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<mods:title id="DF06D1992EBBE906F11ABFD714644A4F">A taxonomic review of Korean Liogluta Thomson (Coleoptera, Staphylinidae, Aleocharinae) with descriptions of three new species</mods:title>
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<taxonomicName id="19CEFFBFE84AFFC5609BFEB4FE57FE30" ID-CoL="3V66F" authority="Sawada, 1972" authorityName="Sawada" authorityYear="1972" box="[151,505,367,394]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="301" phylum="Arthropoda" rank="species" species="ursi">
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<emphasis id="ECBA582EE84AFFC5609BFEB4FE9DFE33" bold="true" box="[151,307,367,393]" italics="true" pageId="16" pageNumber="301">Liogluta ursi</emphasis>
(
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(
<figureCitation id="46F598B9E84AFFC56093FE49FF47FE11" box="[159,233,402,427]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1989]" captionTargetBox="[160,1427,208,1929]" captionTargetId="figure@2.[152,1435,193,1945]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Liogluta. A, L. changwhani sp. nov.; B, L. distans; C, L. granigera; D, L. infans. E, L. jinilli sp. nov.; F, L. pyonganica; G, L. rufescens sp. nov.; H, L. ursi." httpUri="https://zenodo.org/record/166919/files/figure.png" pageId="16" pageNumber="301">Figs. 1</figureCitation>
H, 9)
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<emphasis id="ECBA582EE84AFFC5609BFE01FEA9FE4A" box="[151,263,474,496]" italics="true" pageId="16" pageNumber="301">Atheta ursi</emphasis>
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: 53
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<emphasis id="ECBA582EE84AFFC5609BFE22FEB4FDB5" box="[151,282,505,527]" italics="true" pageId="16" pageNumber="301">Liogluta ursi</emphasis>
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;
<treatmentCitation id="5F6FA22DE84AFFC56124FE22FD9AFDB5" author="Yosii" box="[296,564,505,527]" page="97" pageId="16" pageNumber="301" year="1976">
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: 97
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<bibRefCitation id="BA5FF9CDE84AFFC56232FE22FD77FDB5" author="Smetana" box="[574,729,505,527]" pageId="16" pageNumber="301" refString="Smetana, A. (2004) Family Staphylinidae: Aleocharinae. pp. 353 - 494. In: Lobl, I., &amp; Smetana A. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 2. Hydrophiloidea - Histeroidea - Staphylinoidea. Apollo Books, Stenstrup, 942 pp." type="book chapter" year="2004">Smetana, 2004</bibRefCitation>
: 411
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<bibRefCitation id="BA5FF9CDE84AFFC56318FE22FB8EFDB5" author="Schulke" box="[788,1056,505,527]" pageId="16" pageNumber="301" refString="Schulke, M. &amp; Smetana, A. (2015) Staphylinidae Latreille, 1802. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 2 / 1. Revised and updated edition. Hydrophiloidea - Staphylinoidea. Brill, Leiden, pp. 304 - 1134." type="journal article" year="2015">Schülke &amp; Smetana, 2015</bibRefCitation>
: 567
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<paragraph id="DE71843CE84AFFC5609BFDE7FDA4F816" blockId="16.[151,1437,572,2037]" pageId="16" pageNumber="301">
<emphasis id="ECBA582EE84AFFC5609BFDE7FE87FDEF" bold="true" box="[151,297,572,597]" pageId="16" pageNumber="301">Description.</emphasis>
Length
<quantity id="193629D9E84AFFC5619BFDE7FD85FDEE" box="[407,555,572,596]" metricMagnitude="-3" metricUnit="m" metricValue="3.0999999999999996" metricValueMax="3.4" metricValueMin="2.8" pageId="16" pageNumber="301" unit="mm" value="3.0999999999999996" valueMax="3.4" valueMin="2.8">2.83.4 mm</quantity>
. Body fusiform; surface fairly glossy and densely pubescent, with fine microsculpture. Body (
<figureCitation id="46F598B9E84AFFC56190FD84FE73FDC2" box="[412,477,607,632]" captionStart="FIGURE 1" captionStartId="2.[151,250,1967,1989]" captionTargetBox="[160,1427,208,1929]" captionTargetId="figure@2.[152,1435,193,1945]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Liogluta. A, L. changwhani sp. nov.; B, L. distans; C, L. granigera; D, L. infans. E, L. jinilli sp. nov.; F, L. pyonganica; G, L. rufescens sp. nov.; H, L. ursi." httpUri="https://zenodo.org/record/166919/files/figure.png" pageId="16" pageNumber="301">Fig. 1</figureCitation>
H) reddish brown to dark brown; head darker than other parts; elytra and legs paler brown.
