878 lines
117 KiB
XML
878 lines
117 KiB
XML
<document ID-CLB-Dataset="24202" ID-DOI="http://dx.doi.org/10.3897/zse.96.48312" ID-GBIF-Dataset="7de3ea60-ba35-4831-ae1d-048dc362f8b0" ID-Pensoft-Pub="1860-0743-1-25" ID-Pensoft-UUID="4EDA9B5B777551B7BBB0152DD188E5EF" ID-ZooBank="F2C8585A1268443693348B64AE20F6EE" ModsDocID="1860-0743-1-25" checkinTime="1580744411276" checkinUser="pensoft" docAuthor="Sands, Arthur F, Gloeer, Peter, Guerlek, Mustafa E, Albrecht, Christian & Neubauer, Thomas A" docDate="2020" docId="55E91AC28D0857A69BA932BD100648AE" docLanguage="en" docName="ZoosystEvol 96(1): 25-66" docOrigin="Zoosystematics and Evolution 96 (1)" docSource="http://dx.doi.org/10.3897/zse.96.48312" docTitle="Theodoxus wesselinghi Sands & Gloeer 2020, sp. nov." docType="treatment" docUuid="97AF034D-EF6D-4AB2-AC17-0C5B037C8DAD" docUuidSource="ZooBank" docVersion="8" id="4EDA9B5B777551B7BBB0152DD188E5EF" lastPageNumber="25" masterDocId="4EDA9B5B777551B7BBB0152DD188E5EF" masterDocTitle="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species" masterLastPageNumber="66" masterPageNumber="25" pageNumber="25" updateTime="1732731517510" updateUser="ExternalLinkService">
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<mods:title id="4AC6F62FAF06B26C97F1EDC2987D3FB3">A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species</mods:title>
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<mods:name id="9735339002597A2A333E51512A03F635" type="personal">
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<mods:roleTerm id="E17055DD39716A6EB896D5E125C204D2">Author</mods:roleTerm>
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<mods:namePart id="3B1F440218E5089CB85E2D4440157B27">Sands, Arthur F</mods:namePart>
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<mods:affiliation id="B5CB2688C83CF01C22D750B9C70ECA15">Justus Liebig Univrsity Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="11C77ADE8CAC15A07F2737910413DCEE">Gloeer, Peter</mods:namePart>
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<mods:affiliation id="E1D13A4E180C6C8EAE829C8681A102E8">Biodiversity Research Laboratory, Hetlingen, Germany</mods:affiliation>
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<mods:roleTerm id="B116135483C0E5D2ED5D034585589CC4">Author</mods:roleTerm>
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<mods:namePart id="6E78561366D1C2CDFA1B5911B81F6BD2">Guerlek, Mustafa E</mods:namePart>
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<mods:affiliation id="7CDCF6DF9CA86163963C25F56376C5AE">Mehmet Akif Ersoy niversity, Burdur, Turkey</mods:affiliation>
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<mods:namePart id="E4DB8B70F7F96339477BDD184304E148">Albrecht, Christian</mods:namePart>
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<mods:affiliation id="83D2C6723F0DC4C069EB22912504DFC9">Justus Liebig University Giessen, Giessen, Germany</mods:affiliation>
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<mods:namePart id="5217DA34114BF2B8EA5F061B4A5BC00F">Neubauer, Thomas A</mods:namePart>
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<mods:affiliation id="24F449C5B8EC6EBFF222BBF113CDA379">Justus Liebig University Giessen, Giessen, Germany & Naturalis Biodiversity Centre, Leiden, Netherlands</mods:affiliation>
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<mods:title id="2DFF311C90D0D94FE7CDB325754A4293">Zoosystematics and Evolution</mods:title>
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<mods:date id="C5AA9AB385C19AA874243CC163A256FA">2020</mods:date>
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<mods:number id="BE746E1B5325BF3E23C7645618AC5489">96</mods:number>
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<mods:url id="4E9C641022682332BD91DD5D6730571F">http://dx.doi.org/10.3897/zse.96.48312</mods:url>
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<mods:classification id="D21BA417F962F36769102D481859C44F">journal article</mods:classification>
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<mods:identifier id="CA40840CDF621B2D88FC014A4C849C6D" type="DOI">http://dx.doi.org/10.3897/zse.96.48312</mods:identifier>
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<treatment id="55E91AC28D0857A69BA932BD100648AE" ID-GBIF-Taxon="161939051" LSID="urn:lsid:zoobank.org:act:97AF034D-EF6D-4AB2-AC17-0C5B037C8DAD" httpUri="http://treatment.plazi.org/id/55E91AC28D0857A69BA932BD100648AE" lastPageNumber="25" pageId="0" pageNumber="25" scope_class="Gastropoda" scope_family="Neritidae" scope_subFamily="Neritininae" scope_superFamily="Neritoidea">
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<subSubSection id="01863E517C017B435673BACC88ABB007" pageId="0" pageNumber="25" type="nomenclature">
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<paragraph id="13D467788E40CDE767252C918554C8F4" pageId="0" pageNumber="25">
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<taxonomicName id="69E57C4E554FFA2DE1B8FB33392F28DB" ID-CoL="7TCKX" LSID="55E91AC2-8D08-57A6-9BA9-32BD100648AE" authority="Sands & Glöer" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi" status="sp. nov.">
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Theodoxus wesselinghi Sands &
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<normalizedToken id="31711BF8FB31CFCDDC5C7CE937962B8F" originalValue="Glöer">Gloeer</normalizedToken>
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</taxonomicName>
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<taxonomicNameLabel id="B8291A20ED0559209E3A718C9230C568" pageId="0" pageNumber="25">sp. nov.</taxonomicNameLabel>
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<figureCitation id="C8B338560C2042B6539D5DD9735D27A0" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">Figures 23A-H</figureCitation>
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<figureCitation id="3269137C64B367FED047D952D9B6E164" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">, 24A-M</figureCitation>
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<figureCitation id="FFBB46DF774C6F8A463C24D2C64A3176" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">, 25A-F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="58DF69A03F770C9330A1D73923CE4F71" pageId="0" pageNumber="25" type="reference_group">
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<paragraph id="83430A054A90035839F1AC7A0A54FF8F" pageId="0" pageNumber="25">
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<taxonomicName id="04CE967A377BBCC2C24DF6E0931B9F00" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Theodoxus fluviatilis</taxonomicName>
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:
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<bibRefCitation id="BB442D3D130AD66AC6416F5A081132EC" author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
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<normalizedToken id="386CAA0F8853ABB65BA0525E0A53F30B" originalValue="Odabaşı">Odabasi</normalizedToken>
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and Arslan 2015
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</bibRefCitation>
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||
: 330-331 (
|
||
<emphasis id="4DBF2332FDB988F1B88AF3435785271B" bold="true" pageId="0" pageNumber="25">non</emphasis>
|
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<taxonomicName id="F9E408A5C918808BC480061D33A04507" authorityName="Linnaeus" authorityYear="1758" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita fluviatilis" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="fluviatilis">Nerita fluviatilis</taxonomicName>
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Linnaeus, 1758).
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||
</paragraph>
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<paragraph id="7C0CBD0F510F423FF286930928658AFC" pageId="0" pageNumber="25">
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<taxonomicName id="DCDD566DA9B02E0D488A8CE039ED9686" baseAuthorityName="Recluz" baseAuthorityYear="1841" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus anatolicus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="anatolicus">Theodoxus anatolicus</taxonomicName>
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:
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<bibRefCitation id="2D776418F3868CABD6800123A3350BCA" author="Yildirim, MZ" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="115 - 119" refId="B163" refString="Yildirim, MZ, Koca, SB, Guerlek, ME, Gloeer, P, 2018. A new genus and a species from Turkey, Isparta felei n. gen. n. sp.(Gastropoda: Truncatelloidea: Hydrobiidae). Ecologica Montenegrina 18: 115 - 119" title="A new genus and a species from Turkey, Isparta felei n. gen. n. sp. (Gastropoda: Truncatelloidea: Hydrobiidae)." volume="18" year="2018">
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<normalizedToken id="841C546199A2AB44E0EB217E03A9DCB4" originalValue="Yıldırım">Yildirim</normalizedToken>
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et al. 2018
|
||
</bibRefCitation>
|
||
: 118 (
|
||
<emphasis id="6179E05619329FA36B05D5C8271E183E" bold="true" pageId="0" pageNumber="25">non</emphasis>
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<taxonomicName id="700610A2CCE66F986FBD4EB9E597F4F2" authorityName="Recluz" authorityYear="1841" class="Gastropoda" family="Neritidae" genus="Nerita" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nerita anatolica" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="anatolica">Nerita anatolica</taxonomicName>
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<normalizedToken id="0F0F50549DE2657685F0D7F21CD7585F" originalValue="Récluz">Recluz</normalizedToken>
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, 1841).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="33655C6BB4A9ACE2678D6E0E997899E0" pageId="0" pageNumber="25" type="type locality">
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||
<paragraph id="CF615F96CC60085CDBCDF92D0C5AB1B0" pageId="0" pageNumber="25">
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<typeStatus id="574BF8CDDDEC954DD82B467307B79613">Type</typeStatus>
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locality.
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</paragraph>
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<paragraph id="E89D21A7EA46306213493C25760F78E0" pageId="0" pageNumber="25">
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Sakarya River,
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<normalizedToken id="2519D0AA683F45A94BDED42073BEFBE8" originalValue="Çayköy">Caykoey</normalizedToken>
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, Bilecik, Turkey;
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<geoCoordinate id="17CCADC308D798ADDB1D358D3C8F977B" degrees="40.0439" direction="north" orientation="latitude" precision="5" value="40.0439">40.0439°N</geoCoordinate>
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,
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<geoCoordinate id="92F5DEACA461E232F1866026C31E4E72" degrees="30.452" direction="east" orientation="longitude" precision="55" value="30.452">30.452°E</geoCoordinate>
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(Figs
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<figureCitation id="49FB7AE03E37EC1A0038013F963F3018" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3D</figureCitation>
|
||
,
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||
<figureCitation id="15D160BA904E4986805EFE0EE0F29622" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23A, B</figureCitation>
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).
|
||
</paragraph>
|
||
<paragraph id="5B2F3BD582B64156657C8423564E7E1B" pageId="0" pageNumber="25">
|
||
<materialsCitation id="BC13CBBEE20857B4B83706A6D5B9B368" collectionCode="RMNH" country="Turkey" latitude="40.0439" longitude="30.452" stateProvince="Nmnl" typeStatus="Holotype">
|
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<emphasis id="6AB8CF2DF75461D4DDF6EEF93275F275" bold="true" pageId="0" pageNumber="25">
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||
<typeStatus id="9B72405B29AD866A9CCF3AE144D92F51">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectionCode id="7C90FE70330AC71AD94C4A4FEB1B4DD0" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
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||
<collectionCode id="1F78B383A843626762CF524E3AFCAFAE" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342200 (Sakarya River,
|
||
<normalizedToken id="D6A893CC4E4208800D3097223EA450DC" originalValue="Çayköy">Caykoey</normalizedToken>
|
||
, Bilecik,
|
||
<collectingCountry id="2FBF5CC902EACD528AFD87B61093377D" name="Turkey" value="Turkey">Turkey</collectingCountry>
|
||
;
|
||
<geoCoordinate id="BB0560F67F732A7BA22DB724C110166B" degrees="40.0439" direction="north" orientation="latitude" precision="5" value="40.0439">40.0439°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A93ACDCD093F14980E0B51605A307AC0" degrees="30.452" direction="east" orientation="longitude" precision="55" value="30.452">30.452°E</geoCoordinate>
|
||
) stored in
|
||
<collectingRegion id="4661CD28773D20263B6D919DAC478270">NMNL</collectingRegion>
|
||
: Shell height 6.0 mm, width 6.0 mm (Fig.
|
||
<figureCitation id="610F91FC2246BF1CDFBC5608475BF48F" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24A-D</figureCitation>
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||
)
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||
</materialsCitation>
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||
.
|
||
</paragraph>
|
||
<paragraph id="DB933C65C0FD6B57BC752242A70C74C7" pageId="0" pageNumber="25">
|
||
<materialsCitation id="49775681BDD8E690A35C22A38ECC7D25" collectionCode="A" country="Turkey" latitude="40.0439" longitude="30.452" typeStatus="Paratypes">
|
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<emphasis id="FB55F5EC4019A1493C965D9A2846F9B4" bold="true" pageId="0" pageNumber="25">
|
||
<typeStatus id="919B9F8839FD729BF3567945ABA38FD8">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Twenty-four specimens from Sakarya River,
|
||
<normalizedToken id="BD3457FEE15A4A1F36AFE7E172873183" originalValue="Çayköy">Caykoey</normalizedToken>
|
||
, Bilecik,
|
||
<collectingCountry id="DF828705BA1984D18D6865D9905732E4" name="Turkey" value="Turkey">Turkey</collectingCountry>
|
||
;
|
||
<geoCoordinate id="F6BC7C5399703F9102EEF3605C6DC48A" degrees="40.0439" direction="north" orientation="latitude" precision="5" value="40.0439">40.0439°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="10E2761D308F8D00ED3F5A858D167CF3" degrees="30.452" direction="east" orientation="longitude" precision="55" value="30.452">30.452°E</geoCoordinate>
|
||
(Fig.
|
||
<figureCitation id="A2425ED2373036A76E8052E6E81440A4" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23A, B</figureCitation>
|
||
):
|
||
<quantity id="03D21B1E05F244AD502D584A06CB5DD6" metricMagnitude="-1" metricUnit="m" metricValue="2.794" unit="in" value="11.0">11 in</quantity>
|
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<collectionCode id="8289359E556FC6EFE65111672558E8EA" collectionName="NMNL" country="Netherlands" httpUri="http://grbio.org/cool/1yj9-ymxi" name="Natuurmuseum Nijmegen e.o." type="Herbarium">NMNL</collectionCode>
|
||
(
|
||
<collectionCode id="99F6387FFE14AC9889E26926A6E1AC32" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="470CE8973C13167A3D60A8D198D38C6A" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342201,
|
||
<collectionCode id="7462DD1222651090EB4EC9F73DEF1614" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="AF814BED6DC6B349F7E902ABE516860F" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342202; Fig.
|
||
<figureCitation id="DDFC7C92DFC285F13A998B7DBF280EBF" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24E-G</figureCitation>
|
||
) and
|
||
<quantity id="3B1C46743E9B21C379E0D670C752F416" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
|
||
<collectionCode id="2736AF90806D950C7A51869A61C9A9EF" collectionName="UGSB">UGSB</collectionCode>
|
||
(
|
||
<collectionCode id="0E0BA2DDC6EF6D3A97AAB0561C3D3D7C" collectionName="UGSB">UGSB</collectionCode>
|
||
20688,
|
||
<collectionCode id="C4D1212F4BE6007369CAE5590D1F5727" collectionName="UGSB">UGSB</collectionCode>
|
||
20743,
|
||
<collectionCode id="F68FD3FAA71BE426B45A879A94890F9D" collectionName="UGSB">UGSB</collectionCode>
|
||
20744; Fig.
|
||
<figureCitation id="704CFBB1EB926D01C502614499B0C173" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">25A, B, D</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="6FE9EC45A58A0CB1AC94EF68A8F8B0B2" collectionCode="C" country="Turkey" latitude="38.00358" longitude="31.47217">
|
||
Twenty-four specimens from an unnamed roadside spring in Fele,
|
||
<normalizedToken id="AD07A2BF4887DD1734FA1F37A6A44834" originalValue="İsparta">Isparta</normalizedToken>
|
||
province,
|
||
<collectingCountry id="2E8695587E5D110DF26FFC20288FA087" name="Turkey" value="Turkey">Turkey</collectingCountry>
|
||
;
|
||
<geoCoordinate id="0C8FE79EB1F8FEF79EEDEF54669964ED" degrees="38.00358" direction="north" orientation="latitude" precision="1" value="38.00358">38.00358°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="54EC39AC5BF8A4A098EA56CA91793541" degrees="31.47217" direction="east" orientation="longitude" precision="1" value="31.47217">31.47217°E</geoCoordinate>
|
||
(Fig.
|
||
<figureCitation id="AE35F184C7C87A8934DDFB758EC0E66D" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23C, D</figureCitation>
|
||
):
|
||
<quantity id="383A765EF5BE4C917C517A44DDC64AA7" metricMagnitude="-1" metricUnit="m" metricValue="2.794" unit="in" value="11.0">11 in</quantity>
|
||
<collectionCode id="FC37DBFF8442A5A86C750FC9DEB932E2" collectionName="NMNL" country="Netherlands" httpUri="http://grbio.org/cool/1yj9-ymxi" name="Natuurmuseum Nijmegen e.o." type="Herbarium">NMNL</collectionCode>
|
||
(
|
||
<collectionCode id="73E7903DB48F6753C02BFB7C7486738B" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="F4B301241DB31475DC7992E7A8B64AFF" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342203,
|
||
<collectionCode id="2D26AAA2E007E30F2999A9E6CE9E6BDF" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="8802A414BCECF89D16BE687C89B96DF7" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342204; Fig.
|
||
<figureCitation id="397468C1970787A883E6195692388BC4" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24H-J</figureCitation>
|
||
) and
|
||
<quantity id="B3B15A33D9C37A85B51E61BF66FEFAA4" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
|
||
<collectionCode id="A7F0C9431DB1AD78D795D7081124255E" collectionName="UGSB">UGSB</collectionCode>
|
||
(
|
||
<collectionCode id="A81ACBD51A8A1889A72662CAF515E89F" collectionName="UGSB">UGSB</collectionCode>
|
||
20684,
|
||
<collectionCode id="14EDD8FD885343553CFBCF8BCFD8E413" collectionName="UGSB">UGSB</collectionCode>
|
||
20735,
|
||
<collectionCode id="FD7F833E633EA49F228C13754EDD7AD0" collectionName="UGSB">UGSB</collectionCode>
|
||
20736)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="7E4252BD7424838ED28F3238A0DD02C5" collectionCode="E" country="Turkey" latitude="37.8256" longitude="31.6748">
|
||
Twenty-five specimens from Eflatun
|
||
<normalizedToken id="84C8E7D58CD4729D1C22A846B217E93E" originalValue="Pınarı">Pinari</normalizedToken>
|
||
, near
|
||
<normalizedToken id="277F63E401BFD5E26F333B637AAD8CD9" originalValue="Sadıkhacı">Sadikhaci</normalizedToken>
|
||
, Konya,
|
||
<collectingCountry id="E80858D578C721AD43DBA8C2BE8ED16F" name="Turkey" value="Turkey">Turkey</collectingCountry>
|
||
;
|
||
<geoCoordinate id="25E2A0C690B7DE201CB5ADE055621F1D" degrees="37.8256" direction="north" orientation="latitude" precision="5" value="37.8256">37.8256°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9F1135D2D90FF9A4ABD4B5A8396612BA" degrees="31.6748" direction="east" orientation="longitude" precision="5" value="31.6748">31.6748°E</geoCoordinate>
|
||
(Fig.
|
||
<figureCitation id="06EBB289F843C0A6B093C91AB14FDF1F" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23E, F</figureCitation>
|
||
):
|
||
<quantity id="17A1FC4262ABE55AD3206C2C5361BF8F" metricMagnitude="-1" metricUnit="m" metricValue="2.794" unit="in" value="11.0">11 in</quantity>
|
||
<collectionCode id="5347EC91E3A9462780666A752BD69FC4" collectionName="NMNL" country="Netherlands" httpUri="http://grbio.org/cool/1yj9-ymxi" name="Natuurmuseum Nijmegen e.o." type="Herbarium">NMNL</collectionCode>
|
||
(
|
||
<collectionCode id="0B32C14329A44841CB9C86B200FD03EF" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="18407054B2FD4C52755023E75F00F0CE" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342205,
|
||
<collectionCode id="652E323DBECEAD655F866BC12E2FA861" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="67DEDB5458E5546B638889872FD72E6D" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342206; Fig.
|
||
<figureCitation id="4E8B7177121FC25680F5A47481CD264E" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24K-M</figureCitation>
|
||
) and
|
||
<quantity id="0EF7A699B218A2A1C0DB76FF867C7648" metricMagnitude="-1" metricUnit="m" metricValue="3.556" unit="in" value="14.0">14 in</quantity>
|
||
<collectionCode id="C67E2B3B068463D9A77A83A067FA70DB" collectionName="UGSB">UGSB</collectionCode>
|
||
(
|
||
<collectionCode id="CBA64711F547ECAFD9FEF1C852CBDF59" collectionName="UGSB">UGSB</collectionCode>
|
||
20685,
|
||
<collectionCode id="13E250551C3D5D2AABFC6F0C0C97A499" collectionName="UGSB">UGSB</collectionCode>
|
||
20737,
|
||
<collectionCode id="F46F96AFF67D5F9B09E4B6DDFE02DDD5" collectionName="UGSB">UGSB</collectionCode>
|
||
20738; Fig.
|
||
<figureCitation id="6D517E110BECE792A88316029A695AB5" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">25F</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="8DDDDDA2855EBA6D2455479E0D85FB71" collectionCode="G" country="Turkey" latitude="39.15277" longitude="31.61562">
|
||
Thirty specimens from
|
||
<normalizedToken id="D208ED8350B79DA415B5FD4F6E53A3D6" originalValue="Balıkdamı">Balikdami</normalizedToken>
|
||
Wetland spring,
|
||
<normalizedToken id="69179728953EC0627D7E779B2AC25F0F" originalValue="Eskişehir">Eskisehir</normalizedToken>
|
||
,
|
||
<collectingCountry id="4E45EECD2D5AF6CA3C33EED3D6D0CAA2" name="Turkey" value="Turkey">Turkey</collectingCountry>
|
||
;
|
||
<geoCoordinate id="602C9B69687BCC9FC344D9B50CE046BF" degrees="39.15277" direction="north" orientation="latitude" precision="1" value="39.15277">39.15277°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="0BF5C9F34FAA75AE913CC5B29C8F585A" degrees="31.61562" direction="east" orientation="longitude" precision="1" value="31.61562">31.61562°E</geoCoordinate>
|
||
(Fig.
|
||
<figureCitation id="0B4DDE7324236B7A2EC7405264EC6565" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23G, H</figureCitation>
|
||
):
|
||
<quantity id="0EBB75FE8C214D982335C1C914007338" metricMagnitude="-1" metricUnit="m" metricValue="3.302" unit="in" value="13.0">13 in</quantity>
|
||
<collectionCode id="9810B7075DBF6E2FA71034BF31CE5F7A" collectionName="NMNL" country="Netherlands" httpUri="http://grbio.org/cool/1yj9-ymxi" name="Natuurmuseum Nijmegen e.o." type="Herbarium">NMNL</collectionCode>
|
||
(
|
||
<collectionCode id="B44A16B192BF76A5671A6E813487D018" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="067B8D86B0838732123618F441C824FE" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342207) and
|
||
<quantity id="7A048B09FF11E19D5A74366646EA7010" metricMagnitude="-1" metricUnit="m" metricValue="4.318" unit="in" value="17.0">17 in</quantity>
|
||
<collectionCode id="7D78BE349EB7A41E8AFFD7808DFFD620" collectionName="UGSB">UGSB</collectionCode>
|
||
(
|
||
<collectionCode id="11C2902516C293CA6D3A9F31AED7712C" collectionName="UGSB">UGSB</collectionCode>
|
||
20686,
|
||
<collectionCode id="D37A7AB67649A53A968175FA7FC08DE9" collectionName="UGSB">UGSB</collectionCode>
|
||
20739,
|
||
<collectionCode id="A29ED5B6E939334A9AD8CF54632A8854" collectionName="UGSB">UGSB</collectionCode>
|
||
20740; Fig.
|
||
<figureCitation id="D3EC2E76841B58302A29467A531075FF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">25C, E</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="E9060704B4FB6216FC5A57B5CBAB6A52" doi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25" start="Figure 23" startId="F23">
|
||
<paragraph id="87A4D4DFC971D3F1C05425925B62DFCC" pageId="0" pageNumber="25">
|
||
<emphasis id="483D639DDA0BF7630F8C17F410EF337F" bold="true" pageId="0" pageNumber="25">Figure 23.</emphasis>
|
||
<typeStatus id="CE02C59C44CF8A0AC490E577C8AE4A02">Type</typeStatus>
|
||
locality of
|
||
<taxonomicName id="2CABB854423ABDCDB78D8C1EFE27DFD5" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi">
|
||
<emphasis id="A93C38FD0EFFF323E56C10E5A96AE684" italics="true" pageId="0" pageNumber="25">Theodoxus wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
Sands &
|
||
<normalizedToken id="11543B756578E8B59F5C5267E94546CD" originalValue="Glöer">Gloeer</normalizedToken>
|
||
sp. nov.
|
||
<emphasis id="67CAE78263E746792A97B9BC836B0F27" bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
,
|
||
<emphasis id="3F46CAF15B4AB98ECC40D0ED85FA106A" bold="true" pageId="0" pageNumber="25">B</emphasis>
|
||
. Sakarya River,
|
||
<normalizedToken id="5F35F4A39913C0BD1D7AD2E75EBD2C97" originalValue="Çayköy">Caykoey</normalizedToken>
|
||
, Bilecik, Turkey,
|
||
<geoCoordinate id="C345DC1BF979E4A21BC6324B69C8A158" degrees="40.0439" direction="north" orientation="latitude" precision="5" value="40.0439">40.0439°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="A5D59C7884501F9BCF14ACE7D3F40EAC" degrees="30.452" direction="east" orientation="longitude" precision="55" value="30.452">30.452°E</geoCoordinate>
|
||
. Further
|
||
<typeStatus id="ECF7ACF81456E4800350035362E3F51C">paratype</typeStatus>
|
||
localities:
|
||
<emphasis id="40C761F31AC41F5489563205CC129567" bold="true" pageId="0" pageNumber="25">C</emphasis>
|
||
,
|
||
<emphasis id="EC82C8C0994E46192AB70AA272F2FD74" bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
. Unnamed roadside spring in Fele,
|
||
<taxonomicName id="AA6838117FCF34973CF53EA01E3D4D2B" authorityName="Yildirim, Koca, Gurlek & Gloer" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Isparta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isparta" order="Littorinimorpha" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Isparta</taxonomicName>
|
||
province, Turkey,
|
||
<geoCoordinate id="AABA2455082D7D48B28F60C646A2D71A" degrees="38.00358" direction="north" orientation="latitude" precision="1" value="38.00358">38.00358°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="AEC21302DFD335A493FF537728E8EF0D" degrees="31.47217" direction="east" orientation="longitude" precision="1" value="31.47217">31.47217°E</geoCoordinate>
|
||
;
|
||
<emphasis id="EE218300B98E9FE257622AB851CEAA33" bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
,
|
||
<emphasis id="CB86727ABDDC7FDDA835C05E0154F1FB" bold="true" pageId="0" pageNumber="25">F</emphasis>
|
||
. Eflatun
|
||
<normalizedToken id="B0880F291EE2ACAEFA7C6552E251173B" originalValue="Pınarı">Pinari</normalizedToken>
|
||
, near
|
||
<normalizedToken id="CDDD3ECE379E95DE78BEE428D75245E7" originalValue="Sadıkhacı">Sadikhaci</normalizedToken>
|
||
, Konya, Turkey,
|
||
<geoCoordinate id="6B28C42199EBBBCD505C19D77CE4FBC9" degrees="37.8256" direction="north" orientation="latitude" precision="5" value="37.8256">37.8256°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="235572C84D0E4CD10213C5F45881D576" degrees="31.6748" direction="east" orientation="longitude" precision="5" value="31.6748">31.6748°E</geoCoordinate>
|
||
;
|
||
<emphasis id="5F6D3C120213FB5788D87BD80A6F288F" bold="true" pageId="0" pageNumber="25">G</emphasis>
|
||
,
|
||
<emphasis id="94642063BA60E9FFFB07258EEF65735F" bold="true" pageId="0" pageNumber="25">H</emphasis>
|
||
.
|
||
<normalizedToken id="C654BCD4FA9878EE892BE20589534CB4" originalValue="Balıkdamı">Balikdami</normalizedToken>
|
||
Wetland spring,
|
||
<normalizedToken id="14627CDF1F5262E4D19BF64CE38099FB" originalValue="Eskişehir">Eskisehir</normalizedToken>
|
||
, Turkey,
|
||
<geoCoordinate id="3C926675A807008344DAB18CA2F25CBA" degrees="39.15277" direction="north" orientation="latitude" precision="1" value="39.15277">39.15277°N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="9EEF1E2E3DC172BFEF60D405CC6885E4" degrees="31.61562" direction="east" orientation="longitude" precision="1" value="31.61562">31.61562°E</geoCoordinate>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="626F4E4DE4AF402C1BECBD94F9D19D61" doi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25" start="Figure 24" startId="F24">
|
||
<paragraph id="3195CDE601509AAD3B28CE5A3DE284DD" pageId="0" pageNumber="25">
|
||
<emphasis id="903E482D5D64064662422F09374E1B21" bold="true" pageId="0" pageNumber="25">Figure 24.</emphasis>
|
||
<taxonomicName id="98B25E16505A31F4CC7151ABA25797DD" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi">
|
||
<emphasis id="C7D2D18ED8F14F7CDD02892074E684F6" italics="true" pageId="0" pageNumber="25">Theodoxus wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
Sands &
|
||
<normalizedToken id="7E9CD5C1A54D6A53BAB3F07CAE9E6944" originalValue="Glöer">Gloeer</normalizedToken>
|
||
sp. nov.
|
||
<emphasis id="2684A9AA817A6D10CDDEA57F4125DE35" bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
-
|
||
<emphasis id="9EF54943EF4F08470606C14586334A1B" bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
.
|
||
<typeStatus id="D4F50E72A5950DF193575A9EC805B9D8">Holotype</typeStatus>
|
||
collected in the Sakarya River,
|
||
<normalizedToken id="D3A8D262198DFAED25A6E0F84B5F45BB" originalValue="Çayköy">Caykoey</normalizedToken>
|
||
, Bilecik, Turkey (
|
||
<collectionCode id="5B332F54EC96ED2F9095177A6C6CCF01" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="FC73BBF34AA3BADEF03616E3D17E06BF" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342200);
|
||
<emphasis id="BAD2D93CDA2BA518DE2588116CD11E9E" bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
-
|
||
<emphasis id="12AEB8712DF77E16A7B1C9A2385CA0D9" bold="true" pageId="0" pageNumber="25">G</emphasis>
|
||
.
|
||
<typeStatus id="3766E085E383CA7912D5D41107399673">Paratype</typeStatus>
|
||
from the same location as the
|
||
<typeStatus id="C23DA10B9C713CC3A261C75A09F2F394">holotype</typeStatus>
|
||
(
|
||
<collectionCode id="771C509A78296A8A7DF4B7C2FEF0BE89" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="5C6FFCE9A9E2324A54345FA0CBC0F13B" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342202);
|
||
<emphasis id="7E1D0E0B5310758C9E04C5F3CC872D23" bold="true" pageId="0" pageNumber="25">H</emphasis>
|
||
-
|
||
<emphasis id="2FBDFC7D8FD0C21D7147365231ABE38E" bold="true" pageId="0" pageNumber="25">J</emphasis>
|
||
.
|
||
<typeStatus id="C1F28C70FED23824A87EDA2F5EFF655F">Paratype</typeStatus>
|
||
from Fele,
|
||
<taxonomicName id="4A08F36E180AF2ACCD0FBC12E5E7C42F" authorityName="Yildirim, Koca, Gurlek & Gloer" authorityYear="2018" class="Gastropoda" family="Hydrobiidae" genus="Isparta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isparta" order="Littorinimorpha" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">Isparta</taxonomicName>
|
||
province, Turkey (
|
||
<collectionCode id="9C9045CE106177992F337F6596071C96" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="E03E366667E53C490B843461849E478F" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342204);
|
||
<emphasis id="A857479A74AE69D198A3A4AA0EB7A74E" bold="true" pageId="0" pageNumber="25">K</emphasis>
|
||
-
|
||
<emphasis id="64290CD5C40624B4983F686661403C24" bold="true" pageId="0" pageNumber="25">M</emphasis>
|
||
.
|
||
<typeStatus id="A883918EC19DADE8570400B5968FBE4B">Paratype</typeStatus>
|
||
from Eflatun
|
||
<normalizedToken id="94D0DE5CE78C7B0E6261197E30DBC376" originalValue="Pınarı">Pinari</normalizedToken>
|
||
, Konya province, Turkey (
|
||
<collectionCode id="CF29F8C0710A07050230D522C525EBF3" collectionName="Netherlands, Leiden, Nationaal Natuurhistorische Museum ("Naturalis") [formerly Rijksmuseum van Natuurlijke Historie]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" type="Museum">RMNH</collectionCode>
|
||
.
|
||
<collectionCode id="C1506C6FF6361B970EA90D2F611659D2" collectionName="MOL" country="Peru" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15530" name="Universidad Nacional Agraria La Molina" type="Herbarium">MOL</collectionCode>
|
||
.342206). All photographed material (A-M) is stored in
|
||
<collectionCode id="EF313D24B92D5D8213CFBAFE2FDAD54E" collectionName="NMNL" country="Netherlands" httpUri="http://grbio.org/cool/1yj9-ymxi" name="Natuurmuseum Nijmegen e.o." type="Herbarium">NMNL</collectionCode>
|
||
. Scale bars:
|
||
<quantity id="D6471592474869BCDCF422AB4AAA3BB3" metricMagnitude="-3" metricUnit="m" metricValue="1.0" unit="mm" value="1.0">1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="0B762897D049C99E892858146900B473" doi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25" start="Figure 25" startId="F25">
|
||
<paragraph id="0A3C3D5661136AAF5431FFCC4058CBCA" pageId="0" pageNumber="25">
|
||
<emphasis id="A51DEEE419C0E6EE2B5B6C5804B128E2" bold="true" pageId="0" pageNumber="25">Figure 25.</emphasis>
|
||
Radula of
|
||
<taxonomicName id="5DEC9718D3C4A514C6CAF66F9D6B3C8F" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi">
|
||
<emphasis id="0FD10B2FA7A8D7E56B77B3E43CE1813F" italics="true" pageId="0" pageNumber="25">Theodoxus wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
Sands &
|
||
<normalizedToken id="DEE3A90459A3EC2FBEFFA2AED15A03EF" originalValue="Glöer">Gloeer</normalizedToken>
|
||
sp. nov.
|
||
<typeStatus id="0B8FC5AE9BE5D3916A10E7E04165EBAE">paratypes</typeStatus>
|
||
.
|
||
<emphasis id="8E6F050731FB1D05F8C69D71880F4031" bold="true" pageId="0" pageNumber="25">A</emphasis>
|
||
. Portion of the radula showing full sets of teeth (
|
||
<collectionCode id="31B74FB80347459806B91BA57EEF61A8" collectionName="UGSB">UGSB</collectionCode>
|
||
20688);
|
||
<emphasis id="9C4F03ABC0154A44E6C32ACBB5E6D5F4" bold="true" pageId="0" pageNumber="25">B</emphasis>
|
||
. Magnified view of the central teeth (
|
||
<collectionCode id="C829F8669A10E9B29F0CF02E44ABE6C2" collectionName="UGSB">UGSB</collectionCode>
|
||
20688);
|
||
<emphasis id="58599A14B01304440761BFCAB1211873" bold="true" pageId="0" pageNumber="25">C</emphasis>
|
||
. Magnified view of the lateral and marginal teeth (
|
||
<collectionCode id="6795511A896A33A74DFD3092152B205D" collectionName="UGSB">UGSB</collectionCode>
|
||
20686);
|
||
<emphasis id="5E8D9D5829C50385743361D436C04866" bold="true" pageId="0" pageNumber="25">D</emphasis>
|
||
. Magnified view of the first and second rows of marginal teeth (
|
||
<collectionCode id="0C63330D8B6B0B23866390B59C6090CB" collectionName="UGSB">UGSB</collectionCode>
|
||
20688);
|
||
<emphasis id="529A323DB0D3FAB9F32652558AFD35B1" bold="true" pageId="0" pageNumber="25">E</emphasis>
|
||
. Magnified view of the first row of marginal teeth (
|
||
<collectionCode id="A421CB1CD5886408915F93E8136E82EB" collectionName="UGSB">UGSB</collectionCode>
|
||
20686);
|
||
<emphasis id="044B85E6C0B5203AD4C472FB0575947D" bold="true" pageId="0" pageNumber="25">F</emphasis>
|
||
. Magnified view of the faces of inner marginal teeth belonging to the first row (
|
||
<collectionCode id="AAAD7C97B9D9D971448B2BD702BB0197" collectionName="UGSB">UGSB</collectionCode>
|
||
20685). Scale bars: 100
|
||
<normalizedToken id="6556E8FCAB34A842918A2755E9D4EF7F" originalValue="μm">μm</normalizedToken>
|
||
(A-E), 20
|
||
<normalizedToken id="1FDF218ED502C767ED630159AAC1CACE" originalValue="μm">μm</normalizedToken>
|
||
(F).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="B1CA1A247F058C927272E4C9B6A12167" pageId="0" pageNumber="25" type="etymology">
|
||
<paragraph id="DBFE0AB020A8DD7991E61201C9455FCC" pageId="0" pageNumber="25">Etymology.</paragraph>
|
||
<paragraph id="B5714FDEEC995DD48B80A5EB34B4625B" pageId="0" pageNumber="25">The species is named in honour of the molluscan palaeontologist Frank P. Wesselingh (Naturalis Biodiversity Center, Leiden, The Netherlands) for his contributions to malacology.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="10644A879D4FAC9C714571094F091B75" pageId="0" pageNumber="25" type="description">
|
||
<paragraph id="D518D44340057DC49C97DB4CD0BC280A" pageId="0" pageNumber="25">Description.</paragraph>
|
||
<paragraph id="756DA63597619973F8FE5131E534E4FA" pageId="0" pageNumber="25">
|
||
Shell (Fig.
|
||
<figureCitation id="875127EE7547AF2F1A870C05F630E424" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24A-C, E, F, H-M</figureCitation>
|
||
): Hemispherical, transversely elongate, consisting of typically three whorls that rapidly grow. Spire well defined, moderate height for
|
||
<taxonomicName id="2BD248EF038B82781E62835B306A7F1E" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="ABB09F3D75CAEBD5AE9DB0A1FA3883A9" italics="true" pageId="0" pageNumber="25">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
; often corroded along with other parts of shell. Shell height ranges from 4.0-
|
||
<quantity id="C2F20AE71BD7781A9422892AA1804941" metricMagnitude="-2" metricUnit="m" metricValue="6.8" unit="mm" value="68.0">6.8 mm</quantity>
|
||
, width from
|
||
<quantity id="415C60E9E1D3FD7C73A8B3EC3C955AC5" metricMagnitude="-2" metricUnit="m" metricValue="5.45" metricValueMax="7.1" metricValueMin="3.8" unit="mm" value="54.5" valueMax="71.0" valueMin="38.0">3.8-7.1 mm</quantity>
|
||
. Juveniles appear more globular. Periostracum is uniformly ivory or solid black, intermediate forms with broad brown-black smudged diagonal stripes also exist; surface glossy, finely striated with growth lines. Aperture semicircular, no serrations on inner lip. Columellar plate smooth, flat to slightly concave, inclined towards aperture; colouration blue-grey in darker shelled individuals to white in lighter forms.
|
||
</paragraph>
|
||
<paragraph id="E186C63498715E841B81105ED68A14D8" pageId="0" pageNumber="25">
|
||
Operculum (Fig.
|
||
<figureCitation id="C7C836B6B68C9D2F541AE1B2C9A40F26" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24D, G</figureCitation>
|
||
): Operculum plate made of two parts, calcareous base and conchioline lamella; operculum base light, mostly ivory to white, white lamella with distinct orange edge on border with operculum base. Operculum base left adductor can be blunt and rounded, weak callus on top right edge. Apophysis follows same colour scheme as operculum calcareous base, broader at top and narrower and attenuated at bottom. Very narrow rib-shield, deep rib-pouch present on operculum. Operculum lacks a pseudo-apophysis.
|
||
</paragraph>
|
||
<paragraph id="51ADBDA87A04D57856C4277AA42B6657" pageId="0" pageNumber="25">
|
||
Radula (Fig.
|
||
<figureCitation id="DC403B1AD696285FEDB7BFBA87EC183C" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">25A-F</figureCitation>
|
||
): R-central tooth flanked by A-central, B-central, C-central, E-lateral on each side. Additionally, two interconnected layers of marginal teeth encase central and lateral teeth. R-central shows some variation among populations, slightly spherical or more squared face with slightly concave anterior edge. A-central large and flat with thin ridge that becomes broad and folded right at end of cusp. B-central diminished, forms irregular
|
||
<normalizedToken id="5EF0AA598A920A824DE0E77D9B1FF46D" originalValue="“S”">"S"</normalizedToken>
|
||
shape. C-central equally diminished, hidden below lower edge of E-lateral. E-lateral simple with smooth upper edge. First layer of marginal teeth consists of 37-44 teeth that decrease in size away from E-lateral but increase in size and bear serrations on edges of small faces; semidetached from second layer, which is fused and forms outer wall.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8FADA018042E247F81FCAFD40126D92" pageId="0" pageNumber="25" type="differentiating features">
|
||
<paragraph id="D3BA39A255BCA3EDB20CB0206A5741EB" pageId="0" pageNumber="25">Differentiating features.</paragraph>
|
||
<paragraph id="B7B9A543D758EC19CF62DF90F1AAC056" pageId="0" pageNumber="25">
|
||
Based on conchological features of periostracum colouration and patterning and shell shape, it is difficult to differentiate
|
||
<taxonomicName id="B1EFC4FCFB7DAC81254DD1A40678AF39" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="6BBE7361EAEF455EF3407A0160E573C0" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from most Asian
|
||
<taxonomicName id="B4A7D6EB6E8694AA1104263525E7CFF6" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="6B4FB3B5436264AF9C21627E586D3361" italics="true" pageId="0" pageNumber="25">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
spp. given the variety in colour and patterns among the
|
||
<typeStatus id="30AF5854D660D612314BD28C009D6E14">type</typeStatus>
|
||
material (Fig.
|
||
<figureCitation id="0D8CE7B6DB9D4F4B4E46EAACED39ECAB" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24A-M</figureCitation>
|
||
). However, in some instances it can be distinguished from Anatolian morphotypes of
|
||
<taxonomicName id="C31D367F6D5B3681611AA220BB2F2DAF" lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis id="7B9876B3DC87ED261B987FC42BF7CA03" italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
, which typically displays ivory blotches on a brown background (Fig.
|
||
<figureCitation id="DEE44F01EA02FACA0608B38E5E1A7670" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Theodoxus baeticus (Lamarck, 1822). A - D. Specimen (T. callosus - morphoype) collected in Akyaka, Turkey (UGSB 19118); E - H. Specimen (T. callosus - morphotype) collected in Kato Tragana, Greece (UGSB 2488); I - L. Specimen (T. varius - morphotype) collected at the Blue Eye Nature Monument, Albania (UGSB 24170); M - P. Specimen (T. meridionalis - morphotype) collected at the Ketana Oasis, Gabes, Tunisia (UGSB 18111). All specimens, except A-H, were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure7" httpUri="https://binary.pensoft.net/fig/376456" pageId="0" pageNumber="25">7A-D</figureCitation>
|
||
);
|
||
<taxonomicName id="0F1AC6BFEDA9E2C4D87FA0423CF1773E" lsidName="T. altenai" pageId="0" pageNumber="25" rank="species" species="altenai">
|
||
<emphasis id="8FB4F71A7E9DC5EA03195ECCB4DE8585" italics="true" pageId="0" pageNumber="25">T. altenai</emphasis>
|
||
</taxonomicName>
|
||
with clear ivory checks on a dark brown-black background (Fig.
|
||
<figureCitation id="54EADB9D878FD7C38D0D9B01BD6A3047" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Theodoxus altenai Schuett, 1965. A - C. Holotype from Lake Kirkgoez, Doesemealti, Antalya, Turkey (RNL V. 56 / 1, NMNL). D - G. Topotype (UGSB 24167) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure4" httpUri="https://binary.pensoft.net/fig/376453" pageId="0" pageNumber="25">4A-G</figureCitation>
|
||
) and
|
||
<taxonomicName id="690EBBBF58B33167BE63CA7D33726FD7" lsidName="T. gloeri" pageId="0" pageNumber="25" rank="species" species="gloeri">
|
||
<emphasis id="B758F6E754B7B38C1C344432272C2801" italics="true" pageId="0" pageNumber="25">T. gloeri</emphasis>
|
||
</taxonomicName>
|
||
lacking shell pigmentation and bearing strong axial ribs on the shell (Fig.
|
||
<figureCitation id="F753D55C8060B46939ABAAC4572E1FF6" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Theodoxus gloeri Odabasi & Arslan, 2015. A - C. Paratype from Balikdami Wetland-Sakarya River, Eskisehir, Turkey (coll. Gloeer). Scale bar: 1 mm." figureDoi="10.3897/zse.96.48312.figure11" httpUri="https://binary.pensoft.net/fig/376460" pageId="0" pageNumber="25">11A-C</figureCitation>
|
||
). The light ivory-coloured operculum calcareous base makes this species distinct from
|
||
<taxonomicName id="B1BF981A322533C14726DECC4F856736" lsidName="T. gurur" pageId="0" pageNumber="25" rank="species" species="gurur">
|
||
<emphasis id="90BF49F579B64F967623994C1DC53297" italics="true" pageId="0" pageNumber="25">T. gurur</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (light to dark brown; Figs
|
||
<figureCitation id="C44605E616D92D09B9FA1912FD773C76" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Theodoxus gurur Sands & Gloeer sp. nov. A - D. Holotype from the Suyunbasi spring complex, Ayrancilar, Izmir, Turkey stored in NMNL (RMNH. MOL. 342197); E - G. Paratype from same locality and storage facility (RMNH. MOL. 342199). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure13" httpUri="https://binary.pensoft.net/fig/376462" pageId="0" pageNumber="25">13D</figureCitation>
|
||
,
|
||
<figureCitation id="8E6CB5AFF86C118639246171357D2A21" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24D, G</figureCitation>
|
||
) and
|
||
<taxonomicName id="E6767F4F63EDEED5653C355954CBC98F" lsidName="T. wilkei" pageId="0" pageNumber="25" rank="species" species="wilkei">
|
||
<emphasis id="88602C4E48CAE15923FD6EF84BFB3BB1" italics="true" pageId="0" pageNumber="25">T. wilkei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (bright orange; Figs
|
||
<figureCitation id="D06D6CA5460B6F364929BE25610E30E2" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24D, G</figureCitation>
|
||
,
|
||
<figureCitation id="68158FC054D0ECADF408F1EF809CA86B" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Theodoxus wilkei Sands & Gloeer sp. nov. A - D. Holotype (RMNH. MOL. 342208); E - G. Paratype (RMNH. MOL. 342210); H - J. Paratype (RMNH. MOL. 342211). All photographed material is from the Cifteler Spring, Cifteler, Eskisehir, Turkey and stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure27" httpUri="https://binary.pensoft.net/fig/376476" pageId="0" pageNumber="25">27D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="A98606DD5576074CDF71BBD64716340A" pageId="0" pageNumber="25">
|
||
More differentiating features occur in the operculum structure (Fig.
