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<document id="62D149871873A09862435AC415522ADC" ID-CLB-Dataset="3376" ID-DOI="10.5852/ejt.2021.767.1481" ID-GBIF-Dataset="9b3e85f1-1856-49bc-8cde-b2d197897698" ID-ISSN="2118-9773" ID-Zenodo-Dep="5715526" ID-ZooBank="32E080F8-A27A-4CBB-BA48-044DF82335A9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637362086142" checkinUser="admin" docAuthor="Liang, Zulong, Angus, Robert B. &amp; Jia, Fenglong" docDate="2021" docId="56557D6EFFEAFFFE557ACE1D3347FE99" docLanguage="en" docName="ejt-767_1-39.pdf.imf" docOrigin="European Journal of Taxonomy 767" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Patrus hainanensis Liang &amp; Angus &amp; Jia 2021, sp. nov." docType="treatment" docUuid="1001055C-63EB-4EC7-A1E4-92FED32C3A58" docUuidSource="ZooBank" docVersion="7" lastPageNumber="8" masterDocId="AA6C0516FFE8FFF95709CD48375DFFC8" masterDocTitle="Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae)" masterLastPageNumber="39" masterPageNumber="1" pageNumber="3" updateTime="1699469142537" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="856E86247C7259437BD4F24BDFFAD3E2">Three new species of Patrus Aubé with additional records of Gyrinidae from China (Coleoptera, Gyrinidae)</mods:title>
</mods:titleInfo>
<mods:name id="1DFFCF5AE53F5B8C00ADEB50685D08CC" type="personal">
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<mods:namePart id="DABCB9F2260D05DCB9B1576ABB9FA5BC">Liang, Zulong</mods:namePart>
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<mods:affiliation id="CC0287E20BD36AF5D74FC31BFC0D6434">Institute of Entomology, Life Science School, Sun Yat-sen University, Guangzhou, 510275, Guangdong, China. Department of Life Sciences (Insects), The Natural History Museum, Cromwell Road, London SW 7 5 BD, UK.</mods:affiliation>
<mods:nameIdentifier id="23CF003E72C038691B13626B61551FF2" type="email">liangzl3@mail2.sysu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="3D0680E376DCF77A59EF2D5D7C070993">Angus, Robert B.</mods:namePart>
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<mods:namePart id="60AF7E8ED60977D4BD0501FF75824095">Jia, Fenglong</mods:namePart>
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<mods:title id="E696CEB7D475AD56F5D0B0DE422A011F">European Journal of Taxonomy</mods:title>
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<mods:date id="9331093C8830BB64EE9807280B9D0B7A">2021</mods:date>
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<mods:number id="EBC6CCF76D935F6D08D8D8AFEF67198F">2021-09-02</mods:number>
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<treatment id="56557D6EFFEAFFFE557ACE1D3347FE99" ID-DOI="http://doi.org/10.5281/zenodo.5715633" ID-GBIF-Taxon="190335232" ID-Zenodo-Dep="5715633" LSID="urn:lsid:zoobank.org:act:1001055C-63EB-4EC7-A1E4-92FED32C3A58" httpUri="http://treatment.plazi.org/id/56557D6EFFEAFFFE557ACE1D3347FE99" lastPageId="7" lastPageNumber="8" pageId="2" pageNumber="3">
<subSubSection id="96E69FF3FFEAFFFB557ACE1D3398FC5B" pageId="2" pageNumber="3" type="nomenclature">
<paragraph id="DE43CC78FFEAFFFB557ACE1D349CFCB8" blockId="2.[366,1221,853,950]" box="[627,961,853,880]" pageId="2" pageNumber="3">
<heading id="850B7B14FFEAFFFB557ACE1D349CFCB8" box="[627,961,853,880]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="3" reason="2">
<taxonomicName id="19FCB7FBFFEAFFFB557ACE1D3402FCB8" authority="Liang &amp; Angus &amp; Jia, 2021" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[627,863,853,880]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="EC88106AFFEAFFFB557ACE1D3402FCB8" bold="true" box="[627,863,853,880]" italics="true" pageId="2" pageNumber="3">Patrus hainanensis</emphasis>
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<taxonomicNameLabel id="F7BBAD11FFEAFFFB546ECE1E349CFCB8" box="[871,961,854,880]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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<subSubSection id="96E69FF3FFEAFFFB55BACED334DFFC7D" box="[691,898,923,950]" pageId="2" pageNumber="3" type="description">
<paragraph id="DE43CC78FFEAFFFB55BACED334DFFC7D" blockId="2.[366,1221,853,950]" box="[691,898,923,950]" pageId="2" pageNumber="3">
<figureCitation id="46C7D0FDFFEAFFFB55BACED33468FC7E" box="[691,821,923,950]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[189,1398,359,1889]" captionTargetId="figure-21@4.[189,1398,359,1889]" captionTargetPageId="4" captionText="Fig. 1. Habitus. AC. Patrus hainanensis sp. nov., ♂, paratype (SYSU). DF. Patrus jiangxiensis. sp. nov., ♂, paratypes (SYSU). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5715528" httpUri="https://zenodo.org/record/5715528/files/figure.png" pageId="2" pageNumber="3">Figs 1AC</figureCitation>
,
<figureCitation id="46C7D0FDFFEAFFFB5448CED4340CFC7E" box="[833,849,924,950]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="2" pageNumber="3">3</figureCitation>
,
<figureCitation id="46C7D0FDFFEAFFFB5456CED334DFFC7D" box="[863,898,923,949]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[204,1379,1111,1818]" captionTargetId="figure-216@11.[189,1398,1111,1818]" captionTargetPageId="11" captionText="Fig. 6. Female reproductive tract of new species, paratype with the same collection data as for holotype (SYSU). A. Patrus hainanensis sp. nov. B. Patrus jiangxiensis sp. nov. C. Patrus shangchuanensis sp. nov.Abbreviations: AG = accessory gland; BC = bursa copulatrix; BS = bursa sclerite; CO = common oviduct; FD = fertilization duct; GC = gonocoxa; LT = laterogtergite; SP = spermatheca; VA = vagina. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.5715542" httpUri="https://zenodo.org/record/5715542/files/figure.png" pageId="2" pageNumber="3">6A</figureCitation>
</paragraph>
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<subSubSection id="96E69FF3FFEAFFFB57B4CE95336EF9BF" pageId="2" pageNumber="3" type="diagnosis">
<paragraph id="DE43CC78FFEAFFFB57B4CE95366AFC3F" blockId="2.[189,311,989,1015]" box="[189,311,989,1015]" pageId="2" pageNumber="3">
<heading id="850B7B14FFEAFFFB57B4CE95366AFC3F" bold="true" box="[189,311,989,1015]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="EC88106AFFEAFFFB57B4CE95366AFC3F" bold="true" box="[189,311,989,1015]" pageId="2" pageNumber="3">Diagnosis</emphasis>
</heading>
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<paragraph id="DE43CC78FFEAFFFB57B4C94335CDFA61" blockId="2.[189,1400,1034,1449]" pageId="2" pageNumber="3">
