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<document ID-DOI="http://dx.doi.org/10.3897/jhr.33.5078" ID-GBIF-Taxon="182249314" ID-Pensoft-Pub="1314-2607-33-1" ID-Pensoft-UUID="5F20FFE2E95AAC21FFD1FFA0D8160112" ID-Zenodo-Dep="574825" ID-ZooBank="88E03F8FD9B947F1B6E90456D47B1E37" ModsDocID="1314-2607-33-1" checkinTime="1621553083643" checkinUser="pensoft" docAuthor="Kimsey, Lynn S. &amp; Wasbauer, Marius S." docDate="2013" docId="5745AE48A254A5604FDC06E147AE5A95" docLanguage="en" docOrigin="Journal of Hymenoptera Research 33" docPubDate="2013-08-01" docSource="http://dx.doi.org/10.3897/jhr.33.5078" docTitle="Colocistis crassa" docType="treatment" docVersion="2" id="5F20FFE2E95AAC21FFD1FFA0D8160112" lastPageId="8" lastPageNumber="9" masterDocId="5F20FFE2E95AAC21FFD1FFA0D8160112" masterDocTitle="Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae)" masterLastPageNumber="24" masterPageNumber="1" pageId="7" pageNumber="8" updateTime="1643536911076" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the brachycistidine genus Colocistis Krombein, 1942 (Hymenoptera, Tiphiidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kimsey, Lynn S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology, University of California, Davis, 95616 USA</mods:affiliation>
<mods:nameIdentifier type="email">lskimsey@ucdavis.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Wasbauer, Marius S.</mods:namePart>
<mods:affiliation>Bohart Museum of Entomology, Department of Entomology, University of California, Davis, 95616 USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2013</mods:date>
<mods:detail type="pubDate">
<mods:number>2013-08-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>33</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>24</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.33.5078</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.33.5078</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-33-1</mods:identifier>
<mods:identifier type="ZooBank">88E03F8FD9B947F1B6E90456D47B1E37</mods:identifier>
<mods:identifier type="Pensoft-UUID">5F20FFE2E95AAC21FFD1FFA0D8160112</mods:identifier>
<mods:identifier type="Zenodo-Dep">574825</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="182249314" LSID="urn:lsid:plazi:treatment:5745AE48A254A5604FDC06E147AE5A95" httpUri="http://treatment.plazi.org/id/5745AE48A254A5604FDC06E147AE5A95" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8">
<subSubSection pageId="7" pageNumber="8" type="nomenclature">
<paragraph pageId="7" pageNumber="8">
<taxonomicName LSID="http://species-id.net/wiki/Colocistis_crassa" authority="(Bradley)" baseAuthorityName="Bradley" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Colocistis crassa" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassa">Colocistis crassa (Bradley)</taxonomicName>
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="7" pageNumber="8">Figures 4</figureCitation>
<figureCitation captionStart="Figures 1324" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="7" pageNumber="8">, 19, 22</figureCitation>
<figureCitation captionStart="Figures 3336" captionStartId="F4" captionText="Figures 33 - 36. Distribution maps of Colocistis species." httpUri="https://binary.pensoft.net/fig/11790" pageId="7" pageNumber="8">, 35</figureCitation>
<figureCitation captionStart="Figures 4148" captionStartId="F6" captionText="Figures 41 - 48. Ventral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11792" pageId="7" pageNumber="8">, 44</figureCitation>
<figureCitation captionStart="Figures 4956" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="7" pageNumber="8">, 52</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="reference_group">
<paragraph pageId="7" pageNumber="8">
<taxonomicName class="Insecta" family="Formicidae" genus="Brachycistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassa">Brachycistis crassa</taxonomicName>
Bradley, 1917:277. Holotype males; USA: California, Fresno County, Coalinga (CUIC).*
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="8" type="diagnosis">
<paragraph pageId="7" pageNumber="8">Diagnosis.</paragraph>
<paragraph pageId="7" pageNumber="8">
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassa">
<emphasis italics="true" pageId="7" pageNumber="8">Colocistis crassa</emphasis>
</taxonomicName>
and
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="7" pageNumber="8">Colocistis pilosa</emphasis>
</taxonomicName>
share a distinctive modification of metasomal sternum II, which has the basal area elevated and separated from the rest of the sternum by a transverse declivity. These two species can be separated by the castaneous body color in
<taxonomicName class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="crassa">
<emphasis italics="true" pageId="7" pageNumber="8">Colocistis crassa</emphasis>
</taxonomicName>
(black in
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="7" pageNumber="8">Colocistis pilosa</emphasis>
</taxonomicName>
), clypeus with a well-developed medial tubercle (no tubercle in
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="7" pageNumber="8">Colocistis pilosa</emphasis>
</taxonomicName>
) and large compound eyes and ocelli (small in
<taxonomicName authorityName="Krombein" authorityYear="1942" class="Insecta" family="Tiphiidae" genus="Colocistis" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Hymenoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="pilosa">
<emphasis italics="true" pageId="7" pageNumber="8">Colocistis pilosa</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="description">
<paragraph pageId="8" pageNumber="9">
<pageBreakToken pageId="8" pageNumber="9" start="start">Description</pageBreakToken>
.
