150 lines
16 KiB
XML
150 lines
16 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.66.684" ID-GBIF-Dataset="35e99d78-af9c-400e-b111-6abc6e1d9b1d" ID-PMC="PMC3088417" ID-Pensoft-Pub="1313-2970-66-1" ID-Pensoft-UUID="AD1FFFAF9122FFEAFF8EEF3BE966F439" ID-PubMed="21594029" ID-Zenodo-Dep="576888" ID-ZooBank="176306EB63424E75AD76C4A82040A002" ModsDocAuthor="" ModsDocDate="2010" ModsDocID="1313-2970-66-1" ModsDocOrigin="ZooKeys 66" ModsDocTitle="Agra, arboreal beetles of Neotropical forests:pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)" checkinTime="1451250911506" checkinUser="pensoft" docAuthor="Erwin, Terry L." docDate="2010" docId="55FB64AAF91FBFF0EF1510282D17CF98" docLanguage="en" docName="ZooKeys 66: 1-28" docOrigin="ZooKeys 66" docSource="http://dx.doi.org/10.3897/zookeys.66.684" docTitle="Agra cruciaria Erwin 2010, sp. n." docType="treatment" docUuid="7906629D-6846-4348-A3CD-B7A3031C7AE9" docUuidSource="ZooBank" docVersion="7" id="AD1FFFAF9122FFEAFF8EEF3BE966F439" lastPageNumber="6" masterDocId="AD1FFFAF9122FFEAFF8EEF3BE966F439" masterDocTitle="Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)" masterLastPageNumber="28" masterPageNumber="1" pageNumber="5" updateTime="1732481568501" updateUser="ExternalLinkService">
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<mods:mods id="FAD8C3C07C23D82EFB935148C5E3DFA8" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BA3BD9EFE14A62A26CE3DA999CE07DE4">
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<mods:title id="6C3FBF9BA24A557BB29895532A8CF7AC">Agra, arboreal beetles of Neotropical forests: pusilla group and piranha group systematics and notes on their ways of life (Coleoptera, Carabidae, Lebiini, Agrina)</mods:title>
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<mods:name id="F969B4C292DE58655BDBD58BAE4AE87E" type="personal">
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<mods:roleTerm id="EA0B1FD94DD785AB6CAF771388481CF7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0465900AD2547924A82280A5B6C6C007">Erwin, Terry L.</mods:namePart>
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<mods:affiliation id="1B47E9B9176C222D9431DC01907C0E2F">Washington DC</mods:affiliation>
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<mods:nameIdentifier id="63C6BFC36286143A0B9565FF49043281" type="email">erwint@si.edu</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource id="3AF7B77F8F0E8E3639EA75C9D25FA141">text</mods:typeOfResource>
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<mods:title id="34F4C13A45C78D87942DE9AFC237B475">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="B7EC23B99CF591A773D091B31B684AA3">
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<mods:date id="A684C887A9005C6CF6B47BA83CECF282">2010</mods:date>
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<mods:detail id="F69AD64DFB6BB82C67F0C5FC93401859" type="volume">
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<mods:number id="C83FF2223F845F277F16CF927C71CDFF">66</mods:number>
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</mods:detail>
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<mods:extent id="6C9979A6ED51CD6463A57B9C98E7EF43" unit="page">
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<mods:start id="87A6EFA5120E2F3D0F1FB4C8D304A50B">1</mods:start>
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<mods:end id="919494AF9C60DCFD8DCD15CB0BD80553">28</mods:end>
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<mods:location id="04CA28517C93A969EA4C949F2177774F">
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<mods:url id="5604FAD0D38B860F2469ACED51A15461">http://dx.doi.org/10.3897/zookeys.66.684</mods:url>
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</mods:location>
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<mods:classification id="6BF6CA9D5D51A5B06FF2CD7BD1D6F7A9">journal article</mods:classification>
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<mods:identifier id="AC6363DA3E5352231C6CBB9FD3FF0F59" type="DOI">http://dx.doi.org/10.3897/zookeys.66.684</mods:identifier>
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<mods:identifier id="DCA99BB35CBE208BF2EF370E153D8C1D" type="Pensoft-Pub">1313-2970-66-1</mods:identifier>
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<mods:identifier id="041D0C5C79930BBF8B88677782A1FC37" type="ZooBank">176306EB63424E75AD76C4A82040A002</mods:identifier>
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<mods:identifier id="36C9E9EC4F5B422717DC84D975C67922" type="Pensoft-UUID">AD1FFFAF9122FFEAFF8EEF3BE966F439</mods:identifier>
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<mods:identifier id="AF52637C169B21946503BA7889918FE1" type="Zenodo-Dep">576888</mods:identifier>
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</mods:mods>
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<treatment id="55FB64AAF91FBFF0EF1510282D17CF98" ID-GBIF-Taxon="159360645" LSID="urn:lsid:zoobank.org:act:7906629D-6846-4348-A3CD-B7A3031C7AE9" httpUri="http://treatment.plazi.org/id/55FB64AAF91FBFF0EF1510282D17CF98" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" scope_family="Carabidae" scope_order="Coleoptera" scope_subTribe="Agrina" scope_tribe="Lebiini">
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<subSubSection id="9BDF9E4DE73665998031A3DFBC106575" pageId="4" pageNumber="5" type="nomenclature">
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<paragraph id="CB382579D34AF84530187699A060F55E" pageId="4" pageNumber="5">
