics);
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<bibRefCitation id="9479E667C46454BB8403F3411590F27A" author="Nilsson, AN" journalOrPublisher="Apollo books, Stenstrup" pageId="260" pageNumber="261" title="A World Catalogue of the Family Dytiscidae, or the Diving Beetles (Coleoptera, Adephaga). Version 1. I. 2015" url="http://www2.emg.umu.se/projects/biginst/andersn/" year="2015">Nilsson 2015</bibRefCitation>
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: 216 (catalogue, faunistics).
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</paragraph>
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</subSubSection>
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<subSubSection id="90C02869C76D22B58032B46FDF6BF996" pageId="151" pageNumber="152" type="type locality">
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<paragraph id="5DFC6622ADD0B7F9EB41B19A1212BA00" pageId="151" pageNumber="152">Type locality.</paragraph>
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<paragraph id="C95DAC52BD2FCAE23B9D63950F70EC03" pageId="151" pageNumber="152">
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Gabon: Riv.
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<normalizedToken id="D983B6B456DD5F7EF750A39FF5D3F3F9" originalValue="Louayé">Louaye</normalizedToken>
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,
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<normalizedToken id="5DAC4780C7ED9AB5583F59676E4CCF15" originalValue="Mékambo">Mekambo</normalizedToken>
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, Malouma (
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<geoCoordinate id="7C35BE9126CBF2E756D212B79A59D2A7" direction="north" orientation="latitude" precision="555" value="0.36">0.36N</geoCoordinate>
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,
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<geoCoordinate id="23D7C48ED57E45D6CFD2894496EFD447" direction="east" orientation="longitude" precision="555" value="13.5">13.50E</geoCoordinate>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="0DBC3CF9E107C0A4B7021FB011A675E8" pageId="151" pageNumber="152" type="type material">
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<paragraph id="3029CE3535D78ACF37A37175B4446A59" pageId="151" pageNumber="152">Type material studied</paragraph>
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<paragraph id="36B48007D9817C81A7AC53DE176F51F1" pageId="151" pageNumber="152">
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(2 exs.): Holotype: male: "Gabon
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<normalizedToken id="E1638D3F887A361BB8E7DFE9481414C2" originalValue="Mékambo">Mekambo</normalizedToken>
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(Malouma) Riv.
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<normalizedToken id="5B63946EBE057E326B8416A8BD63AFEC" originalValue="Louayé">Louaye</normalizedToken>
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(Borne) 24/07/01 A. Bilardo / 4.12 x 2.24 / Holotypus /
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<taxonomicName id="10F01630B05EE26C9D19A558DFE1B44C" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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Bil. & Roc. Det. Bilardo 04" (MSNM). - Paratype: female: "Gabon, VIII. Bissok (Oyem) F. Lara 8/8/91 A. Bilardo / 4,00 x 2,24 / Paratypus /
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<taxonomicName id="6AE5B96E3EB218B7985ECD73CD4BEDFC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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Bil. & Roc. Det. Bilardo 04" (1 ex. MSNM; habitus in Fig. 457).
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</paragraph>
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</subSubSection>
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<subSubSection id="1D21EBFA0792AB2D1582E0F525F726DC" pageId="151" pageNumber="152" type="notes on taxonomy">
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<paragraph id="D2292EB359B15BD9FC4283CE7D968654" pageId="151" pageNumber="152">Notes on taxonomy.</paragraph>
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<paragraph id="648D856A4A53A96A272E145EF13CACB5" pageId="151" pageNumber="152">
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Reflecting solely at the shape of penis
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<taxonomicName id="63A8E9F6437CB144A7B76AB875FCD6E0" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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resembles much of
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<taxonomicName id="91BD3742FFCF52AE7848F163C427D666" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus saegeri" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="saegeri">Laccophilus saegeri</taxonomicName>
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, in this revision placed in another species group (see Figs 283-284). The penis of
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<taxonomicName id="5E06AEDDA9711B8D2024AE6261DF84DC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus saegeri" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="saegeri">Laccophilus saegeri</taxonomicName>
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is, however, more evenly curved than in
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<taxonomicName id="AD3AF7ED0925FB88DA5F48F82F3AA0BB" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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. Moreover, elytra are extensively dark coloured in
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<taxonomicName id="2A2FC750F09FC6AC3EA7DC5C724887DC" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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, while they are pale ferrugineous and provided with irrorations in
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<taxonomicName id="3A6794C2173F4F21F2ADCFF2A097D71A" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus saegeri" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="saegeri">Laccophilus saegeri</taxonomicName>
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. Similarity in male genitalia between these two species still indicates need of further study in the delimitation of species groups in
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<taxonomicName id="1E0FAA9FE5896EB6017F4893B9E2A04D" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="genus">Laccophilus</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="36094C2FEC024FDE88B727EBB3129157" pageId="151" pageNumber="152" type="diagnosis">
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<paragraph id="4A71B7292E4F9743C7DC39ED12F8AD97" pageId="151" pageNumber="152">Diagnosis.</paragraph>
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<paragraph id="1A16C18B68FA72917ACB9A4D4166A1A1" pageId="151" pageNumber="152">
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<taxonomicName id="939015CA2731238420D529C0AC0E09FE" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="151" pageNumber="152" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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is easy to distinguish by study of male genitalia. No other species in this species group has a penis of same type; penis in lateral aspect long, external outline curved, somewhat undulate. Extreme apex with a fine hook.