</paragraph>
<paragraph pageId="8" pageNumber="9">Male.</paragraph>
<paragraph pageId="8" pageNumber="9">
<emphasis italics="true" pageId="8" pageNumber="9">Body length</emphasis>
. 9-16 mm.
</paragraph>
<paragraph pageId="8" pageNumber="9">
<emphasis italics="true" pageId="8" pageNumber="9">Head</emphasis>
(
<figureCitation captionStart="Figures 112" captionStartId="F1" captionText="Figures 1 - 12. 1 - 8 Front view of male head. 9 Lateral view of male head 10 - 12 Ventral view of female head. Abbreviations: LID = least interocular distance, OcC = occipital carina; OOL = ocellocular distance, THC = transverse hypostomal carina, UID = upper interocular distance." httpUri="https://binary.pensoft.net/fig/11787" pageId="8" pageNumber="9">Fig. 4</figureCitation>
). Compound eyes enlarged, extending well beyond temples; ocelli enlarged, diameter of lateral ocellus at least 0.7
<normalizedToken originalValue="×">x</normalizedToken>
OOL; impressed line present between lateral ocelli; antennal socket rim thickened and carinate ventrolaterally; clypeus with median tubercle, visible above mandible in lateral view, apex not bounded laterally by strong teeth; gular carina not raised; LID 0.7
<normalizedToken originalValue="×">x</normalizedToken>
UID; flagellomere I length 3.2-3.3
<normalizedToken originalValue="×">x</normalizedToken>
breadth; flagellomere II length 2.7-2.8
<normalizedToken originalValue="×">x</normalizedToken>
breadth; flagellomere IX length 5.7-5.8
<normalizedToken originalValue="×">x</normalizedToken>
breadth; occipital carina strong, incomplete ventrally; hypostomal carina not raised; mandible narrowest submedially, tapering apically toward ventroapical tooth.
</paragraph>
<paragraph pageId="8" pageNumber="9">
<emphasis italics="true" pageId="8" pageNumber="9">Mesosoma</emphasis>
. Pronotum nearly vertical anteriorly, punctures separated by 0.5-1.0 puncture diameters, shoulders well developed, not produced laterally; forefemur without stout spines; foretibia without spines except apically; basitarsus with two spines on outer surface; tibial spur 0.9
<normalizedToken originalValue="×">x</normalizedToken>
length of basitarsus; hindcoxal longitudinal dorsal carina forming obtuse angle basally; scutal and mesopleural punctures 1-2 puncture diameters apart; scutellum somewhat curved above level of propodeum, coarsely punctured with large, polished impunctate area medially; postscutellar punctures 1-2 puncture diameters apart; propodeum with long dorsal surface, nearly impunctate, with wide medial, longitudinal trough, reaching carina separating dorsal from posterior surfaces, posterior surface with shallow punctures, 0.5-1.0 puncture diameters apart, lateral surface punctures 0.5-1.0 puncture diameters apart dorsally, nearly impunctate ventrally; forewing with 3 submarginal cells; costa extending along wing margin beyond stigma.