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<taxonomicName id="FDFA05B23B4CDE05A8940D2DA9434026" ID-CoL="B2KQ" LSID="55FB64AA-F91F-BFF0-EF15-10282D17CF98" authority="Erwin" authorityName="Erwin" authorityYear="2010" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Agra cruciaria" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="cruciaria" status="sp. n.">
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<pageBreakToken id="8E95EB78365F1E4FB462683A0EC40CFE" pageId="4" pageNumber="5" start="start">Agra</pageBreakToken>
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cruciaria Erwin
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</taxonomicName>
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<taxonomicNameLabel id="2B156AEB1CB7AE323F2F8BDC3B031B9B" pageId="4" pageNumber="5">sp. n.</taxonomicNameLabel>
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<figureCitation id="EFE3EE7B481211B4988D645771B56DE7" captionStart="Figures 3–6" captionStartId="F3" captionText="Figures 3 - 6. 3 Agra cruciaria Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 070044) 4 Agra grace Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 087638) 5 Agra pseuopusilla Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 070045) 6 Agra pusilla Chaudoir, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 060087)." httpUri="https://binary.pensoft.net/fig/14826" pageId="4" pageNumber="5">Figs 3</figureCitation>
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<figureCitation id="9090972876C41AA7BC10521209702316" captionStart="Figure 10" captionStartId="F6" captionText="Figure 10. Distribution map of the species of the pusilla group. Names marked with a "? " do not have precise localities on the specimen label (s)." httpUri="https://binary.pensoft.net/fig/14829" pageId="4" pageNumber="5">10</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="DFB1346843C73CB97F9044937F60D9C7" pageId="4" pageNumber="5" type="holotype">
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<paragraph id="46A30B0BBEDEEC05C10DA051CB600EC1" pageId="4" pageNumber="5">Holotype:</paragraph>
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<paragraph id="DCE9467142F6CD7EFC8252C3660BF25B" pageId="4" pageNumber="5">
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<emphasis id="3A031C6489A83E95C4093AFFAC760CB4" bold="true" pageId="4" pageNumber="5">Brazil</emphasis>
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: Rio de Janeiro, Rio de Janeiro, Corcovado, 585m, 22.9517°S, 43.2116°W, 5 May 1958 (C.A. Campos Seabra & M. Alvarenga)(NMNH: ADP 070044, male).
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</paragraph>
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</subSubSection>
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<subSubSection id="86E09C6AEFA1810E2B77479351EB7C16" pageId="4" pageNumber="5" type="derivation of specific epithet">
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<paragraph id="013B0AAD980BEAEEC470B80BBD04D975" pageId="4" pageNumber="5">Derivation of specific epithet.</paragraph>
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<paragraph id="B9ECEBB47F2331C229AF3A5596766324" pageId="4" pageNumber="5">
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The epithet "
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<emphasis id="5BAF864CDAD1ABAFBA583A9490330A13" italics="true" pageId="4" pageNumber="5">cruciaria</emphasis>
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" is a Latin adjective meaning "of/pertaining to the cross/torture" and is based upon the large cross on Corcovado flooded each night with high powered lights that attract insects by the millions and is the type locality of this species.
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</paragraph>
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</subSubSection>
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<subSubSection id="F5FF2399BC01C20A8F6C574B1EE06DF9" pageId="4" pageNumber="5" type="proposed english vernacular name">
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<paragraph id="696F9217BB7A53EF1E8273D5886C94C0" pageId="4" pageNumber="5">Proposed English vernacular name.</paragraph>
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<paragraph id="A5E9369393E93057588A1AE9FB9657E3" pageId="4" pageNumber="5">Cross Elegant Canopy Beetle.</paragraph>
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</subSubSection>
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<subSubSection id="0585B3402034325A0273F51295A12FE3" pageId="4" pageNumber="5" type="diagnosis">
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<paragraph id="F6761C5416F442001A87FA8E35C66FA1" pageId="4" pageNumber="5">Diagnosis.</paragraph>
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<paragraph id="CCA8405192E6455A9EE6AD082E159383" pageId="4" pageNumber="5">
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With the attributes of the genus and species-group as described above and medium sized for the
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<emphasis id="7C6FBC90B0E5A71A2C0E3F5FBC328CFA" italics="true" pageId="4" pageNumber="5">pusilla</emphasis>
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group. Adults with black integument; head behind eyes and prothorax with slight brassy reflections. Frons laterad unicarinate and rugose. Occiput with sparse punctures, some punctures with short setae.