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</paragraph>
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</subSubSection>
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<subSubSection id="17D3B6421404CC12B5AFC645D7F1076F" lastPageId="152" lastPageNumber="153" pageId="151" pageNumber="152" type="description">
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<paragraph id="CBFCCBE69D02CCBEDB24A0688C113F74" pageId="151" pageNumber="152">Description.</paragraph>
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<paragraph id="1CFEAF0F57189AF6DABF34531042C631" pageId="151" pageNumber="152">Body length 4.0-4.1 mm, width 2.2 mm. Dorsal, colour pattern of body exhibits only minor variation (Fig. 457).</paragraph>
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<paragraph id="AB7E59BF44056FD0627C2948171A5096" pageId="151" pageNumber="152">Head: Pale ferrugineous. Rather shiny, although finely microsculptured; reticulation double but difference between size classes minimal. Large meshes extensively hardly discernible. At eyes with very fine, irregular punctures.</paragraph>
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<paragraph id="61A82225AA61B4665A23B06E701D1602" pageId="152" pageNumber="153">
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<pageBreakToken id="41B7E8156F156C8A697AAAFCA97AC54C" pageId="152" pageNumber="153" start="start">Pronotum</pageBreakToken>
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: Ferrugineous to dark ferrugineous, with pale ferrugineous anterior corners. Rather shiny, although distinctly microsculptured. Reticulation double but difference between size classes small. Large meshes, when discernible, contain 2-4 small meshes. At margin, except basally in the middle, with fine, scattered punctures.
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</paragraph>
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<paragraph id="2F1BD134FEA9E763B79101A9E8260A3D" pageId="152" pageNumber="153">Elytra: Blackish ferrugineous to dark brown, with a subbasal, transversely located, pale ferrugineous markings. Furthermore, with a transverse, pale area discernible posterior to middle. Additionally, extreme apex of elytra pale (Fig. 457). Rather shiny, although finely microsculptured. Reticulation double; large meshes strongly reduced and only in part discernible. Laterally and apically almost solely with fine meshes. Discal row of punctures formed by fine, somewhat irregular punctures. Dorsolateral and lateral rows of punctures indicated by sparse, scattered punctures. Laterally, with a pre-apical, finely pubescent furrow.</paragraph>
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<paragraph id="7F27A8D963651B04501EE23F3343A880" pageId="152" pageNumber="153">Ventral aspect: Ferrugineous to dark ferrugineous; no distinct colour pattern discernible. Rather shiny, although finely microsculptured. Abdomen with fine, curved striae. Prosternal process slender, apex extended, pointed. Metacoxal plates in anterior half with 8-9, almost transversely located, shallow furrows; in posterior half furrows strongly reduced and only minor fragments discernible. Apical ventrite asymmetric, with lateral keel (Fig. 118).</paragraph>
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<paragraph id="5E5D38A065444FF6DBF0D869AFE67107" pageId="152" pageNumber="153">Legs: Pro- and mesotarsus rather slender, somewhat extended, provided with suckers.</paragraph>
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<paragraph id="57472EB5D0BF01B39974F6BD8A2096DE" pageId="152" pageNumber="153">Male genitalia: Penis in lateral aspect long, external outline curved, somewhat undulate. Extreme apex finely hooked (Figs 307-308).</paragraph>
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<paragraph id="87E766CC416A1919C25184EA886D287B" pageId="152" pageNumber="153">Female: Pro- and mesotarsus slender. Apical ventrite symmetric, no lateral keel (Fig. 119).</paragraph>
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</subSubSection>
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<subSubSection id="52BEAAD70D63E5D1CB12E825866BB279" pageId="152" pageNumber="153" type="distribution">
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<paragraph id="986140642E967BA329EABAC953E850D0" pageId="152" pageNumber="153">Distribution.</paragraph>
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<paragraph id="C6A3D7BB46DB58CB918C4829B549B378" pageId="152" pageNumber="153">Gabon (Fig. 562).</paragraph>
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</subSubSection>
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<subSubSection id="4A5AB1A61BDB7A31CD11E194823DDB87" pageId="152" pageNumber="153" type="collecting circumstances">
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<paragraph id="30AE10F427D842A7A9F7C1911FC895BE" pageId="152" pageNumber="153">Collecting circumstances.</paragraph>
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<paragraph id="05054CAB1C3EDA0D9D34724ED2A3F791" pageId="152" pageNumber="153">
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Almost unknown. Some information may be found in available literature (e.g.
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<bibRefCitation id="387A17F5730DE971105AEC3FA6F06397" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="283 - 299" title="Dytiscidae (Coleoptera) del Gabon. (Parte quarta)." volume="145" year="2004">Bilardo and Rocchi 2004</bibRefCitation>
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,
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<bibRefCitation id="9FF1D4B66C15D91E83D3E1FAEA3DDA7D" author="Bilardo, A" journalOrPublisher="Atti della Societa Italiana di Scienze Naturali e del Museo Civico di Storia Naturale di Milano" pageId="254" pageNumber="255" pagination="195 - 238" title="Haliplidae, Noteridae, Dytiscidae (Coleoptera) du Gabon (6 eme partie). Parc National des Plateaux Bateke (missions 2005 et 2006)." volume="149" year="2008">2008</bibRefCitation>
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):
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<taxonomicName id="D5B805DF1306F5A3F32DEE3B74E49D60" class="Insecta" family="Dytiscidae" genus="Laccophilus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Laccophilus pulcher" order="Coleoptera" pageId="152" pageNumber="153" phylum="Arthropoda" rank="species" species="pulcher">Laccophilus pulcher</taxonomicName>
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was collected in puddles of a residual stream in forest. It is also listed as a general forest species by the present authors.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |