treatments-xml/data/56/11/A1/5611A119FFB6C27BFF2A6E1BFDBBC43B.xml

<document id="8C5709EE7F51A7ADEB16B91AFE00F979" ID-CLB-Dataset="29101" ID-DOI="10.11646/zootaxa.4455.2.7" ID-GBIF-Dataset="631d3896-6dff-4a16-8d5b-c96e11a50d9b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1457325" ID-ZooBank="D9D80F34-BF73-4C57-81A7-AF46A0FFACFA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1539262124874" checkinUser="plazi" docAuthor="Kundrаtа, Robin, Sormovа, Eliskа &amp; Prosvirov, Аlexаnder S." docDate="2018" docId="5611A119FFB6C27BFF2A6E1BFDBBC43B" docLanguage="en" docName="zootaxa.4455.2.7.pdf" docOrigin="Zootaxa 4455 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ludioctenus pakistanicus Schimmel &amp; Tarnawski 2012" docType="treatment" docVersion="7" lastPageNumber="386" masterDocId="AA28D961FFB2C272FFBD6851FF9CC606" masterDocTitle="Revisiting the diversity of Ludioctenus Fairmaire (Elateridae: Agrypninae), with description of a new species from Afghanistan, and discussion on the systematic position of Tetrigusina" masterLastPageNumber="388" masterPageNumber="377" pageNumber="381" updateTime="1698575614057" updateUser="ExternalLinkService">
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<mods:title id="8B6C88B0A141C7D16F21FDDBAA73641B">Revisiting the diversity of Ludioctenus Fairmaire (Elateridae: Agrypninae), with description of a new species from Afghanistan, and discussion on the systematic position of Tetrigusina</mods:title>
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<mods:namePart id="C4FF4B6A29E397288F2EA851B4F39FC2">Kundrаtа, Robin</mods:namePart>
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<mods:namePart id="AF0294A9F31B89893E9A3BFBCE2B6661">Sormovа, Eliskа</mods:namePart>
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<mods:namePart id="C096DFF6687E58B8CD895DC481F544F9">Prosvirov, Аlexаnder S.</mods:namePart>
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<heading id="854FA763FFB6C276FF2A6E1BFCDFC063" bold="true" box="[151,835,1610,1637]" fontSize="11" level="1" pageId="4" pageNumber="381" reason="1">
<taxonomicName id="19B86B8CFFB6C276FF2A6E1BFCDFC063" authority="Schimmel &amp; Tarnawski, 2012" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[151,835,1610,1637]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="pakistanicus">
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<emphasis id="ECCCCC1DFFB6C276FF2A6E1BFE57C062" bold="true" box="[151,459,1610,1636]" italics="true" pageId="4" pageNumber="381">Ludioctenus pakistanicus</emphasis>
<bibRefCitation id="BA296DFEFFB6C276FE6F6E1AFCDFC063" box="[466,835,1611,1637]" pageId="4" pageNumber="381" refId="ref6384" refString="Schimmel, R. &amp; Tarnawski, D. (2012) Monograph of the monophylum Tetrigusina (Insecta: Coleoptera, Elateridae, Agrypninae). Polish Entomological Monographs. Vol. 8. Polish Entomological Society, Poznan, 288 pp." type="book">Schimmel &amp; Tarnawski, 2012</bibRefCitation>
</emphasis>
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(
<figureCitation id="46830C8AFFB6C276FF226E3CFE82C080" box="[159,286,1645,1670]" captionStart-0="FIGURES 42–55" captionStart-1="FIGURES 56–61. 56" captionStartId-0="7.[151,264,1876,1899]" captionStartId-1="8.[151,264,1769,1792]" captionTargetBox-0="[230,1348,205,1846]" captionTargetBox-1="[161,1420,861,1748]" captionTargetId-0="figure@7.[229,1363,193,1861]" captionTargetId-1="figure@8.[151,1436,841,1748]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." captionText-1="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri-0="https://zenodo.org/record/1457333/files/figure.png" httpUri-1="https://zenodo.org/record/1457335/files/figure.png" pageId="4" pageNumber="381">Figs 42–61</figureCitation>
)
</paragraph>
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<taxonomicName id="19B86B8CFFB6C276FF2A6EE4FC9FC0CD" authority="Schimmel &amp; Tarnawski, 2012: 127" authorityName="Schimmel &amp; Tarnawski" authorityPageNumber="127" authorityYear="2012" box="[151,771,1717,1739]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="381" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFB6C276FF2A6EE4FE0BC0CD" box="[151,407,1717,1739]" italics="true" pageId="4" pageNumber="381">Ludioctenus pakistanicus</emphasis>
<treatmentCitation id="5F19361EFFB6C276FE236EE4FC9FC0CD" author="" box="[414,771,1717,1739]" page="127" pageId="4" pageNumber="381">
<bibRefCitation id="BA296DFEFFB6C276FE236EE4FD4DC0CD" box="[414,721,1717,1739]" pageId="4" pageNumber="381" refId="ref6384" refString="Schimmel, R. &amp; Tarnawski, D. (2012) Monograph of the monophylum Tetrigusina (Insecta: Coleoptera, Elateridae, Agrypninae). Polish Entomological Monographs. Vol. 8. Polish Entomological Society, Poznan, 288 pp." type="book">Schimmel &amp; Tarnawski, 2012</bibRefCitation>
: 127
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.