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</paragraph>
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<paragraph id="7E57D666C17DC4ED297765DB2D05DA69" pageId="4" pageNumber="5">Elytra with moderately lobed apex.</paragraph>
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</subSubSection>
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<subSubSection id="F0993FDDACB5C06FA64905EDF4C34572" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="description">
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<paragraph id="B8D9C9A2551370DBA96E9425B5285B9C" pageId="4" pageNumber="5">Description.</paragraph>
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<paragraph id="784B9AE190DDF4DA293F97661D6B4E33" pageId="4" pageNumber="5">
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(
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<figureCitation id="AA3AB86AEB1DD7EE464DE385D5901751" captionStart="Figures 3–6" captionStartId="F3" captionText="Figures 3 - 6. 3 Agra cruciaria Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 070044) 4 Agra grace Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 087638) 5 Agra pseuopusilla Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 070045) 6 Agra pusilla Chaudoir, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 060087)." httpUri="https://binary.pensoft.net/fig/14826" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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).
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<emphasis id="4C25163FD3DFF884D4105B07DE70E190" italics="true" pageId="4" pageNumber="5">Size</emphasis>
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: Small, ABL = 6.4 - 7.7 mm, SBL = 5.75 - 6.48 mm, TW = 1.74 -1.8 mm.
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<emphasis id="AE502F48BF78BC72250088CD1ED39EBC" italics="true" pageId="4" pageNumber="5">Color:</emphasis>
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Head black with brassy reflection posteriorly, pronotum with brassy reflections; antennae blackish blue with scape testaceous and with slight metallic blue reflections, mouthparts piceous, and legs and tarsi blackish blue, with mostly testaceous femur.
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<emphasis id="54CA21A954060EA9C473A4D3A1BE179E" italics="true" pageId="4" pageNumber="5">Luster:</emphasis>
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Head, pronotum and legs shiny metallic, elytra matte smoky-blue.
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<emphasis id="716C1FC554D26E9588EFACE65FAB399E" italics="true" pageId="4" pageNumber="5">Head:</emphasis>
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Labrum moderately elongate and truncate apically, barely emarginate medially. Frons medially raised and smooth, laterally depressed and unicarinate and rugose. Gena with hind angles broadly rounded to constricted neck in males. Genae and occiput moderately densely and coarsely some punctures setiferous.
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</paragraph>
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<paragraph id="3650BDAB6583247BA22639307A895B17" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
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<emphasis id="399369FE70ADD4B0682F411BF99A34E5" italics="true" pageId="4" pageNumber="5">Prothorax:</emphasis>
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Slightly broader medially, flared basally; surface with dense punctures, many setiferous; lateral elongate callous with single row of setiferous punctures along middle.
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<emphasis id="5D8B4A32D1D5AC89A08C5AE145E881D8" italics="true" pageId="4" pageNumber="5">Pterothorax:</emphasis>
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Elytron markedly convex, intervals slightly costate, interneurs of rows of somewhat irregularly shaped punctures that are double in some places, apex slightly oblique and moderately lobate, apical dentation asymmetric, lateral tooth small, acute, sutural apex slightly produced, narrowly pointed. Metasternum sparsely setiferous in male.
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<emphasis id="FF78993C96E81F4BFE6F8A8542AF225E" italics="true" pageId="4" pageNumber="5">Legs:</emphasis>
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Normal in male.
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<emphasis id="50877CE4D11DCBB9C7D60F04BD98FF2C" italics="true" pageId="4" pageNumber="5">Abdomen</emphasis>
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: Abdominal sterna III to VII of male moderately and bilaterally setiferous; sternum VII of male barely emarginated, corners rounded.