This species can be diagnosed by the following characters combined: 1) labrum less than ⅓ as long as wide, with anterior margin straight; 2) pronotal setose region anteriorly expanded inward onto pronotal disc to middle of dorsal eye; 3) punctation absent on head and pronotum; 4) punctures on surface of elytral glabrous region elongate transverse or oblique, several narrow longitudinal stripes shown on elytral glabrous region, micro-reticulation on elytral glabrous region weak and subtle, very elongate (
<figureCitation id="46C7D0FDFFEAFFFB57CFC9F33614FB1E" box="[198,329,1211,1238]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="2" pageNumber="3">Fig. 3A, G</figureCitation>
); 5) in lateral view elytral pubescent area in basal ½ distinctly narrower than on pronotum, expanded to similar width apically (
<figureCitation id="46C7D0FDFFEAFFFB556DC99735E2FB31" box="[612,703,1247,1273]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[189,1398,359,1889]" captionTargetId="figure-21@4.[189,1398,359,1889]" captionTargetPageId="4" captionText="Fig. 1. Habitus. AC. Patrus hainanensis sp. nov., ♂, paratype (SYSU). DF. Patrus jiangxiensis. sp. nov., ♂, paratypes (SYSU). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5715528" httpUri="https://zenodo.org/record/5715528/files/figure.png" pageId="2" pageNumber="3">Fig. 1C</figureCitation>
); 6) elytral apex rounded, epipleural angle acute; 7) female elytra similar to those of male; 8) male protarsus as in
<figureCitation id="46C7D0FDFFEAFFFB545CC84A3489FAD4" box="[853,980,1282,1308]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="2" pageNumber="3">Fig. 3BC</figureCitation>
, adhesive pallet with few adhesive setae and large suction discs, ⅔ length of protibia; 9) aedeagus as in
<figureCitation id="46C7D0FDFFEAFFFB531EC86D3328FAF7" box="[1047,1141,1317,1343]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="2" pageNumber="3">Fig. 3D</figureCitation>
, median lobe shorter than parameres, extending to ¾ of their length, strongly laterally expanded in apical ⅓; 10) female reproductive tract as in
<figureCitation id="46C7D0FDFFEAFFFB56DDC823356FFA4D" box="[468,562,1387,1413]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[204,1379,1111,1818]" captionTargetId="figure-216@11.[189,1398,1111,1818]" captionTargetPageId="11" captionText="Fig. 6. Female reproductive tract of new species, paratype with the same collection data as for holotype (SYSU). A. Patrus hainanensis sp. nov. B. Patrus jiangxiensis sp. nov. C. Patrus shangchuanensis sp. nov.Abbreviations: AG = accessory gland; BC = bursa copulatrix; BS = bursa sclerite; CO = common oviduct; FD = fertilization duct; GC = gonocoxa; LT = laterogtergite; SP = spermatheca; VA = vagina. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.5715542" httpUri="https://zenodo.org/record/5715542/files/figure.png" pageId="2" pageNumber="3">Fig. 6A</figureCitation>
, spermatheca elongate, reniform; fertilization duct broad and flattened, forming several compact hairpin hoops.
</paragraph>
<paragraph id="DE43CC78FFEAFFFB57B4C898336EF9BF" blockId="2.[187,1399,1488,1655]" pageId="2" pageNumber="3">
This species is a member of Group
<quantity id="1904619DFFEAFFFB5559C8983526FA22" box="[592,635,1488,1514]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="3" unit="in" value="2.0">2 in</quantity>
Vaziranis (1984) key (Group V in the key of
<bibRefCitation id="BA6DB189FFEAFFFB538CC898325BFA22" author="Ochs" box="[1157,1286,1488,1514]" firstAuthor="Ochs" journalOrPublisher="Entomologische Blatter" pageId="2" pageNumber="3" pagination="41 - 45" part="24" refId="ref17189" refString="Ochs G. 1928. Uber die Gyriniden Borneo's. Entomologische Blatter 24: 41 - 45, 78 - 85, 93 - 103." title="Uber die Gyriniden Borneo's" type="journal article" year="1928">Ochs 1928</bibRefCitation>
). The few adhesive setae and large suction discs on the male protarsus and characteristic elytral surface sculpture can easily distinguish this species from other known Chinese species of this group. This species is similar to
<taxonomicName id="19FCB7FBFFEAFFFB563ACB72352BF99C" authority="(Peschet, 1925)" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" baseAuthorityName="Peschet" baseAuthorityYear="1925" box="[307,630,1593,1620]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="coomani">
<emphasis id="EC88106AFFEAFFFB563ACB7236EBF99B" box="[307,438,1594,1619]" italics="true" pageId="2" pageNumber="3">P. coomani</emphasis>
(
<bibRefCitation id="BA6DB189FFEAFFFB56CCCB723530F99C" author="Peschet" box="[453,621,1593,1620]" firstAuthor="Peschet" pageId="2" pageNumber="3" pagination="253 - 254" refId="ref17414" refString="Peschet R. 1925. Description d'un Orectochilus nouveau du Tonkin [Col. Gyrinidae]. Bulletin de la Societe entomologique de France 30 (15): 253 - 254." type="journal article" year="1925">Peschet, 1925</bibRefCitation>
)
</taxonomicName>
, but can still be easily seperated by its narrow setose region at the elytral base and by the median lobe strongly laterally expanded in apical ⅓.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFEAFFFB57B4CBD634F8F92F" box="[189,933,1694,1767]" pageId="2" pageNumber="3" type="etymology">
<paragraph id="DE43CC78FFEAFFFB57B4CBD63618F970" blockId="2.[189,325,1694,1720]" box="[189,325,1694,1720]" pageId="2" pageNumber="3">
<heading id="850B7B14FFEAFFFB57B4CBD63618F970" bold="true" box="[189,325,1694,1720]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="EC88106AFFEAFFFB57B4CBD63618F970" bold="true" box="[189,325,1694,1720]" pageId="2" pageNumber="3">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFEAFFFB57B4CB8534F8F92F" blockId="2.[189,933,1741,1767]" box="[189,933,1741,1767]" pageId="2" pageNumber="3">
This species is named after the
<typeStatus id="014772DAFFEAFFFB5524CB85353DF92F" box="[557,608,1741,1767]" pageId="2" pageNumber="3">type</typeStatus>
locality ʻHainan Provinceʼ.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFEAFFFA57B4CA463265FC54" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="DE43CC78FFEAFFFB57B4CA4636F0F8E0" blockId="2.[189,429,1806,1832]" box="[189,429,1806,1832]" pageId="2" pageNumber="3">
<heading id="850B7B14FFEAFFFB57B4CA4636F0F8E0" bold="true" box="[189,429,1806,1832]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="EC88106AFFEAFFFB57B4CA4636F0F8E0" bold="true" box="[189,429,1806,1832]" pageId="2" pageNumber="3">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFEAFFFB57E3CA753600F89F" blockId="2.[189,1399,1853,1950]" box="[234,349,1853,1879]" pageId="2" pageNumber="3">
<emphasis id="EC88106AFFEAFFFB57E3CA753600F89F" bold="true" box="[234,349,1853,1879]" pageId="2" pageNumber="3">
<typeStatus id="014772DAFFEAFFFB57E3CA753600F89F" box="[234,349,1853,1879]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DE43CC78FFEAFFFB57B4CA28351CF855" blockId="2.[189,1399,1853,1950]" pageId="2" pageNumber="3">
<materialsCitation id="6E94C625FFEAFFFB57B4CA28351CF855" ID-GBIF-Occurrence="3409120336" collectingDate="1981-07-08" collectorName="Hua Lizhong" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="holotype">
<collectingCountry id="A6EB8CE8FFEAFFFB57B4CA283645F8B2" box="[189,280,1888,1914]" name="China" pageId="2" pageNumber="3">CHINA</collectingCountry>
<emphasis id="EC88106AFFEAFFFB563DCA2836CFF8B2" bold="true" box="[308,402,1888,1914]" pageId="2" pageNumber="3">
<collectingRegion id="1C38029AFFEAFFFB563DCA2836CFF8B2" box="[308,402,1888,1914]" country="China" name="Hainan" pageId="2" pageNumber="3">Hainan</collectingRegion>
</emphasis>
<specimenCount id="C8FA07F1FFEAFFFB56A3CA29369EF8B3" box="[426,451,1889,1915]" pageId="2" pageNumber="3" type="male"></specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEAFFFB55C1CA28347EF8B2" box="[712,803,1888,1914]" pageId="2" pageNumber="3">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEAFFFB5427CA2834AFF8B3" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEAFFFB5427CA2834AFF8B3" box="[814,1010,1888,1915]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="2" pageNumber="3" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEAFFFB53A5CA28326DF8B2" box="[1196,1328,1888,1914]" pageId="2" pageNumber="3" value="1981-07-08">
<collectingDate id="BA061350FFEAFFFB53A5CA28326DF8B2" box="[1196,1328,1888,1914]" pageId="2" pageNumber="3" value="1981-07-08">8 Jul. 1981</collectingDate>
</date>
; 华立 中采 [
<collectorName id="7309A9AEFFEAFFFB560FCACC36FEF855" box="[262,419,1923,1950]" pageId="2" pageNumber="3">Hua Lizhong</collectorName>
leg.]; SYSU.
</materialsCitation>
</paragraph>
<paragraph id="DE43CC78FFEBFFFA57E3CC413634FEEB" blockId="3.[188,1399,265,925]" box="[234,361,265,291]" pageId="3" pageNumber="4">
<emphasis id="EC88106AFFEBFFFA57E3CC413634FEEB" bold="true" box="[234,361,265,291]" pageId="3" pageNumber="4">
<typeStatus id="014772DAFFEBFFFA57E3CC413634FEEB" box="[234,361,265,291]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
</emphasis>
</paragraph>
<paragraph id="DE43CC78FFEBFFFA57B4CC643265FC54" blockId="3.[188,1399,265,925]" pageId="3" pageNumber="4">
<materialsCitation id="6E94C625FFEBFFFA57B4CC643305FE8E" ID-GBIF-Occurrence="3409120330" box="[189,1112,299,326]" collectingDate="1981-07-08" collectorName="Hua Lizhong" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="7" specimenCount-female="2" specimenCount-male="5" stateProvince="Hainan" typeStatus="paratype">
<collectingCountry id="A6EB8CE8FFEBFFFA57B4CC643645FE8E" box="[189,280,300,326]" name="China" pageId="3" pageNumber="4">CHINA</collectingCountry>
<emphasis id="EC88106AFFEBFFFA563ECC6336C8FE8D" bold="true" box="[311,405,299,325]" pageId="3" pageNumber="4">
<collectingRegion id="1C38029AFFEBFFFA563ECC6336C8FE8D" box="[311,405,299,325]" country="China" name="Hainan" pageId="3" pageNumber="4">Hainan</collectingRegion>
</emphasis>
<specimenCount id="C8FA07F1FFEBFFFA56A6CC6336AEFE8E" box="[431,499,299,326]" pageId="3" pageNumber="4" type="male">5 ♂♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA5508CC643519FE8E" box="[513,580,300,326]" pageId="3" pageNumber="4" type="female">2 ♀♀</specimenCount>
; same collection data as for holotype; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA537BCC64322BFEA1" ID-GBIF-Occurrence="3409120353" collectingDate="1981-07-05" collectorName="Zhang Xinfu" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA537BCC6433FCFE8E" box="[1138,1185,300,326]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA53A7CC6433AFFE8E" box="[1198,1266,300,326]" pageId="3" pageNumber="4" type="female">2 ♀♀</specimenCount>
; 海南尖峰 岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA5648CC0736C1FEA1" box="[321,412,335,361]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA56AECC073531FEA1" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA56AECC073531FEA1" box="[423,620,335,361]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division,];
<date id="AA42EAB8FFEBFFFA5439CC0734E8FEA1" box="[816,949,335,362]" pageId="3" pageNumber="4" value="1981-07-05">
<collectingDate id="BA061350FFEBFFFA5439CC0734E8FEA1" box="[816,949,335,362]" pageId="3" pageNumber="4" value="1981-07-05">5 Jul. 1981</collectingDate>
</date>
; 张新府采 [
<collectorName id="7309A9AEFFEBFFFA534CCC073382FEA1" box="[1093,1247,335,361]" pageId="3" pageNumber="4">Zhang Xinfu</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA57D9CC3A3613FE67" ID-GBIF-Occurrence="3409120429" collectingDate="1981-07-07" collectorName="Huang Yun" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA57D9CC3A365DFE45" box="[208,256,370,397]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA550ECC3A353FFE44" box="[519,610,370,396]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA5567CC3A3469FE45" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA5567CC3A3469FE45" box="[622,820,370,397]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA54FACC3B3327FE44" box="[1011,1146,370,397]" pageId="3" pageNumber="4" value="1981-07-07">
<collectingDate id="BA061350FFEBFFFA54FACC3B3327FE44" box="[1011,1146,370,397]" pageId="3" pageNumber="4" value="1981-07-07">7 Jul. 1981</collectingDate>
</date>
; 黄云采 [
<collectorName id="7309A9AEFFEBFFFA53E6CC3B322BFE45" box="[1263,1398,371,397]" pageId="3" pageNumber="4">Huang Yun</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA5662CCDD35DBFE1B" ID-GBIF-Occurrence="3409120316" collectingDate="1981-07-07" collectorName="Chen Ningmei" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA5662CCDD36C6FE78" box="[363,411,405,432]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA56A0CCDD36B3FE78" box="[425,494,405,432]" pageId="3" pageNumber="4" type="female">3 ♀♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA55F0CCDD3409FE67" box="[761,852,405,431]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA5468CCDD337AFE78" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA5468CCDD337AFE78" box="[865,1063,405,432]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA53E0CCDE322FFE67" box="[1257,1394,405,432]" pageId="3" pageNumber="4" value="1981-07-07">
<collectingDate id="BA061350FFEBFFFA53E0CCDE322FFE67" box="[1257,1394,405,432]" pageId="3" pageNumber="4" value="1981-07-07">7 Jul. 1981</collectingDate>
</date>
; 陈宁美采 [
<collectorName id="7309A9AEFFEBFFFA5649CCF036ADFE1A" box="[320,496,440,466]" pageId="3" pageNumber="4">Chen Ningmei</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA5596CCF13438FE3E" ID-GBIF-Occurrence="3409120363" collectingDate="1981-07-07" collectorName="Liu Yuanfu" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA5596CCF13590FE1B" box="[671,717,441,467]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA55D3CCF1345AFE1B" box="[730,775,441,467]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA5307CCF03334FE1A" box="[1038,1129,440,466]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA537CCCF03264FE1B" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA537CCCF03264FE1B" box="[1141,1337,440,467]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA563CCC9436E4FE3E" box="[309,441,476,502]" pageId="3" pageNumber="4" value="1981-07-07">
<collectingDate id="BA061350FFEBFFFA563CCC9436E4FE3E" box="[309,441,476,502]" pageId="3" pageNumber="4" value="1981-07-07">7 Jul. 1981</collectingDate>
</date>
; 刘元福采 [
<collectorName id="7309A9AEFFEBFFFA5541CC943592FE3E" box="[584,719,476,502]" pageId="3" pageNumber="4">Liu Yuanfu</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA5477CC9433DAFDD1" ID-GBIF-Occurrence="3409120338" collectingDate="1981-07-07" collectorName="Li Zongran" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="17" specimenCount-female="8" specimenCount-male="9" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA5477CC94349CFE3E" box="[894,961,476,502]" pageId="3" pageNumber="4" type="male">9 ♂♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA54C7CC94334CFE3E" box="[974,1041,476,502]" pageId="3" pageNumber="4" type="female">8 ♀♀</specimenCount>
; 海南尖峰岭三分区 [
<collectingMunicipality id="3E275602FFEBFFFA521ECC94322FFE3E" box="[1303,1394,476,502]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA57B4CCB736DEFDD1" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA57B4CCB736DEFDD1" box="[189,387,511,537]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, third division];
<date id="AA42EAB8FFEBFFFA5543CCB7358CFDD1" box="[586,721,511,537]" pageId="3" pageNumber="4" value="1981-07-07">
<collectingDate id="BA061350FFEBFFFA5543CCB7358CFDD1" box="[586,721,511,537]" pageId="3" pageNumber="4" value="1981-07-07">7 Jul. 1981</collectingDate>
</date>
; 李宗冉采 [
<collectorName id="7309A9AEFFEBFFFA546DCCB734B0FDD1" box="[868,1005,511,537]" pageId="3" pageNumber="4">Li Zongran</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA53AACCB73613FD97" ID-GBIF-Occurrence="3409120339" collectingDate="1981-07-07" collectorName="Liang Shaoying" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="16" specimenCount-female="6" specimenCount-male="10" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA53AACCB733AAFDD1" box="[1187,1271,511,537]" pageId="3" pageNumber="4" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA520CCCB73214FDD1" box="[1285,1353,511,537]" pageId="3" pageNumber="4" type="female">6 ♀♀</specimenCount>
; 海 南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA5695CF6A36AAFDF4" box="[412,503,546,572]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA550FCF6A3590FDF5" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA550FCF6A3590FDF5" box="[518,717,546,573]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA549DCF6B3342FDF4" box="[916,1055,546,573]" pageId="3" pageNumber="4" value="1981-07-07">
<collectingDate id="BA061350FFEBFFFA549DCF6B3342FDF4" box="[916,1055,546,573]" pageId="3" pageNumber="4" value="1981-07-07">7 Jul. 1981</collectingDate>
</date>
; 梁少营采 [
<collectorName id="7309A9AEFFEBFFFA53BCCF6A322AFDF4" box="[1205,1399,546,572]" pageId="3" pageNumber="4">Liang Shaoying</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA5662CF0D352AFD4B" ID-GBIF-Occurrence="3409120390" collectingDate="1981-07-08" collectorName="He Guofeng" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="6" specimenCount-female="5" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA5662CF0D36C6FDA8" box="[363,411,581,608]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA56A0CF0D36B3FDA8" box="[425,494,581,608]" pageId="3" pageNumber="4" type="female">5 ♀♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA55F0CF0D3409FD97" box="[761,852,581,607]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA5468CF0D337AFDA8" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA5468CF0D337AFDA8" box="[865,1063,581,608]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA53E0CF0D322FFD97" box="[1257,1394,581,607]" pageId="3" pageNumber="4" value="1981-07-08">
<collectingDate id="BA061350FFEBFFFA53E0CF0D322FFD97" box="[1257,1394,581,607]" pageId="3" pageNumber="4" value="1981-07-08">8 Jul. 1981</collectingDate>
</date>
; 何国锋采 [
<collectorName id="7309A9AEFFEBFFFA564DCF213687FD4A" box="[324,474,616,643]" pageId="3" pageNumber="4">He Guofeng</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA559FCF21349EFD6E" ID-GBIF-Occurrence="3409120304" collectingDate="1981-07-08" collectorName="Lei Qunbang" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="10" specimenCount-female="6" specimenCount-male="4" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA559FCF213580FD4B" box="[662,733,617,643]" pageId="3" pageNumber="4" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA55E4CF20346EFD4B" box="[749,819,616,643]" pageId="3" pageNumber="4" type="female">6 ♀♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA5348CF2033C1FD4A" box="[1089,1180,616,642]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA53A3CF20322FFD4B" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA53A3CF20322FFD4B" box="[1194,1394,616,643]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA5679CFC436A5FD6E" box="[368,504,652,678]" pageId="3" pageNumber="4" value="1981-07-08">
<collectingDate id="BA061350FFEBFFFA5679CFC436A5FD6E" box="[368,504,652,678]" pageId="3" pageNumber="4" value="1981-07-08">8 Jul. 1981</collectingDate>
</date>
; 雷群邦采 [
<collectorName id="7309A9AEFFEBFFFA5582CFC43474FD6E" box="[651,809,652,678]" pageId="3" pageNumber="4">Lei Qunbang</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA54D6CFC433DEFD01" ID-GBIF-Occurrence="3409120466" collectingDate="1981-07-08" collectorName="Li Yizhao" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="1" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA54D6CFC43352FD6E" box="[991,1039,652,678]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA521ECFC4322FFD6E" box="[1303,1394,652,678]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA57B7CFE736DBFD01" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA57B7CFE736DBFD01" box="[190,390,687,713]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA5545CFE73585FD01" box="[588,728,687,713]" pageId="3" pageNumber="4" value="1981-07-08">
<collectingDate id="BA061350FFEBFFFA5545CFE73585FD01" box="[588,728,687,713]" pageId="3" pageNumber="4" value="1981-07-08">8 Jul. 1981</collectingDate>
</date>
; 李逸昭采 [
<collectorName id="7309A9AEFFEBFFFA5464CFE734BBFD01" box="[877,998,687,713]" pageId="3" pageNumber="4">Li Yizhao</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA53AACFE7322BFD24" ID-GBIF-Occurrence="3409120397" collectingDate="1981-07-08" collectorName="Chen Runji" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA53AACFE7338FFD01" box="[1187,1234,687,713]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
, 海南尖峰岭 五分区 [
<collectingMunicipality id="3E275602FFEBFFFA562ECF9A36DFFD24" box="[295,386,722,748]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA569BCF9A3506FD25" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA569BCF9A3506FD25" box="[402,603,722,749]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA542DCF9A34ECFD24" box="[804,945,722,748]" pageId="3" pageNumber="4" value="1981-07-08">
<collectingDate id="BA061350FFEBFFFA542DCF9A34ECFD24" box="[804,945,722,748]" pageId="3" pageNumber="4" value="1981-07-08">8 Jul. 1981</collectingDate>
</date>
; 陈润吉采 [
<collectorName id="7309A9AEFFEBFFFA5340CF9A338AFD24" box="[1097,1239,722,748]" pageId="3" pageNumber="4">Chen Runji</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA57DBCFBD3509FCFB" ID-GBIF-Occurrence="3409120370" collectingDate="1983-07-29" collectorName="Chen Huanqiang" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA57DBCFBD3644FCD8" box="[210,281,757,784]" pageId="3" pageNumber="4" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA5620CFBD3632FCD8" box="[297,367,757,784]" pageId="3" pageNumber="4" type="female">2 ♀♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA5574CFBD3585FCC7" box="[637,728,757,783]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA55EECFBD34F2FCD8" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA55EECFBD34F2FCD8" box="[743,943,757,784]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA537CCFBD3252FCC7" box="[1141,1295,757,783]" pageId="3" pageNumber="4" value="1983-07-29">
<collectingDate id="BA061350FFEBFFFA537CCFBD3252FCC7" box="[1141,1295,757,783]" pageId="3" pageNumber="4" value="1983-07-29">29 Jul. 1983</collectingDate>
</date>
; 陈焕强 采 [
<collectorName id="7309A9AEFFEBFFFA57E2CE5036EAFCFA" box="[235,439,792,818]" pageId="3" pageNumber="4">Chen Huanqiang</collectorName>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA557BCE513423FC9E" ID-GBIF-Occurrence="3409120357" collectingDate="1983-07-29" collectorName="Jiang Shigui" country="China" location="Jianfengling Mt." municipality="Hainan" pageId="3" pageNumber="4" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA557BCE5135FEFCFB" box="[626,675,793,819]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA55BACE5135A4FCFB" box="[691,761,793,819]" pageId="3" pageNumber="4" type="female">3 ♀♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingMunicipality id="3E275602FFEBFFFA530FCE50333CFCFA" box="[1030,1121,792,818]" pageId="3" pageNumber="4">Hainan</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA5366CE50326AFCFB" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA5366CE50326AFCFB" box="[1135,1335,792,819]" country="China" municipality="Hainan" name="Jianfengling Mt." pageId="3" pageNumber="4" stateProvince="Hainan">Jianfengling Mt.</location>
, fifth division];
<date id="AA42EAB8FFEBFFFA563DCE74369BFC9E" box="[308,454,828,854]" pageId="3" pageNumber="4" value="1983-07-29">
<collectingDate id="BA061350FFEBFFFA563DCE74369BFC9E" box="[308,454,828,854]" pageId="3" pageNumber="4" value="1983-07-29">29 Jul. 1983</collectingDate>
</date>
; 江世贵采 [
<collectorName id="7309A9AEFFEBFFFA555CCE7435B4FC9E" box="[597,745,828,854]" pageId="3" pageNumber="4">Jiang Shigui</collectorName>
leg,]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA549FCE7433F1FCB1" ID-GBIF-Occurrence="3409120438" collectingDate="1983-09-08" country="China" county="Hainan" location="Reserve station" municipality="Jianfengling Mt." pageId="3" pageNumber="4" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA549FCE74349EFC9E" box="[918,963,828,854]" pageId="3" pageNumber="4" type="male">1 ♂</specimenCount>
,
<specimenCount id="C8FA07F1FFEBFFFA54C6CE74334CFC9E" box="[975,1041,828,854]" pageId="3" pageNumber="4" type="female">3 ♀♀</specimenCount>
; 海南尖峰岭五分区 [
<collectingCounty id="3722B4F4FFEBFFFA521ECE74322FFC9E" box="[1303,1394,828,854]" pageId="3" pageNumber="4">Hainan</collectingCounty>
,
<collectingMunicipality id="3E275602FFEBFFFA57B5CE1736DCFCB1" box="[188,385,863,889]" pageId="3" pageNumber="4">Jianfengling Mt.</collectingMunicipality>
, fifth division];
<date id="AA42EAB8FFEBFFFA5537CE173590FCB1" box="[574,717,863,889]" pageId="3" pageNumber="4" value="1983-09-08">
<collectingDate id="BA061350FFEBFFFA5537CE173590FCB1" box="[574,717,863,889]" pageId="3" pageNumber="4" value="1983-09-08">8 Sep. 1983</collectingDate>
</date>
; 保护站采 [
<location id="DB239AA3FFEBFFFA5454CE173349FCB1" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA5454CE173349FCB1" box="[861,1044,863,889]" country="China" county="Hainan" municipality="Jianfengling Mt." name="Reserve station" pageId="3" pageNumber="4" stateProvince="Hainan">Reserve station</location>
leg.]; SYSU
</materialsCitation>
<materialsCitation id="6E94C625FFEBFFFA53CCCE173269FC54" ID-GBIF-Occurrence="3409120422" collectingDate="1983-09-08" country="China" county="Hainan" location="Tianchi" municipality="Jianfengling Mt." pageId="3" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Hainan" typeStatus="paratype">
<specimenCount id="C8FA07F1FFEBFFFA53CCCE1733AFFCB1" box="[1221,1266,863,889]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
; 海南尖峰 岭天池 [
<collectingCounty id="3722B4F4FFEBFFFA562ACECA3623FC54" box="[291,382,898,924]" pageId="3" pageNumber="4">Hainan</collectingCounty>
,
<collectingMunicipality id="3E275602FFEBFFFA5680CECA3513FC55" box="[393,590,898,925]" pageId="3" pageNumber="4">Jianfengling Mt.</collectingMunicipality>
,
<location id="DB239AA3FFEBFFFA5550CEC935E5FC54" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA5550CEC935E5FC54" box="[601,696,897,924]" country="China" county="Hainan" municipality="Jianfengling Mt." name="Tianchi" pageId="3" pageNumber="4" stateProvince="Hainan">Tianchi</location>
];
<date id="AA42EAB8FFEBFFFA55C2CECA3407FC54" box="[715,858,898,924]" pageId="3" pageNumber="4" value="1983-09-08">
<collectingDate id="BA061350FFEBFFFA55C2CECA3407FC54" box="[715,858,898,924]" pageId="3" pageNumber="4" value="1983-09-08">8 Sep. 1983</collectingDate>
</date>
; 保护站采 [
<location id="DB239AA3FFEBFFFA54E0CECB33FDFC54" LSID="urn:lsid:plazi:treatment:56557D6EFFEAFFFE557ACE1D3347FE99:DB239AA3FFEBFFFA54E0CECB33FDFC54" box="[1001,1184,898,925]" country="China" county="Hainan" municipality="Jianfengling Mt." name="Reserve station" pageId="3" pageNumber="4" stateProvince="Hainan">Reserve station</location>
leg]; SYSU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="96E69FF3FFEBFFFF57B4CE8B3648FCCE" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4" type="description">
<paragraph id="DE43CC78FFEBFFFA57B4CE8B3612FC15" blockId="3.[189,335,963,989]" box="[189,335,963,989]" pageId="3" pageNumber="4">
<heading id="850B7B14FFEBFFFA57B4CE8B3612FC15" bold="true" box="[189,335,963,989]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
<emphasis id="EC88106AFFEBFFFA57B4CE8B3612FC15" bold="true" box="[189,335,963,989]" pageId="3" pageNumber="4">Description</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFEBFFFA57B4CEBB34C9FBE7" blockId="3.[188,1399,1010,1072]" pageId="3" pageNumber="4">
MEASUREMENTS.
<typeStatus id="014772DAFFEBFFFA5675CEBA36B3FBC4" box="[380,494,1010,1036]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
: 7.0 mm in length, 3.0 mm in width; male: 6.99.0 mm in length, 3.0
<quantity id="1904619DFFEBFFFA5216CEBA322AFBC5" box="[1311,1399,1010,1037]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="4" unit="mm" value="3.8">3.8 mm</quantity>
in width; female: 7.0
<quantity id="1904619DFFEBFFFA56C8C95D3544FBF8" box="[449,537,1045,1072]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="3" pageNumber="4" unit="mm" value="8.5">8.5 mm</quantity>
in length,
<quantity id="1904619DFFEBFFFA559FC95D3479FBF8" box="[662,804,1045,1072]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" metricValueMax="3.4" metricValueMin="3.2" pageId="3" pageNumber="4" unit="mm" value="3.3" valueMax="3.4" valueMin="3.2">3.23.4 mm</quantity>
in width.
</paragraph>
<paragraph id="DE43CC78FFEBFFFA57B4C91F34BBFB12" blockId="3.[188,1399,1111,1243]" pageId="3" pageNumber="4">
HABITUS AND COLOUR (
<figureCitation id="46C7D0FDFFEBFFFA56C4C91F3509FBB9" box="[461,596,1111,1137]" captionStart="Fig" captionStartId="4.[189,232,1928,1954]" captionTargetBox="[189,1398,359,1889]" captionTargetId="figure-21@4.[189,1398,359,1889]" captionTargetPageId="4" captionText="Fig. 1. Habitus. AC. Patrus hainanensis sp. nov., ♂, paratype (SYSU). DF. Patrus jiangxiensis. sp. nov., ♂, paratypes (SYSU). A. Dorsal view. B. Ventral view. C. Lateral view. D. Dorsal view. E. Ventral view. F. Lateral view. Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5715528" httpUri="https://zenodo.org/record/5715528/files/figure.png" pageId="3" pageNumber="4">Fig. 1AC</figureCitation>
). Medium-sized species, oblong-oval in shape, widest just before middle, moderately convex in lateral view. Dorsal side black with strong bronze sheen, stronger on elytron than on head and pronotum. Pronotum and elytra with yellow lateral borders. Ventral side mostly black, with legs and rear of abdomen ferruginous, epipleuron yellow.
</paragraph>
<paragraph id="DE43CC78FFEBFFFA57B4C84A330AFA4D" blockId="3.[189,1398,1280,1414]" pageId="3" pageNumber="4">
HEAD. Labrum wide and short, ⅓ as long as wide, with anterior margin straight; anterior surface smooth; posterior surface with strong punctate-tomentose. Micro-reticulation on surface of clypeus distinct with polygonal meshes. Micro-reticulation on frons and vertex rather distinct, with meshes transverse or oblique (
<figureCitation id="46C7D0FDFFEBFFFA562CC8233622FA4D" box="[293,383,1387,1413]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="3" pageNumber="4">Fig. 3E</figureCitation>
), gradually obliterated backwards. Punctation absent on head.
</paragraph>
<paragraph id="DE43CC78FFEBFFFA57B4C8E5357BF8A5" blockId="3.[189,1400,1453,1901]" pageId="3" pageNumber="4">
THORAX. Pronotum regularly attenuated from base to apex, with margins weakly convex. Surface of pronotal glabrous region with weak and subtle micro-reticulation formed by elongate transverse or oblique meshes (
<figureCitation id="46C7D0FDFFEBFFFA5684C8BB36B6F9C5" box="[397,491,1523,1549]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="3" pageNumber="4">Fig. 3F</figureCitation>
); punctation absent on pronotum. Lateral pubescence anteriorly expanded inward onto pronotal disc to middle of dorsal eye, posteriorly narrower and parallel to lateral borders. Scutellum wide and short, ½ as long as wide, with surface smooth. Lateral pubescence on elytra anteriorly strongly reduced, distinctly narrower than on basal pronotum until middle of elytra, then abruptly expanded and reaching suture just before truncature (
<figureCitation id="46C7D0FDFFEBFFFA549DCBC834AFF952" box="[916,1010,1664,1690]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="3" pageNumber="4">Fig. 3A</figureCitation>
). Pronoto-elytral glabrous region irregular oval, terminating posteriorly with obtuse and wide angle. Surface of elytral glabrous region covered by distinct and rather dense punctation formed by elongate transverse or oblique strioles and with several narrow longitudinal stripes, micro-reticulation invisible under optical microscope, but visible under SEM, similar to that on pronotum (
<figureCitation id="46C7D0FDFFEBFFFA540ACA45343FF8EF" box="[771,866,1805,1831]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="3" pageNumber="4">Fig. 3G</figureCitation>
). Strioles at outer side of elytra much longer than that at inner side. Truncature of elytra straight; epipleural angle almost 90°, acute; sutural angle almost 90°, narrowly rounded.
</paragraph>
<caption id="8A839CF0FFECFFFD57B4CAC03479F822" ID-DOI="http://doi.org/10.5281/zenodo.5715528" ID-Zenodo-Dep="5715528" httpUri="https://zenodo.org/record/5715528/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1928,1954]" targetBox="[189,1398,359,1889]" targetPageId="4">
<paragraph id="DE43CC78FFECFFFD57B4CAC03479F822" blockId="4.[189,1399,1928,2026]" pageId="4" pageNumber="5">
<emphasis id="EC88106AFFECFFFD57B4CAC03652F86A" bold="true" box="[189,271,1928,1954]" pageId="4" pageNumber="5">Fig. 1.</emphasis>
Habitus.
<emphasis id="EC88106AFFECFFFD568ECAC0369CF86A" bold="true" box="[391,449,1928,1954]" pageId="4" pageNumber="5">AC</emphasis>
.
<taxonomicName id="19FCB7FBFFECFFFD56DACAC135E6F86B" authority="Liang &amp; Angus &amp; Jia, 2021" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[467,699,1929,1955]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="EC88106AFFECFFFD56DACAC135E6F86B" box="[467,699,1929,1955]" italics="true" pageId="4" pageNumber="5">Patrus hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BBAD11FFECFFFD55CFCAC1347AF86B" box="[710,807,1929,1955]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
, ♂, paratype (SYSU).
<emphasis id="EC88106AFFECFFFD5336CAC13328F86B" bold="true" box="[1087,1141,1929,1955]" pageId="4" pageNumber="5">DF</emphasis>
.
<taxonomicName id="19FCB7FBFFECFFFD538ECAC13232F86A" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[1159,1391,1929,1954]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="jiangxiensis">
<emphasis id="EC88106AFFECFFFD538ECAC13232F86A" box="[1159,1391,1929,1954]" italics="true" pageId="4" pageNumber="5">Patrus jiangxiensis</emphasis>
</taxonomicName>
. sp. nov., ♂, paratypes (SYSU).
<emphasis id="EC88106AFFECFFFD554FCAE43506F80E" bold="true" box="[582,603,1964,1990]" pageId="4" pageNumber="5">A</emphasis>
. Dorsal view.
<emphasis id="EC88106AFFECFFFD541ACAE4347AF80E" bold="true" box="[787,807,1964,1990]" pageId="4" pageNumber="5">B</emphasis>
. Ventral view.
<emphasis id="EC88106AFFECFFFD54EDCAE434A4F80E" bold="true" box="[996,1017,1964,1990]" pageId="4" pageNumber="5">C</emphasis>
. Lateral view.
<emphasis id="EC88106AFFECFFFD53BCCAE43397F80E" bold="true" box="[1205,1226,1964,1990]" pageId="4" pageNumber="5">D</emphasis>
. Dorsal view.
<emphasis id="EC88106AFFECFFFD57B4CA87378CF821" bold="true" box="[189,209,1999,2025]" pageId="4" pageNumber="5">E</emphasis>
. Ventral view.
<emphasis id="EC88106AFFECFFFD5689CA8736CFF821" bold="true" box="[384,402,1999,2025]" pageId="4" pageNumber="5">F</emphasis>
. Lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<caption id="8A839CF0FFEDFFFC57B4CAEE3322F82C" ID-DOI="http://doi.org/10.5281/zenodo.5715530" ID-Zenodo-Dep="5715530" httpUri="https://zenodo.org/record/5715530/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1958,1984]" targetBox="[189,1398,388,1918]" targetPageId="5">
<paragraph id="DE43CC78FFEDFFFC57B4CAEE3322F82C" blockId="5.[189,1398,1958,2020]" pageId="5" pageNumber="6">
<emphasis id="EC88106AFFEDFFFC57B4CAEE3656F808" bold="true" box="[189,267,1958,1984]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
Habitus of
<taxonomicName id="19FCB7FBFFEDFFFC569BCAEF35EFF808" authority="Liang &amp; Angus &amp; Jia, 2021" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[402,690,1958,1984]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="shangchuanensis" status="sp. nov.">
<emphasis id="EC88106AFFEDFFFC569BCAEF35EFF808" box="[402,690,1958,1984]" italics="true" pageId="5" pageNumber="6">Patrus shangchuanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BBAD11FFEDFFFC55B0CAEF3448F809" box="[697,789,1959,1985]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, paratypes (SYSU).
<emphasis id="EC88106AFFEDFFFC530DCAEE3344F808" bold="true" box="[1028,1049,1958,1984]" pageId="5" pageNumber="6">A</emphasis>
. ♂, dorsal view.
<emphasis id="EC88106AFFEDFFFC53D6CAEE33AEF808" bold="true" box="[1247,1267,1958,1984]" pageId="5" pageNumber="6">B</emphasis>
. ♂, ventral view.
<emphasis id="EC88106AFFEDFFFC560ACA813645F82B" bold="true" box="[259,280,1993,2019]" pageId="5" pageNumber="6">C</emphasis>
. ♂, lateral view.
<emphasis id="EC88106AFFEDFFFC56EBCA8136AAF82B" bold="true" box="[482,503,1993,2019]" pageId="5" pageNumber="6">D</emphasis>
. ♀, dorsal view.
<emphasis id="EC88106AFFEDFFFC55B6CA81358EF82B" bold="true" box="[703,723,1993,2019]" pageId="5" pageNumber="6">E</emphasis>
. ♀, lateral view. Scale bars = 1 mm.
</paragraph>
</caption>
<paragraph id="DE43CC78FFEEFFFF57B4CC4136D1FEA1" blockId="6.[189,1399,265,361]" pageId="6" pageNumber="7">
MALE GENITALIA (
<figureCitation id="46C7D0FDFFEEFFFF568ECC4136BAFEEB" box="[391,487,265,291]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="6" pageNumber="7">Fig. 3D</figureCitation>
). Median lobe shorter than parameres, extending ¾ of their length, basal half parallel, strongly laterally expanded in apical ⅓, apical part regularly narrowed toward apex, apex point shortly narrowed.
</paragraph>
<paragraph id="DE43CC78FFEEFFFF57B4CCD9358EFDDC" blockId="6.[187,1398,401,533]" pageId="6" pageNumber="7">
FEMALE REPRODUCTIVE TRACT (
<figureCitation id="46C7D0FDFFEEFFFF551ECCD9352AFE64" box="[535,631,401,428]" captionStart="Fig" captionStartId="11.[189,232,1857,1883]" captionTargetBox="[204,1379,1111,1818]" captionTargetId="figure-216@11.[189,1398,1111,1818]" captionTargetPageId="11" captionText="Fig. 6. Female reproductive tract of new species, paratype with the same collection data as for holotype (SYSU). A. Patrus hainanensis sp. nov. B. Patrus jiangxiensis sp. nov. C. Patrus shangchuanensis sp. nov.Abbreviations: AG = accessory gland; BC = bursa copulatrix; BS = bursa sclerite; CO = common oviduct; FD = fertilization duct; GC = gonocoxa; LT = laterogtergite; SP = spermatheca; VA = vagina. Scale bars = 500 μm." figureDoi="http://doi.org/10.5281/zenodo.5715542" httpUri="https://zenodo.org/record/5715542/files/figure.png" pageId="6" pageNumber="7">Fig. 6A</figureCitation>
). Gonocoxosternite elongate, apically narrowly rounded, anterior process elongate; laterotergite elongate, slightly curved apically; vagina elongate; spermatheca elongate, reniform; fertilization duct broad and flattened, forming several compact hairpin hoops; accessory gland short and slender, inserting dorsally on bursa.
</paragraph>
<paragraph id="DE43CC78FFEEFFFF57B4CF743648FCCE" blockId="6.[189,1401,572,774]" pageId="6" pageNumber="7">
SEXUAL DIMORPHISM. This species exhibits very little sexual dimorphism. Protibia of both sexes rather short and wide, basal third regularly expanded from base to apex, apical two-thirds subparallel, anteroexternal angle rather obliterated. Protarsus in male expanded, sub-oval, slightly more than ½ as long as protibia, slightly wider (
<figureCitation id="46C7D0FDFFEEFFFF550CCFED353EFD08" box="[517,611,677,704]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="6" pageNumber="7">Fig. 3B</figureCitation>
), inner margin straight, external margin convex, ventral side with large sucking disc on top of adhesive setae (
<figureCitation id="46C7D0FDFFEEFFFF55CBCF813443FD2B" box="[706,798,713,739]" captionStart="Fig" captionStartId="6.[189,232,1888,1914]" captionTargetBox="[208,1379,882,1850]" captionTargetId="figure-189@6.[208,1379,882,1853]" captionTargetPageId="6" captionText="Fig. 3. Patrus hainanensis sp. nov., ♂ (SYSU). A. Left elytron, paratype.B. Front leg., paratype C. Front tarsus, paratype. D. Genitalia, holotype. E. Surface sculpture on head, paratype. F. Surface sculpture on pronotum, paratype. G. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm." figureDoi="http://doi.org/10.5281/zenodo.5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="6" pageNumber="7">Fig. 3C</figureCitation>
), while not expanded and without adhesive setae in female.
</paragraph>
</subSubSection>
<caption id="8A839CF0FFEEFFFF57B4CA2836B8F82C" ID-DOI="http://doi.org/10.5281/zenodo.5715532" ID-Zenodo-Dep="5715532" httpUri="https://zenodo.org/record/5715532/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1888,1914]" targetBox="[208,1379,882,1850]" targetPageId="6">
<paragraph id="DE43CC78FFEEFFFF57B4CA2836B8F82C" blockId="6.[187,1399,1888,2020]" pageId="6" pageNumber="7">
<emphasis id="EC88106AFFEEFFFF57B4CA283654F8B2" bold="true" box="[189,265,1888,1914]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
<taxonomicName id="19FCB7FBFFEEFFFF5606CA2936AFF8B2" authority="Liang &amp; Angus &amp; Jia, 2021" authorityName="Liang &amp; Angus &amp; Jia" authorityYear="2021" box="[271,498,1888,1914]" class="Insecta" family="Gyrinidae" genus="Patrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="hainanensis" status="sp. nov.">
<emphasis id="EC88106AFFEEFFFF5606CA2936AFF8B2" box="[271,498,1888,1914]" italics="true" pageId="6" pageNumber="7">Patrus hainanensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F7BBAD11FFEEFFFF56FECA29350FF8B3" box="[503,594,1889,1915]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
, ♂ (SYSU).
<emphasis id="EC88106AFFEEFFFF55EACA2835A5F8B2" bold="true" box="[739,760,1888,1914]" pageId="6" pageNumber="7">A</emphasis>
. Left elytron, paratype.
<emphasis id="EC88106AFFEEFFFF5302CA283342F8B2" bold="true" box="[1035,1055,1888,1914]" pageId="6" pageNumber="7">B</emphasis>
. Front leg., paratype
<emphasis id="EC88106AFFEEFFFF521ACA283275F8B2" bold="true" box="[1299,1320,1888,1914]" pageId="6" pageNumber="7">C</emphasis>
. Front tarsus, paratype.
<emphasis id="EC88106AFFEEFFFF568BCACC36CAF856" bold="true" box="[386,407,1924,1950]" pageId="6" pageNumber="7">D</emphasis>
. Genitalia, holotype.
<emphasis id="EC88106AFFEEFFFF559DCACC35F5F856" bold="true" box="[660,680,1924,1950]" pageId="6" pageNumber="7">E</emphasis>
. Surface sculpture on head, paratype.
<emphasis id="EC88106AFFEEFFFF536FCACC3325F856" bold="true" box="[1126,1144,1924,1950]" pageId="6" pageNumber="7">F</emphasis>
. Surface sculpture on pronotum, paratype.
<emphasis id="EC88106AFFEEFFFF56BBCAEE3694F808" bold="true" box="[434,457,1958,1984]" pageId="6" pageNumber="7">G</emphasis>
. Surface sculpture on elytra, paratype. Scale bars: A = 1 mm; B, D = 0.5 mm; C = 200 μm; EG = 30 μm.
</paragraph>
</caption>
<subSubSection id="96E69FF3FFEFFFFE57B4CC403347FE99" box="[189,1050,264,338]" pageId="7" pageNumber="8" type="distribution">
<paragraph id="DE43CC78FFEFFFFE57B4CC403605FEEA" blockId="7.[189,344,264,290]" box="[189,344,264,290]" pageId="7" pageNumber="8">
<heading id="850B7B14FFEFFFFE57B4CC403605FEEA" bold="true" box="[189,344,264,290]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="EC88106AFFEFFFFE57B4CC403605FEEA" bold="true" box="[189,344,264,290]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="DE43CC78FFEFFFFE57B4CC7F3347FE99" blockId="7.[189,1050,311,338]" box="[189,1050,311,338]" pageId="7" pageNumber="8">
Only known from
<typeStatus id="014772DAFFEFFFFE569ECC703697FE9A" box="[407,458,312,338]" pageId="7" pageNumber="8">type</typeStatus>
locality Jianfengling Mt,
<collectingRegion id="1C38029AFFEFFFFE55F0CC7F349DFE99" box="[761,960,311,337]" country="China" name="Hainan" pageId="7" pageNumber="8">Hainan Province</collectingRegion>
,
<collectingCountry id="A6EB8CE8FFEFFFFE54C5CC7F334BFE99" box="[972,1046,311,337]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>