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</paragraph>
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<paragraph id="DE07100FFFB6C276FF2A6EA6FC17C1A6" blockId="4.[151,1436,1783,2025]" pageId="4" pageNumber="381">
<materialsCitation id="6ED01A52FFB6C276FF2A6EA6FD63C132" ID-GBIF-Occurrence="1917075844" collectingDate="1971-07-04" collectionCode="NKME" collectorName="Material &amp; Prov. Swat &amp; Madyan &amp; Licht &amp; C. Holzschuh" country="Pakistan" elevation="1400" location="PAKISTAN" pageId="4" pageNumber="381" specimenCount="1" typeStatus="holotype">
<emphasis id="ECCCCC1DFFB6C276FF2A6EA6FEE4C116" bold="true" box="[151,376,1783,1808]" pageId="4" pageNumber="381">
<collectorName id="734D75D9FFB6C276FF2A6EA6FF63C116" box="[151,255,1783,1808]" pageId="4" pageNumber="381">Material</collectorName>
examined
</emphasis>
. 
<emphasis id="ECCCCC1DFFB6C276FE3B6EA6FD86C116" bold="true" box="[390,538,1783,1808]" pageId="4" pageNumber="381">
<collectingCountry id="A6AF509FFFB6C276FE3B6EA6FD89C116" box="[390,533,1783,1808]" name="Pakistan" pageId="4" pageNumber="381">PAKISTAN</collectingCountry>
.
</emphasis>
<typeStatus id="0103AEADFFB6C276FD9D6EA6FD17C116" box="[544,651,1783,1808]" pageId="4" pageNumber="381" type="holotype">Holotype</typeStatus>
, male, &quot;NW-Pakistan, 
<collectorName id="734D75D9FFB6C276FC356EA9FB9FC116" box="[904,1027,1784,1808]" pageId="4" pageNumber="381">Prov. Swat</collectorName>
, 71°90'L/35°70'B, 
<collectorName id="734D75D9FFB6C276FB656EA6FAABC116" box="[1240,1335,1783,1808]" pageId="4" pageNumber="381">Madyan</collectorName>
, 
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<quantity id="1940BDEAFFB6C276FAFC6EA9FA0BC109" box="[1345,1431,1784,1808]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="381" unit="m" value="1400.0">1400 m</quantity>
</elevation>
, am 
<collectorName id="734D75D9FFB6C276FF026F4DFF61C133" box="[191,253,1820,1845]" pageId="4" pageNumber="381">Licht</collectorName>
, 19.6.– 
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<date id="AA0636CFFFB6C276FEF66F4DFE2EC133" box="[331,434,1820,1845]" pageId="4" pageNumber="381" value="1971-07-04">4.7.1971</date>
</collectingDate>
, leg. 
<collectorName id="734D75D9FFB6C276FE506F4DFD14C133" box="[493,648,1820,1845]" pageId="4" pageNumber="381">C. Holzschuh</collectorName>
&quot; (
<collectionCode id="B8A988CAFFB6C276FD236F4DFD6BC132" box="[670,759,1820,1844]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13752" name="Naturkundemuseum Erfurt" pageId="4" pageNumber="381">NKME</collectionCode>
)
</materialsCitation>
. 