<emphasis id="ECBA582EE84AFFC560E3FD5FFE85FD27" box="[239,299,644,669]" italics="true" pageId="16" pageNumber="301">Head</emphasis>
. More or less circular, approximately 1.01.1 times as wide as long, widest across eyes, distinctly narrower than pronotum; eyes relatively small, about 0.81.0 times as long as temples; gular sutures moderately separated, diverged basally; infraorbital carina complete; cervical carina absent. Antennae (
<figureCitation id="46F598B9E84AFFC564CAFD17FAA3FD5E" box="[1222,1293,716,741]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Fig. 9</figureCitation>
A) long and slender; antennomeres 13 elongate, 2 slightly longer than 3, 4 about as long as wide, 510 subquadrate to slightly transverse, 11 longest, about as long as preceding two combined.
<emphasis id="ECBA582EE84AFFC563CDFCCFFBEAFC97" box="[961,1092,788,813]" italics="true" pageId="16" pageNumber="301">Mouthparts</emphasis>
. Labrum transverse, slightly emarginate in anterior margin, with ε-sensillum and about 8 macrosetae on each side of midline; epipharynx with several sensilla, including 2 lateral sensory rows on each side of midline; α-sensillum setaceous, slightly longer than ε-sensillum, β- and γ-sensilla very short, convergent at apex. Mandibles asymmetrical, subtriangular, decurved and pointed apically, about 1.6 times as long as basal width; right one with small internal tooth, internal margin slightly serrulate; prostheca developed, composited three portions, second portion longer. Galea and lacinia of maxilla long and slender; lacinia having seven spines in distal comb region, two isolated spines present; maxillary palpus elongate with pubescence and long setae; palpomere 1 smallest, 2 about 2.83.0 times as long as wide, 3 slightly longer than 2, about 3.0 times as long as wide, 4 digitiform, filamentous sensilla not reaching to basal half. Labium with ligula long and slender, divided into 2 lobes in basal third; prementum with two medial setae widely separated; two basal pores narrowly separated, about 1.0 times width of basal pore; several medial pseudopores, 1 setal pore and 2 real pores present on each side of midline; labial palpus very elongate, with many setulae; palpomere 1 largest, about 2.02.4 times as long as wide, γ-setula contiguous with b-seta, 2 shortest, about 1.31.5 times as long as wide, 3 parallel-sided and shorter than 1, about 3.5 times as long as wide. Mentum trapezoidal, anterior margin truncate; v-seta relatively long, close to u-seta.
<emphasis id="ECBA582EE84AFFC56350FAD7FC03FA9F" box="[860,941,1292,1317]" italics="true" pageId="16" pageNumber="301">Thorax</emphasis>
. Pronotum approximately 1.4 times as wide as long, widest at middle. Metanotal scutum with 1 long seta and about 57 relatively short setae on each side of midline. Mesocoxal cavities narrowly separated; mesoventral process pointed at apex; length ratio of mesoventral process, isthmus and metaventral process 6:4:1. Elytra slightly longer and slightly wider than pronotum; elytron approximately 1.41.5 times as long as wide, postero-lateral margin almost straight; pubescence directed posterior and postero-laterally; hind wings fully developed, flabellum composed of about 910 setose lobes.
<emphasis id="ECBA582EE84AFFC5650EFA1AFA99FA62" box="[1282,1335,1473,1496]" italics="true" pageId="16" pageNumber="301">Legs</emphasis>
. Slender and long, with pubescence and macrosetae; tibiae with different length of two spurs at apex; tarsal formula 4-5-5, length ratio of tarsomeres 32:36:40:90 (protarsus); 42:51:52:48:90 (mesotarsus); 75:71:57:57:110 (metatarsus); one empodial seta present, shorter than claw.
<emphasis id="ECBA582EE84AFFC562CFF9F7FC83F9FF" box="[707,813,1580,1605]" italics="true" pageId="16" pageNumber="301">Abdomen</emphasis>
. Narrow apically; surface fairly glossy and densely pubescent, fine and transverse microsculpture; conspicuous macrosetae present in lateral regions; macrochaetal arrangement of tergites
<date id="AA70A2FCE84AFFC56193F9AEFDE5F936" box="[415,587,1652,1676]" pageId="16" pageNumber="301" value="2013-02-01" valueMax="2013-06-12" valueMin="2013-02-01">IIVI 01-12-13</date>
-13-13 (or 23); tergite VII subquadrate; male tergite VIII (
<figureCitation id="46F598B9E84AFFC564D8F9AFFAB8F936" box="[1236,1302,1652,1677]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Fig. 9</figureCitation>
B) with 5 or 6 macrosetae on each side of midline, posterior margin truncate; male sternite VIII (
<figureCitation id="46F598B9E84AFFC5644BF94CFB27F90A" box="[1095,1161,1687,1712]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Fig. 9</figureCitation>
C) with 7 macrosetae on each side of midline, posterior margin subtruncate; posterior margin of female tergite VIII (
<figureCitation id="46F598B9E84AFFC564CDF967FAA8F96E" box="[1217,1286,1724,1749]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Fig. 9</figureCitation>
D) similar to male; female sternite VIII (
<figureCitation id="46F598B9E84AFFC561C6F904FDA3F942" box="[458,525,1759,1784]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Fig. 9</figureCitation>
E) with 6 macrosetae on each side of midline, posterior margin subtruncate, with long marginal setae and minute setae in median region.
<emphasis id="ECBA582EE84AFFC56318F8DFFC2FF8A7" box="[788,897,1796,1821]" italics="true" pageId="16" pageNumber="301">Aedeagus</emphasis>
. Median lobe (
<figureCitation id="46F598B9E84AFFC56423F8DFFBD2F8A6" box="[1071,1148,1796,1821]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Figs. 9</figureCitation>
FG) narrowly ovate and widest in basal third, apical process short and convergent at apex in ventral aspect. Paramerite not pointed apically. Apical lobe of paramerites (
<figureCitation id="46F598B9E84AFFC561D2F897FD8AF8DE" box="[478,548,1868,1893]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">Fig. 9</figureCitation>
H) recurved and pointed apically, with four setae; a-seta longest, other setae subequal in length.
<emphasis id="ECBA582EE84AFFC56175F8B4FDA5F832" box="[377,523,1903,1928]" italics="true" pageId="16" pageNumber="301">Spermatheca</emphasis>
. Bursa elongate, with conical shaped umbilicus; duct long and slender, entirely coiled and apex twisted (
<figureCitation id="46F598B9E84AFFC561A2F84FFE5CF816" box="[430,498,1940,1965]" captionStart="FIGURE 9" captionStartId="17.[151,250,1767,1789]" captionTargetBox="[179,1408,203,1728]" captionTargetId="figure@17.[168,1411,193,1738]" captionTargetPageId="17" captionText="FIGURE 9. Liogluta ursi. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm." httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="16" pageNumber="301">
Fig.
<date id="AA70A2FCE84AFFC561EEF84FFE5CF816" box="[482,498,1940,1964]" pageId="16" pageNumber="301">9</date>
</figureCitation>
I).
</paragraph>
</subSubSection>
<subSubSection id="96D4D7B7E84AFFC560CBF863FC44F84F" pageId="16" pageNumber="301" type="materials_examined">
<paragraph id="DE71843CE84AFFC560CBF863FC44F84F" blockId="16.[151,1437,572,2037]" pageId="16" pageNumber="301">
<materialsCitation id="6EA68E61E84AFFC560CBF863FC44F84F" ID-GBIF-Occurrence="1325928528" collectingDate="2004-06-18" collectorName="Mt. Odaesan, N &amp; SJ Park" country="South Korea" county="Material" location="Jinbumyeon" municipality="Pyeongchang-gun" pageId="16" pageNumber="301" specimenCount="1" stateProvince="Gangwon">
<emphasis id="ECBA582EE84AFFC560CBF863FE1BF86B" bold="true" box="[199,437,1976,2001]" pageId="16" pageNumber="301">
<collectingCounty id="3710FCB0E84AFFC560CBF863FE81F86B" box="[199,303,1976,2001]" pageId="16" pageNumber="301">Material</collectingCounty>
examined.
</emphasis>
<collectingCountry id="A6D9C4ACE84AFFC561CDF863FD25F86A" box="[449,651,1976,2000]" name="South Korea" pageId="16" pageNumber="301">SOUTH KOREA</collectingCountry>
:
<emphasis id="ECBA582EE84AFFC562ACF862FCE2F86A" box="[672,844,1977,2000]" italics="true" pageId="16" pageNumber="301">
<collectingRegion id="1C0A4ADEE84AFFC562ACF862FCA3F86A" box="[672,781,1977,2000]" country="South Korea" name="Gangwon" pageId="16" pageNumber="301">Gangwon</collectingRegion>
Prov
</emphasis>
.: 9 exx. (four on slide),
<collectingMunicipality id="3E151E46E84AFFC56475F863FAE8F86B" box="[1145,1350,1976,2001]" pageId="16" pageNumber="301">Pyeongchang-gun</collectingMunicipality>
,
<location id="DB11D2E7E84AFFC56559F863FF48F84F" LSID="urn:lsid:plazi:treatment:5667352AE84AFFC7609BFEB4FDFCFED9:DB11D2E7E84AFFC56559F863FF48F84F" country="South Korea" county="Material" municipality="Pyeongchang-gun" name="Jinbumyeon" pageId="16" pageNumber="301" stateProvince="Gangwon">Jinbumyeon</location>
,
<collectorName id="733BE1EAE84AFFC560FDF806FE0AF84F" box="[241,420,2012,2037]" pageId="16" pageNumber="301">
<location id="DB11D2E7E84AFFC560FDF806FE2DF84F" LSID="urn:lsid:plazi:treatment:5667352AE84AFFC7609BFEB4FDFCFED9:DB11D2E7E84AFFC560FDF806FE2DF84F" box="[241,387,2012,2037]" country="South Korea" county="Material" municipality="Pyeongchang-gun" name="Mt. Odaesan" pageId="16" pageNumber="301" stateProvince="Gangwon">Mt. Odaesan</location>
, N
</collectorName>
37˚47'03&quot; E128˚33'55&quot;,
<collectingDate id="BA345B14E84AFFC562BEF807FC83F84E" box="[690,813,2012,2037]" pageId="16" pageNumber="301" value="2004-06-18">
<date id="AA70A2FCE84AFFC562BEF807FC83F84E" box="[690,813,2012,2037]" pageId="16" pageNumber="301" value="2004-06-18">18 vi 2004</date>
</collectingDate>
,
<collectorName id="733BE1EAE84AFFC56334F807FC21F84F" box="[824,911,2012,2037]" pageId="16" pageNumber="301">SJ Park</collectorName>
, sifting.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="8AB1D4B4E84BFFC4609BF93CFF68F8E3" httpUri="https://zenodo.org/record/166927/files/figure.png" pageId="17" pageNumber="302" targetBox="[179,1408,203,1728]" targetPageId="17">
<paragraph id="DE71843CE84BFFC4609BF93CFF68F8E3" blockId="17.[151,1436,1767,1881]" pageId="17" pageNumber="302">
<emphasis id="ECBA582EE84BFFC4609BF93CFEBCF946" bold="true" box="[151,274,1767,1789]" pageId="17" pageNumber="302">FIGURE 9.</emphasis>
<taxonomicName id="19CEFFBFE84BFFC46115F93CFE33F947" box="[281,413,1767,1789]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="302" phylum="Arthropoda" rank="species" species="ursi">
<emphasis id="ECBA582EE84BFFC46115F93CFE33F947" box="[281,413,1767,1789]" italics="true" pageId="17" pageNumber="302">Liogluta ursi</emphasis>
</taxonomicName>
. A, antenna; B, male abdominal tergite VIII (dorsal aspect); C, male abdominal sternite VIII (ventral aspect); D, female abdominal tergite VIII (dorsal aspect); E, female abdominal sternite VIII (ventral aspect); F, median lobe of aedeagus (lateral aspect); G, median lobe of aedeagus (ventral aspect); H, paramere (lateral aspect); I: spermatheca. Scales = 0.1 mm.
</paragraph>
</caption>
<subSubSection id="96D4D7B7E84BFFC460CBF85EFD2AF824" box="[199,644,1925,1950]" pageId="17" pageNumber="302" type="distribution">
<paragraph id="DE71843CE84BFFC460CBF85EFD2AF824" blockId="17.[151,1436,1925,2023]" box="[199,644,1925,1950]" pageId="17" pageNumber="302">
<emphasis id="ECBA582EE84BFFC460CBF85EFECEF824" bold="true" box="[199,352,1925,1950]" pageId="17" pageNumber="302">Distribution.</emphasis>
<collectingCountry id="A6D9C4ACE84BFFC4616BF85CFE05F824" box="[359,427,1927,1950]" name="South Korea" pageId="17" pageNumber="302">Korea</collectingCountry>
(South) and
<collectingCountry id="A6D9C4ACE84BFFC46231F85CFD2EF824" box="[573,640,1927,1950]" name="Japan" pageId="17" pageNumber="302">Japan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="96D4D7B7E84BFFC760CBF872FDFCFED9" lastPageId="18" lastPageNumber="303" pageId="17" pageNumber="302" type="discussion">
<paragraph id="DE71843CE84BFFC760CBF872FD21FF6F" blockId="17.[151,1436,1925,2023]" lastBlockId="18.[151,1436,151,355]" lastPageId="18" lastPageNumber="303" pageId="17" pageNumber="302">