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<emphasis id="5BBB3D0FF1156E3FEA1636DA5D336376" italics="true" pageId="4" pageNumber="5">Male genitalia:</emphasis>
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Phallus (
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<figureCitation id="9ED93626C604966C98BB17599AD766A1" captionStart="Figures 3–6" captionStartId="F3" captionText="Figures 3 - 6. 3 Agra cruciaria Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 070044) 4 Agra grace Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 087638) 5 Agra pseuopusilla Erwin, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 070045) 6 Agra pusilla Chaudoir, sp. n., male genitalia (dorsal, ventral, left lateral aspects) (ADP 060087)." httpUri="https://binary.pensoft.net/fig/14826" pageId="4" pageNumber="5">Fig. 3</figureCitation>
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) elongate and narrow with ostium elongate, nearly half the length of the phallus, apex a narrowly lobate expansion of distal end, this slipper-shaped
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<pageBreakToken id="5ECFEEC1D03D0940E8C87DC5775789D4" pageId="5" pageNumber="6" start="start">in</pageBreakToken>
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lateral aspect. Parameres small, left twice the size of the right, both broadly rounded.
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<emphasis id="7103EE0F09EA8D87B998D40D86CE1610" italics="true" pageId="5" pageNumber="6">Female ovipositor:</emphasis>
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Female unknown.
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</paragraph>
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</subSubSection>
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<subSubSection id="4981216332E1B7862FB11EEA53562090" pageId="5" pageNumber="6" type="dispersal potential">
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<paragraph id="6F37E43ED9E28639056DD63FA52C6DCB" pageId="5" pageNumber="6">Dispersal potential.</paragraph>
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<paragraph id="B20B14685094E1DD50192421FDDCC3F4" pageId="5" pageNumber="6">These beetles are macropterous and are probably capable of flight; they are swift and agile runners.</paragraph>
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</subSubSection>
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<subSubSection id="5D898B73789037F08B67E02A01AEC3D0" pageId="5" pageNumber="6" type="way of life">
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<paragraph id="C3D1B590995FB79DCE132040CBCB52BA" pageId="5" pageNumber="6">Way of life.</paragraph>
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<paragraph id="9CA33CCDDD4BE4C05BD04337B18DD9B2" pageId="5" pageNumber="6">
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Adults of other
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<taxonomicName id="8B67365FF06B3A40C8A42C09BA85AA42" authorityName="Fabricius" authorityYear="1801" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Agra</taxonomicName>
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species are found in the canopy of rainforest trees; larvae of this genus are found under the bark of these trees, however they must also roam on the surface, as they have been collected by insecticidal fogging techniques in the very early morning before first light. Members of
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<taxonomicName id="71257F07B6E258C4F2C2FAA6622FA638" authorityName="Erwin" authorityYear="2010" class="Insecta" family="Carabidae" genus="Agra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="cruciaria">Agra cruciaria</taxonomicName>
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occur at midland altitudes in the Mata
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<normalizedToken id="B28DBAC52A61A5C596A757B971078AB4" originalValue="Atlântica">Atlantica</normalizedToken>
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. Adults are active in May, the rainy season.
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</paragraph>
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</subSubSection>
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<subSubSection id="6F7733844F517EB057CBCF4E78668730" pageId="5" pageNumber="6" type="other specimens examined">
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<paragraph id="C1617221D67A467C8A8D6588357A62F6" pageId="5" pageNumber="6">Other specimens examined.</paragraph>
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<paragraph id="CF30D24FA122E864CE2F93AAD70E49BE" pageId="5" pageNumber="6">
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<emphasis id="415BA676CDF8037A035DA182ECACC1A4" bold="true" pageId="5" pageNumber="6">Brazil</emphasis>
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: Minas Gerais, (MNHNP: ADP 060040, male paratype).
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</paragraph>
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</subSubSection>
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<subSubSection id="223C3F5CE1177C601BD7FBF35A9B1B29" pageId="5" pageNumber="6" type="geographic distribution">
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<paragraph id="1DAC2332EA76F97C17C27C3E2AADEF22" pageId="5" pageNumber="6">Geographic distribution.</paragraph>
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<paragraph id="A1200E378E23EF7BBACD2941D3D5AF90" pageId="5" pageNumber="6">
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(
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<figureCitation id="1D39895A09B264BFDAC630F04D42FE92" captionStart="Figure 10" captionStartId="F6" captionText="Figure 10. Distribution map of the species of the pusilla group. Names marked with a "? " do not have precise localities on the specimen label (s)." httpUri="https://binary.pensoft.net/fig/14829" pageId="5" pageNumber="6">Fig. 10</figureCitation>
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). This species is currently known from the type locality and an unknown location in the State of Minas Gerais.
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</paragraph>
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</subSubSection>
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<subSubSection id="FA5D49353E7137D7B574E269164BBFDC" pageId="5" pageNumber="6" type="notes">
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<paragraph id="CAE2FE61F2FF1B0AD148CB45E947CF47" pageId="5" pageNumber="6">Notes.</paragraph>
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<paragraph id="C676E67000CFA732D5D5378345103319" pageId="5" pageNumber="6">Right antenna glued to card of holotype not of this specimen.</paragraph>
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</subSubSection>
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</treatment>
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</document> |