<materialsCitation id="6ED01A52FFB6C276FCB46F4DFA04C132" ID-GBIF-Occurrence="1917075843" box="[777,1432,1820,1845]" collectingDate="1971-07-04" collectionCode="NKME" collectorName="Material &amp; Prov. Swat &amp; Madyan &amp; Licht &amp; C. Holzschuh" country="Pakistan" elevation="1400" location="PAKISTAN" pageId="4" pageNumber="381" specimenCount="2" typeStatus="holotype">
<specimenCount id="C8BEDB86FFB6C276FCB46F4DFC17C133" box="[777,907,1820,1845]" pageId="4" pageNumber="381" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
, males, same data as for the 
<typeStatus id="0103AEADFFB6C276FB7A6F4DFAB5C133" box="[1223,1321,1820,1845]" pageId="4" pageNumber="381" type="holotype">holotype</typeStatus>
(
<collectionCode id="B8A988CAFFB6C276FA8A6F4DFA0CC132" box="[1335,1424,1820,1844]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13752" name="Naturkundemuseum Erfurt" pageId="4" pageNumber="381">NKME</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6ED01A52FFB6C276FF2A6F11FF61C17A" ID-GBIF-Occurrence="1917075864" collectingDate="1998-07-27" collectingDateMax="1998-07-29" collectingDateMin="1998-07-27" collectionCode="NMPC" collectorName="L. Cizek &amp; L. Cerny" country="Pakistan" location="Chitral" pageId="4" pageNumber="381" specimenCount="2" typeStatus="paratype">
<specimenCount id="C8BEDB86FFB6C276FF2A6F11FE87C15E" box="[151,283,1856,1880]" pageId="4" pageNumber="381" type="generic" typeStatus="paratypes">2 paratypes</specimenCount>
, male and female, &quot;Pakistan-N, NWF prov., 
<location id="DB6746D4FFB6C276FCA16F6EFCF6C15E" LSID="urn:lsid:plazi:treatment:5611A119FFB6C27BFF2A6E1BFDBBC43B:DB6746D4FFB6C276FCA16F6EFCF6C15E" box="[796,874,1855,1880]" country="Pakistan" name="Chitral" pageId="4" pageNumber="381">Chitral</location>
env., 
<collectingDate id="BA42CF27FFB6C276FC0D6F6EFBFFC15E" box="[944,1123,1855,1880]" pageId="4" pageNumber="381" value="1998-07-27" valueMax="1998-07-29" valueMin="1998-07-27">
<date id="AA0636CFFFB6C276FC0D6F6EFBFFC15E" box="[944,1123,1855,1880]" pageId="4" pageNumber="381" value="1998-07-27" valueMax="1998-07-29" valueMin="1998-07-27">27.–29.vii.1998</date>
</collectingDate>
, 
<collectorName id="734D75D9FFB6C276FBCD6F11FB4EC15E" box="[1136,1234,1855,1880]" pageId="4" pageNumber="381">L. Čížek</collectorName>
&amp; 
<collectorName id="734D75D9FFB6C276FB476F11FAC2C15E" box="[1274,1374,1855,1880]" pageId="4" pageNumber="381">L. Černý</collectorName>
leg.&quot; (
<collectionCode id="B8A988CAFFB6C276FF226F35FF69C17A" box="[159,245,1892,1916]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="4" pageNumber="381">NMPC</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="6ED01A52FFB6C276FEB16F35FC17C1A6" ID-GBIF-Occurrence="1917075863" collectionCode="MNHN" collectorName="Coll. E. &amp; Fleutiaux" country="India" county="southern India" location="Shembaganur" municipality="Sd. Ind." pageId="4" pageNumber="381" specimenCount="2" typeStatus="paratype">
<specimenCount id="C8BEDB86FFB6C276FEB16F35FE15C17B" box="[268,393,1892,1917]" pageId="4" pageNumber="381" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, male, &quot;T. R. Pell, Karachi, 
<collectorName id="734D75D9FFB6C276FD626F35FC2FC17B" box="[735,947,1892,1917]" pageId="4" pageNumber="381">Coll. E. Fleutiaux</collectorName>
&quot; (
<collectionCode id="B8A988CAFFB6C276FC726F35FBB7C17A" box="[975,1067,1892,1916]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="381">MNHN</collectionCode>
). 
<emphasis id="ECCCCC1DFFB6C276FBFF6F35FB07C17B" bold="true" box="[1090,1179,1892,1917]" pageId="4" pageNumber="381">
<collectingCountry id="A6AF509FFFB6C276FBFF6F35FB0AC17B" box="[1090,1174,1892,1917]" name="India" pageId="4" pageNumber="381">INDIA</collectingCountry>
.
</emphasis>
<specimenCount id="C8BEDB86FFB6C276FB1A6F35FAB8C17B" box="[1191,1316,1892,1917]" pageId="4" pageNumber="381" type="generic" typeStatus="paratype">1 paratype</specimenCount>
, male (?), &quot; 
<location id="DB6746D4FFB6C276FF226FD6FEA0C1A6" LSID="urn:lsid:plazi:treatment:5611A119FFB6C27BFF2A6E1BFDBBC43B:DB6746D4FFB6C276FF226FD6FEA0C1A6" box="[159,316,1927,1952]" country="India" county="southern India" municipality="Sd. Ind." name="Shembaganur" pageId="4" pageNumber="381">Shembaganur</location>
[
<location id="DB6746D4FFB6C276FEF76FD6FE79C1A6" LSID="urn:lsid:plazi:treatment:5611A119FFB6C27BFF2A6E1BFDBBC43B:DB6746D4FFB6C276FEF76FD6FE79C1A6" box="[330,485,1927,1952]" country="India" county="southern India" municipality="Sd. Ind." name="Shenbaganur" pageId="4" pageNumber="381">Shenbaganur</location>
], 
<collectingMunicipality id="3E638A75FFB6C276FE4B6FD6FDD2C1A6" box="[502,590,1927,1952]" pageId="4" pageNumber="381">Sd. Ind.</collectingMunicipality>
[
<collectingCounty id="37666883FFB6C276FDE66FD6FC98C1A6" box="[603,772,1927,1952]" pageId="4" pageNumber="381">southern India</collectingCounty>
]&quot; (
<collectionCode id="B8A988CAFFB6C276FC9E6FD9FCE3C1A6" box="[803,895,1928,1952]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="381">MNHN</collectionCode>
).
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</paragraph>
</subSubSection>
<subSubSection id="96A24384FFB6C27AFF7A6FFDFE8CC513" lastPageId="8" lastPageNumber="385" pageId="4" pageNumber="381" type="description">
<paragraph id="DE07100FFFB6C276FF7A6FFDFA00C1EE" blockId="4.[151,1436,1783,2025]" pageId="4" pageNumber="381">
<emphasis id="ECCCCC1DFFB6C276FF7A6FFDFC54C1C3" bold="true" box="[199,968,1964,1989]" pageId="4" pageNumber="381">Description of terminal abdominal segments and genitalia. Male</emphasis>
. Tergite VIII (
<figureCitation id="46830C8AFFB6C276FBD36FFDFB5CC1C3" box="[1134,1216,1964,1989]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="4" pageNumber="381">Fig. 50</figureCitation>
) about 1.3 times as wide as long, apically widely rounded, finely punctate, covered with dense pubescence. Sternite VIII (
<figureCitation id="46830C8AFFB6C276FA816F81FA08C1EE" box="[1340,1428,2000,2025]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="4" pageNumber="381">Fig. 49</figureCitation>
)
</paragraph>
<caption id="8AC74087FFB7C277FF2A6F02FEBCC1E2" httpUri="https://zenodo.org/record/1457329/files/figure.png" pageId="5" pageNumber="382" startId="5.[151,264,1875,1898]" targetBox="[229,1346,203,1849]" targetPageId="5">
<paragraph id="DE07100FFFB7C277FF2A6F02FEBCC1E2" blockId="5.[151,1436,1875,2020]" pageId="5" pageNumber="382">
<emphasis id="ECCCCC1DFFB7C277FF2A6F02FECAC16F" bold="true" box="[151,342,1875,1898]" pageId="5" pageNumber="382">FIGURES 15–28.</emphasis>
<taxonomicName id="19B86B8CFFB7C277FEE06F05FCBEC16C" authority="(Baudi di Selve, 1871)" authorityName="Baudi di Selve" authorityYear="1871" baseAuthorityName="Baudi di Selve" baseAuthorityYear="1871" box="[349,802,1876,1898]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="382" phylum="Arthropoda" rank="species" species="cyprius">
<emphasis id="ECCCCC1DFFB7C277FEE06F05FDB1C16C" box="[349,557,1876,1898]" italics="true" pageId="5" pageNumber="382">Ludioctenus cyprius</emphasis>
(Baudi di Selve, 1871)
</taxonomicName>
, male (Greece). 15, habitus, dorsal view; 16, habitus, ventral view; 17, habitus, lateral view; 18, antenna; 19, pronotum; 20, scutellar shield; 21, interior part of metacoxal plate, 22, abdominal sternite VIII; 23, abdominal tergite VIII; 24, abdominal sternite IX; 25, abdominal sternite X; 26, abdominal tergites IX–X; 27, aedeagus, ventral view; 28, aedeagus, dorsal view. Scale bars = 5.0 mm (Figs 15–17, 19), 2.0 mm (Fig. 18), 1.0 mm (Figs 20–28).
</paragraph>
</caption>
<caption id="8AC74087FFB4C274FF2A6F03FD3AC1E5" httpUri="https://zenodo.org/record/1457331/files/figure.png" pageId="6" pageNumber="383" startId="6.[151,264,1874,1897]" targetBox="[235,1337,201,1854]" targetPageId="6">
<paragraph id="DE07100FFFB4C274FF2A6F03FD3AC1E5" blockId="6.[151,1435,1874,2019]" pageId="6" pageNumber="383">
<emphasis id="ECCCCC1DFFB4C274FF2A6F03FEC9C16E" bold="true" box="[151,341,1874,1897]" pageId="6" pageNumber="383">FIGURES 29–41.</emphasis>
<taxonomicName id="19B86B8CFFB4C274FEE66F02FC86C16F" authority="(Baudi di Selve, 1871)" authorityName="Baudi di Selve" authorityYear="1871" baseAuthorityName="Baudi di Selve" baseAuthorityYear="1871" box="[347,794,1875,1897]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="383" phylum="Arthropoda" rank="species" species="cyprius">
<emphasis id="ECCCCC1DFFB4C274FEE66F02FDB5C16F" box="[347,553,1875,1897]" italics="true" pageId="6" pageNumber="383">Ludioctenus cyprius</emphasis>
(Baudi di Selve, 1871)
</taxonomicName>
, female (Greece). 29, habitus, dorsal view; 30, habitus, ventral view; 31, habitus, lateral view; 32, antenna; 33, pronotum; 34, scutellar shield; 35, interior part of metacoxal plate, 36, abdominal tergite VII; 37, abdominal tergite VIII and sternite VIII, ventral view; 38, abdominal tergite VIII and sternite VIII, dorsal view; 39, ovipositor, ventral view; 40, ovipositor, dorsal view; 41, female genital tract. Scale bars = 5.0 mm (Figs 29–31, 33), 2.0 mm (Fig. 32–40), 1.0 mm (Figs 34–35, 41).
</paragraph>
</caption>
<caption id="8AC74087FFB5C275FF2A6F05FE63C1E3" httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="7" pageNumber="384" startId="7.[151,264,1876,1899]" targetBox="[230,1348,205,1846]" targetPageId="7">
<paragraph id="DE07100FFFB5C275FF2A6F05FE63C1E3" blockId="7.[151,1436,1876,2021]" pageId="7" pageNumber="384">
<emphasis id="ECCCCC1DFFB5C275FF2A6F05FEC4C16C" bold="true" box="[151,344,1876,1899]" pageId="7" pageNumber="384">FIGURES 42–55.</emphasis>
<taxonomicName id="19B86B8CFFB5C275FEDF6F04FC2DC16D" authority="Schimmel &amp; Tarnawski, 2012" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[354,945,1877,1899]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="384" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFB5C275FEDF6F04FDFBC16D" box="[354,615,1877,1899]" italics="true" pageId="7" pageNumber="384">Ludioctenus pakistanicus</emphasis>
Schimmel &amp; Tarnawski, 2012
</taxonomicName>
, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55).
</paragraph>
</caption>
<paragraph id="DE07100FFFBAC27AFF2A68C6FE8CC513" blockId="8.[151,1437,151,789]" pageId="8" pageNumber="385">
subtriangular, partly membranous, about 1.3 times as wide as long, apically slightly emarginate, medially and apicolaterally finely punctate, covered with short dense pubescence. Tergites IX and X (
<figureCitation id="46830C8AFFBAC27AFB2168EDFB6DC6D3" box="[1180,1265,188,213]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="8" pageNumber="385">Fig. 53</figureCitation>
) connected by membrane; tergite IX about 1.8 times as wide as long, emarginate apically; tergite X small, widely rounded apically. Sternite IX (
<figureCitation id="46830C8AFFBAC27AFE326955FE78C71B" box="[399,484,260,285]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="8" pageNumber="385">Fig. 51</figureCitation>
) elongate, about 2.1 times as long as wide, apically rounded, apicolaterally finely punctate, with short dense pubescence. Sternite X (
<figureCitation id="46830C8AFFBAC27AFCB86976FCC2C746" box="[773,862,295,320]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="8" pageNumber="385">Fig. 52</figureCitation>
) connected by membrane to sternite IX, partly membranous, with two elongate, subparallel-sided sclerotized plates. Аedeagus (
<figureCitation id="46830C8AFFBAC27AFB8D691DFB2EC763" box="[1072,1202,332,357]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="8" pageNumber="385">Figs 54–55</figureCitation>
) elongate, about 3.3 times as long as wide. Median lobe elongate, robust, slightly surpassing parameres, gradually narrowed towards apex, apically abruptly narrowed, apex narrowly rounded, with moderately long basal struts. Paramere elongate, gradually narrowed from base toward apex, gradually constricted subapically, with subapical hook, apical portion covered with short setae, apex narrowly rounded; subapical hook short, thin, its outer portion not or only shallowly emarginate. Phallobase robust, transverse, about 1.2 times as wide as long. 
<emphasis id="ECCCCC1DFFBAC27AFB9069AEFB18C41E" bold="true" box="[1069,1156,511,536]" pageId="8" pageNumber="385">Female</emphasis>
. Tergite VIII (
<figureCitation id="46830C8AFFBAC27AFA8669AEFA0FC41E" box="[1339,1427,511,536]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="8" pageNumber="385">Fig. 61</figureCitation>
) subtriangular, slightly longer than wide, narrowed toward apex, subacute apically, finely punctate, covered with moderately dense pubescence, mainly at margins. Sternite VIII (
<figureCitation id="46830C8AFFBAC27AFCD36A16FC5DC466" box="[878,961,583,608]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="8" pageNumber="385">Fig. 59</figureCitation>
) slightly longer than wide, with sides and apex rounded, finely punctate, covered with dense pubescence, spiculum ventrale slender, long, about 2.6 times sternite length. Ovipositor (
<figureCitation id="46830C8AFFBAC27AFE646ADEFDB3C4AE" box="[473,559,655,680]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="8" pageNumber="385">Fig. 58</figureCitation>
) elongate; paraprocts about 7.5 times as long as gonocoxites, styli absent. Vagina with plate-like sclerotized structures. 
<collectingRegion id="1C7CDEEDFFBAC27AFD266AE4FD40C4CA" box="[667,732,693,716]" country="Turkey" name="Bursa" pageId="8" pageNumber="385">Bursa</collectingRegion>
copulatrix (
<figureCitation id="46830C8AFFBAC27AFCD46AE5FC20C4CB" box="[873,956,692,717]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="8" pageNumber="385">Fig. 60</figureCitation>
) mebranous, sac-like, anteriorly elongate, with long rows of jointly connected spines. Two elongate spermathecae attached to anterior part of bursa copulatrix.
</paragraph>
</subSubSection>
<caption id="8AC74087FFBAC27AFF2A6EB8FC72C13B" httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="8" pageNumber="385" startId="8.[151,264,1769,1792]" targetBox="[161,1420,861,1748]" targetPageId="8">
<paragraph id="DE07100FFFBAC27AFF2A6EB8FC72C13B" blockId="8.[151,1437,1769,1853]" pageId="8" pageNumber="385">
<emphasis id="ECCCCC1DFFBAC27AFF2A6EB8FEC9C0F9" bold="true" box="[151,341,1769,1792]" pageId="8" pageNumber="385">FIGURES 56–61.</emphasis>
56, 
<taxonomicName id="19B86B8CFFBAC27AFE396EBBFC5AC106" authority="Schimmel &amp; Tarnawski, 2012" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[388,966,1770,1792]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="385" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFBAC27AFE396EBBFD1BC106" box="[388,647,1770,1792]" italics="true" pageId="8" pageNumber="385">Ludioctenus pakistanicus</emphasis>
Schimmel &amp; Tarnawski, 2012
</taxonomicName>
, paratype, male? (India), pronotum; 57–61 
<taxonomicName id="19B86B8CFFBAC27AFA3A6EBAFE8AC118" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="385" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFBAC27AFA3A6EBAFE8AC118" italics="true" pageId="8" pageNumber="385">L. pakistanicus</emphasis>
</taxonomicName>
, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61).
</paragraph>
</caption>
<subSubSection id="96A24384FFBAC27AFF7A6F38FD89C184" box="[199,533,1897,1922]" pageId="8" pageNumber="385" type="distribution">
<paragraph id="DE07100FFFBAC27AFF7A6F38FD89C184" blockId="8.[151,1437,1897,2030]" box="[199,533,1897,1922]" pageId="8" pageNumber="385">
<emphasis id="ECCCCC1DFFBAC27AFF7A6F38FEC5C184" bold="true" box="[199,345,1897,1922]" pageId="8" pageNumber="385">Distribution</emphasis>
. 
<collectingCountry id="A6AF509FFFBAC27AFEDA6F38FE3FC184" box="[359,419,1897,1922]" name="India" pageId="8" pageNumber="385">India</collectingCountry>
, 
<collectingCountry id="A6AF509FFFBAC27AFE136F38FD8DC184" box="[430,529,1897,1922]" name="Pakistan" pageId="8" pageNumber="385">Pakistan</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="96A24384FFBAC27BFF7A6FDCFDBBC43B" lastPageId="9" lastPageNumber="386" pageId="8" pageNumber="385" type="discussion">
<paragraph id="DE07100FFFBAC27BFF7A6FDCFCB1C7D6" blockId="8.[151,1437,1897,2030]" lastBlockId="9.[151,1437,151,573]" lastPageId="9" lastPageNumber="386" pageId="8" pageNumber="385">
<emphasis id="ECCCCC1DFFBAC27AFF7A6FDCFEA8C1A0" bold="true" box="[199,308,1933,1958]" pageId="8" pageNumber="385">Remarks</emphasis>
. 
<bibRefCitation id="BA296DFEFFBAC27AFEFC6FDCFD07C1A0" box="[321,667,1933,1958]" pageId="8" pageNumber="385" refId="ref6384" refString="Schimmel, R. &amp; Tarnawski, D. (2012) Monograph of the monophylum Tetrigusina (Insecta: Coleoptera, Elateridae, Agrypninae). Polish Entomological Monographs. Vol. 8. Polish Entomological Society, Poznan, 288 pp." type="book">Schimmel &amp; Tarnawski (2012)</bibRefCitation>
listed only 
<collectingCountry id="A6AF509FFFBAC27AFCA06FDCFCE0C1A0" box="[797,892,1933,1958]" name="Pakistan" pageId="8" pageNumber="385">Pakistan</collectingCountry>
for the distribution of 
<taxonomicName id="19B86B8CFFBAC27AFBC46FDEFABEC1A0" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[1145,1314,1933,1958]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="385" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFBAC27AFBC46FDEFABEC1A0" box="[1145,1314,1933,1958]" italics="true" pageId="8" pageNumber="385">L. pakistanicus</emphasis>
</taxonomicName>
. However, they included in the 
<typeStatus id="0103AEADFFBAC27AFE3E6FE3FE2FC1CC" box="[387,435,1970,1994]" pageId="8" pageNumber="385">type</typeStatus>
series a specimen from MNHN with the data &quot;Shembaganur, without further data&quot;. We examined this specimen and in fact it bears the label &quot;Shembaganur, Sd. Ind.&quot;. Because Shembaganur (or Shenbaganur) lies in the state of 
<collectingRegion id="1C7CDEEDFFBBC27BFDA968C6FD07C6B6" box="[532,667,151,176]" country="India" name="Tamil Nadu" pageId="9" pageNumber="386">Tamil Nadu</collectingRegion>
in southern 
<collectingCountry id="A6AF509FFFBBC27BFC9068C6FCF5C6B6" box="[813,873,151,176]" name="India" pageId="9" pageNumber="386">India</collectingCountry>
, we include here also 
<collectingCountry id="A6AF509FFFBBC27BFBD068C6FB3BC6B6" box="[1133,1191,151,176]" name="India" pageId="9" pageNumber="386">India</collectingCountry>
in the distribution of 
<taxonomicName id="19B86B8CFFBBC27BFF2A68EDFEBCC6D3" authorityName="Fairmaire" authorityYear="1893" box="[151,288,188,213]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="ECCCCC1DFFBBC27BFF2A68EDFEBCC6D3" box="[151,288,188,213]" italics="true" pageId="9" pageNumber="386">Ludioctenus</emphasis>
</taxonomicName>
. 
<bibRefCitation id="BA296DFEFFBBC27BFE8B68EDFD36C6D2" box="[310,682,188,213]" pageId="9" pageNumber="386" refId="ref6384" refString="Schimmel, R. &amp; Tarnawski, D. (2012) Monograph of the monophylum Tetrigusina (Insecta: Coleoptera, Elateridae, Agrypninae). Polish Entomological Monographs. Vol. 8. Polish Entomological Society, Poznan, 288 pp." type="book">Schimmel &amp; Tarnawski (2012)</bibRefCitation>
suggested that the specimen from Shembaganur is a female. Unfortunately, it is in a bad condition and most of abdomen including the soft tissues and genitalia is missing. However, based on the shape of pronotum (
<figureCitation id="46830C8AFFBBC27BFD2D6955FD78C71B" box="[656,740,260,285]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="9" pageNumber="386">Fig. 56</figureCitation>
) and antenna it is most probably a male. What is more, this specimen has much sparser pronotal punctation than the specimens from 
<collectingCountry id="A6AF509FFFBBC27BFC796976FBBFC746" box="[964,1059,295,320]" name="Pakistan" pageId="9" pageNumber="386">Pakistan</collectingCountry>
(
<figureCitation id="46830C8AFFBBC27BFB8C6976FB2DC746" box="[1073,1201,295,320]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="9" pageNumber="386">Figs 56–57</figureCitation>
) and might represent an undescribed species. Since several morphological characters such as the shape of the scutellar shield are variable in 
<taxonomicName id="19B86B8CFFBBC27BFF0E6920FEC0C78E" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[179,348,367,392]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFBBC27BFF0E6920FEC0C78E" box="[179,348,367,392]" italics="true" pageId="9" pageNumber="386">L. pakistanicus</emphasis>
</taxonomicName>
(scutellar shield about 2.1 times as long as wide in the 
<typeStatus id="0103AEADFFBBC27BFC79693EFBBAC78E" box="[964,1062,367,392]" pageId="9" pageNumber="386" type="holotype">holotype</typeStatus>
(
<figureCitation id="46830C8AFFBBC27BFB88693EFB1AC78E" box="[1077,1158,367,392]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="9" pageNumber="386">Fig. 47</figureCitation>
) but 1.8–2.0 times in the examined male 
<typeStatus id="0103AEADFFBBC27BFEF469C5FE25C7AA" box="[329,441,404,428]" pageId="9" pageNumber="386" type="paratype">paratypes</typeStatus>
), we prefer to include the specimen from Shembaganur tentatively under 
<taxonomicName id="19B86B8CFFBBC27BFB4F69C4FA00C7AB" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[1266,1436,404,429]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFBBC27BFB4F69C4FA00C7AB" box="[1266,1436,404,429]" italics="true" pageId="9" pageNumber="386">L. pakistanicus</emphasis>
</taxonomicName>
until more specimens are available for more detailed study.
</paragraph>
<paragraph id="DE07100FFFBBC27BFF7A698DFDBBC43B" blockId="9.[151,1437,151,573]" pageId="9" pageNumber="386">
The male 
<typeStatus id="0103AEADFFBBC27BFE84698DFE07C7F3" box="[313,411,476,501]" pageId="9" pageNumber="386" type="holotype">holotype</typeStatus>
of 
<taxonomicName id="19B86B8CFFBBC27BFE7F698CFDF1C7F3" authorityName="Schimmel &amp; Tarnawski" authorityYear="2012" box="[450,621,476,501]" class="Insecta" family="Elateridae" genus="Ludioctenus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="386" phylum="Arthropoda" rank="species" species="pakistanicus">
<emphasis id="ECCCCC1DFFBBC27BFE7F698CFDF1C7F3" box="[450,621,476,501]" italics="true" pageId="9" pageNumber="386">L. pakistanicus</emphasis>
</taxonomicName>
was not originally dissected and therefore we figure here the pregenital sclerites and aedeagus (
<figureCitation id="46830C8AFFBBC27BFE1E69AEFDB4C41E" box="[419,552,511,536]" captionStart="FIGURES 42–55" captionStartId="7.[151,264,1876,1899]" captionTargetBox="[230,1348,205,1846]" captionTargetId="figure@7.[229,1363,193,1861]" captionTargetPageId="7" captionText="FIGURES 42–55. Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, holotype, male. 42, habitus, dorsal view; 43, habitus, ventral view; 44, habitus, lateral view; 45, antenna; 46, pronotum; 47, scutellar shield; 48, interior part of metacoxal plate, 49, abdominal sternite VIII; 50, abdominal tergite VIII; 51, abdominal sternite IX; 52, abdominal sternite X; 53, abdominal tergites IX–X; 54, aedeagus, dorsal view; 55, aedeagus, ventral view. Scale bars = 5.0 mm (Figs 42–44), 2.0 mm (Fig. 45–46), 1.0 mm (Figs 47–55)." httpUri="https://zenodo.org/record/1457333/files/figure.png" pageId="9" pageNumber="386">Figs 49–55</figureCitation>
). For the first time, we figure here also pregenital segments and genitalia of a female of this species (
<figureCitation id="46830C8AFFBBC27BFE256A75FD86C43B" box="[408,538,548,573]" captionStart="FIGURES 56–61. 56" captionStartId="8.[151,264,1769,1792]" captionTargetBox="[161,1420,861,1748]" captionTargetId="figure@8.[151,1436,841,1748]" captionTargetPageId="8" captionText="FIGURES 56–61. 56, Ludioctenus pakistanicus Schimmel &amp; Tarnawski, 2012, paratype, male? (India), pronotum; 57–61 L. pakistanicus, paratype, female (Pakistan). 57, pronotum; 58, ovipositor; 59, abdominal sternite VIII; 60, female genital tract; 61, abdominal tergite VIII. Scale bars = 5.0 mm (Figs 56–57), 2.0 mm (Figs 58–61)." httpUri="https://zenodo.org/record/1457335/files/figure.png" pageId="9" pageNumber="386">Figs 58–61</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>