<emphasis id="ECBA582EE84BFFC460CBF872FE95F878" bold="true" box="[199,315,1961,1986]" pageId="17" pageNumber="302">Remarks.</emphasis>
<taxonomicName id="19CEFFBFE84BFFC46147F872FE48F878" box="[331,486,1961,1986]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="302" phylum="Arthropoda" rank="species" species="ursi">
<emphasis id="ECBA582EE84BFFC46147F872FE48F878" box="[331,486,1961,1986]" italics="true" pageId="17" pageNumber="302">Liogluta ursi</emphasis>
</taxonomicName>
can be distinguished from other
<taxonomicName id="19CEFFBFE84BFFC46380F872FC45F878" box="[908,1003,1961,1986]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="302" phylum="Arthropoda" rank="genus">
<emphasis id="ECBA582EE84BFFC46380F872FC45F878" box="[908,1003,1961,1986]" italics="true" pageId="17" pageNumber="302">Liogluta</emphasis>
</taxonomicName>
species by the fusiform body; the macrochaetal arrangement of abdominal tergites; and the different shape and structure of the aedeagus and spermatheca. This species was described by
<bibRefCitation id="BA5FF9CDE848FFC7629AFF4CFCEFFF0A" author="Sawada" box="[662,833,151,176]" pageId="18" pageNumber="303" refString="Sawada, K. (1972) Methodological research in the Taxonomy of Aleocharinae. Contribution from the Biological Laboratory Kyoto University, 24 (1), 31 - 59." type="journal article" year="1972">Sawada (1972)</bibRefCitation>
and transferred to the genus
<taxonomicName id="19CEFFBFE848FFC7649FFF4CFB5CFF0A" box="[1171,1266,151,176]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="ECBA582EE848FFC7649FFF4CFB5CFF0A" box="[1171,1266,151,176]" italics="true" pageId="18" pageNumber="303">Liogluta</emphasis>
</taxonomicName>
based on their character systems in
<bibRefCitation id="BA5FF9CDE848FFC7618DFF67FD24FF6F" author="Yosii" box="[385,650,188,213]" pageId="18" pageNumber="303" refString="Yosii, R. &amp; Sawada, K. (1976) Studies on the genus Atheta Thomson and its allies (Coleoptera, Staphylinidae). II: Diagnostic characters of genera and subgenera with description of representative species. Contributions from the Biological Laboratory, Kyoto University, 25 (1), 11 - 140." type="journal article" year="1976">Yosii &amp; Sawada (1976)</bibRefCitation>
.
</paragraph>
<paragraph id="DE71843CE848FFC760CBFF04FDFCFED9" blockId="18.[151,1436,151,355]" pageId="18" pageNumber="303">
The systematic position of
<taxonomicName id="19CEFFBFE848FFC761F1FF3AFDE7FF42" box="[509,585,225,248]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="species" species="ursi">
<emphasis id="ECBA582EE848FFC761F1FF3AFDE7FF42" box="[509,585,225,248]" italics="true" pageId="18" pageNumber="303">L. ursi</emphasis>
</taxonomicName>
is doubtful, considering the fusiform body that distinguishes it from other
<taxonomicName id="19CEFFBFE848FFC7609BFED8FF58FEA6" box="[151,246,259,284]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="ECBA582EE848FFC7609BFED8FF58FEA6" box="[151,246,259,284]" italics="true" pageId="18" pageNumber="303">Liogluta</emphasis>
</taxonomicName>
species, including the
<typeStatus id="01753A9EE848FFC76204FEDFFD96FEA6" box="[520,568,260,284]" pageId="18" pageNumber="303">type</typeStatus>
species. However, the species has most diagnostic characters of
<taxonomicName id="19CEFFBFE848FFC76531FED8FA32FEA6" box="[1341,1436,259,284]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="ECBA582EE848FFC76531FED8FA32FEA6" box="[1341,1436,259,284]" italics="true" pageId="18" pageNumber="303">Liogluta</emphasis>
</taxonomicName>
except for the body form. Therefore, its generic assignment to the
<taxonomicName id="19CEFFBFE848FFC76392FEFDFC53FE85" box="[926,1021,294,319]" class="Insecta" family="Staphylinidae" genus="Liogluta" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="303" phylum="Arthropoda" rank="genus">
<emphasis id="ECBA582EE848FFC76392FEFDFC53FE85" box="[926,1021,294,319]" italics="true" pageId="18" pageNumber="303">Liogluta</emphasis>
</taxonomicName>
is maintained here, pending further comprehensive research of this species.
</paragraph>
</subSubSection>
</treatment>
</document>