</paragraph>
<paragraph pageId="8" pageNumber="9">
<emphasis italics="true" pageId="8" pageNumber="9">Metasoma</emphasis>
. Tergum I with transverse carina at apex of frontal declivity (
<figureCitation captionStart="Figures 1324" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="8" pageNumber="9">Fig. 22</figureCitation>
); sternum II with anterior portion slightly raised and separated from remainder of sternum by a transverse declivity (
<figureCitation captionStart="Figures 1324" captionStartId="F2" captionText="Figures 13 - 24. 13, 14 Male forewing. 15 Ventral view of male hindcoxa 16 Dorsal view of male propodeum 17 dorsal view of male mandible 18 lateral view of mandible 19 Ventral view of male metasomal sterna I-II 20, 22 Lateral view of male metasomal segments I and II 21, 23, 24 Ventral view of female metasomal sternum I. Abbreviations: SM- 1, 2, 3 = submarginal cells 1 - 3." httpUri="https://binary.pensoft.net/fig/11788" pageId="8" pageNumber="9">Figs 19, 22</figureCitation>
).
</paragraph>
<paragraph pageId="8" pageNumber="9">
<emphasis italics="true" pageId="8" pageNumber="9">Genitalia</emphasis>
(
<figureCitation captionStart="Figures 4148" captionStartId="F6" captionText="Figures 41 - 48. Ventral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11792" pageId="8" pageNumber="9">Figs 44</figureCitation>
,
<figureCitation captionStart="Figures 4956" captionStartId="F7" captionText="Figures 49 - 56. Lateral view of male genital capsule." httpUri="https://binary.pensoft.net/fig/11793" pageId="8" pageNumber="9">52</figureCitation>
). Gonostylar apex somewhat excavate in lateral view; aedeagus abruptly decurved in lateral view, apex with dorsally projecting process.
</paragraph>
<paragraph pageId="8" pageNumber="9">
<emphasis italics="true" pageId="8" pageNumber="9">Color</emphasis>
. Head and body unicolorous, castaneous; vestiture pale brown.
</paragraph>
<paragraph pageId="8" pageNumber="9">Female unknown.</paragraph>
</subSubSection>
<subSubSection pageId="8" pageNumber="9" type="distribution">
<paragraph pageId="8" pageNumber="9">Distribution</paragraph>
<paragraph pageId="8" pageNumber="9">
(
<figureCitation captionStart="Figures 3336" captionStartId="F4" captionText="Figures 33 - 36. Distribution maps of Colocistis species." httpUri="https://binary.pensoft.net/fig/11790" pageId="8" pageNumber="9">Fig. 35</figureCitation>
). USA:
<emphasis bold="true" pageId="8" pageNumber="9">Arizona</emphasis>
: Cochise, Coconino, Gila, Graham, Maricopa, Pima, Pinal, Santa Cruz, Yuma counties;
<emphasis bold="true" pageId="8" pageNumber="9">California</emphasis>
: Alpine, Fresno, Imperial, Inyo, Kern, Kings, Lassen, Los Angeles, Madera, Merced, Mono, Monterey, Riverside, San Benito, San Bernardino, San Diego, San Luis Obispo, Santa Barbara, Santa Clara, Sonoma, Stanislaus, Tuolumne, Tulare, Ventura counties;
<emphasis bold="true" pageId="8" pageNumber="9">Idaho</emphasis>
: Ada, Elmore, Owyhee counties; Kansas: Stanton County;
<emphasis bold="true" pageId="8" pageNumber="9">Nevada</emphasis>
: Churchill, Esmeralda, Eureka, Humboldt, Lincoln, Nye, Pershing, Washoe counties;
<emphasis bold="true" pageId="8" pageNumber="9">New Mexico</emphasis>
: Dona Ana, Eddy, Hidalgo, Luna, Quay counties;
<emphasis bold="true" pageId="8" pageNumber="9">Texas</emphasis>
: Alpine, Brewser, Culberson, El Paso, Jeff Davis counties;
<emphasis bold="true" pageId="8" pageNumber="9">Utah</emphasis>
: Ogden County; Mexico:
<emphasis bold="true" pageId="8" pageNumber="9">Baja California</emphasis>
,
<emphasis bold="true" pageId="8" pageNumber="9">Baja California Sur</emphasis>
,
<emphasis bold="true" pageId="8" pageNumber="9">Chihuahua</emphasis>
,
<emphasis bold="true" pageId="8" pageNumber="9">San Luis Potosi</emphasis>
,
<emphasis bold="true" pageId="8" pageNumber="9">Sinaloa</emphasis>
,
<emphasis bold="true" pageId="8" pageNumber="9">Sonora</emphasis>
; 1250 males were examined (BME, CAS, CDFA, EMEC, KSBS, LACM, MCZ).
</paragraph>
</subSubSection>
</treatment>
</document>