|
||
<figureCitation id="274BCED67427203CD2594AB709A8B63F" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24D, G</figureCitation>
|
||
). The presence of an attenuated apophysis distinguishes
|
||
<taxonomicName id="C747317761D1F464A921F8A0E866D95B" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="7277D7413B138A7A86E682E87451772D" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from
|
||
<taxonomicName id="5F8689FD456573A3E2BB488031136BE8" lsidName="T. altenai" pageId="0" pageNumber="25" rank="species" species="altenai">
|
||
<emphasis id="C3371E84191C1FD30F8E05FE44601B91" italics="true" pageId="0" pageNumber="25">T. altenai</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2B1D775560DBEF4666E4CE8CC091A04D" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="A15EE3D6A013C6DE3FB7A76CEEA7FA14" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, which have non-attenuated apophyses (Figs
|
||
<figureCitation id="5548E26A12953309870FABE7A189223B" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Theodoxus altenai Schuett, 1965. A - C. Holotype from Lake Kirkgoez, Doesemealti, Antalya, Turkey (RNL V. 56 / 1, NMNL). D - G. Topotype (UGSB 24167) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure4" httpUri="https://binary.pensoft.net/fig/376453" pageId="0" pageNumber="25">4</figureCitation>
|
||
,
|
||
<figureCitation id="27148A0F2E9BF574D49CB653D231A12A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">15</figureCitation>
|
||
,
|
||
<figureCitation id="B4AE8BDB06B44AD0A26B621BD2188278" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">17</figureCitation>
|
||
). A rib-pouch and rib-shield are either totally lacking or extremely diminished in
|
||
<taxonomicName id="31AE470D12F419652226073F077F008C" lsidName="T. altenai" pageId="0" pageNumber="25" rank="species" species="altenai">
|
||
<emphasis id="3FD1D9A47B7B28380070044A6E59DD9E" italics="true" pageId="0" pageNumber="25">T. altenai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7558B4EC29390B862544A3C8EB39CCFF" lsidName="T. anatolicus" pageId="0" pageNumber="25" rank="species" species="anatolicus">
|
||
<emphasis id="43B59DA668C4F41AD8B13C1C34AC12B9" italics="true" pageId="0" pageNumber="25">T. anatolicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="EDCF77607225C759583AA6C694C3BE7D" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="853F26AC9B3A575B1244FCDF5F63C11B" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="DB0D60B7F7337C00633331D35FA78359" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="299585DF79DBE964704B2210A526C798" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="B2C084FAC64A4841CA0CFF9CC31D4D17" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Theodoxus altenai Schuett, 1965. A - C. Holotype from Lake Kirkgoez, Doesemealti, Antalya, Turkey (RNL V. 56 / 1, NMNL). D - G. Topotype (UGSB 24167) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure4" httpUri="https://binary.pensoft.net/fig/376453" pageId="0" pageNumber="25">4</figureCitation>
|
||
,
|
||
<figureCitation id="9432D5FE2F9B83ACCC8821500A0A7EC9" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Theodoxus anatolicus (Recluz, 1841). A - D. Specimen collected at Isikli, Denizli, Turkey (UGSB 24168) incorporated into the phylogeny (Fig. 2); E - G. Lectotype of N. anatolica from Izmir, Turkey (MHNG-MOLL- 15028). H, I. Paralectotype of N. anatolica from Izmir, Turkey (MHNG-MOLL- 15028); J, K. Paralectotype of N. anatolica from Chios, Greece (MNHN-IM- 2000 - 32519); L, M. Syntype of N. belladonna from Izmir, Turkey (ZMZ 528908). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure5" httpUri="https://binary.pensoft.net/fig/376454" pageId="0" pageNumber="25">5</figureCitation>
|
||
,
|
||
<figureCitation id="0422E7C05DD78F340FDC85C04A86D098" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">15</figureCitation>
|
||
,
|
||
<figureCitation id="E96CA72E9A22478B79BE29A5AC350252" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">17</figureCitation>
|
||
,
|
||
<figureCitation id="E91A77DBD08F6E35D97B820DEACE95C1" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Theodoxus macri (G. B. Sowerby II, 1849). A - D. Specimen collected close to Harbiye Falls, Harbiye, Antakya, Turkey (UGSB 24179); E - H. Specimen from Lake Yenisehir, Reyhanli, Turkey (UGSB 24180); I, J. Specimen from Lake Balik, Adalar, Turkey (UGSB 24181). All specimens (A-J) were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure18" httpUri="https://binary.pensoft.net/fig/376467" pageId="0" pageNumber="25">18</figureCitation>
|
||
), while they are more pronounced in
|
||
<taxonomicName id="2B04A0FEFA485B1459223A94991637D8" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="6D3787CD2A521277479247C63CD64B63" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Fig.
|
||
<figureCitation id="5D99580EE1115A3CD479A658CCD50291" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24</figureCitation>
|
||
). The rib-shield in
|
||
<taxonomicName id="8E7C3B737F69663B98383E88965997D9" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="17D62FA0C078434F6B0E9BF312420469" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is however less broad than that typically observed in
|
||
<taxonomicName id="9CC4390E1F91CCE123B4A2E3B9961D1A" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="63A3F9DC3CF1040C8AA2B9EF324E2A55" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="2996FB3F4144ADDE27BF70A7BC43659E" lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis id="BAB8EBC7EEA596AA2C7452272754D09C" italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="8A93F367133A75D24FBBC26FDFA87CE6" captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Theodoxus baeticus (Lamarck, 1822). A - C. Paralectotype of N. baetica from Andalusia, Spain (MHNG-MOLL- 51319). D) Operculum of a paralectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); E - G. Lectotype of N. baetica from Andalusia (MHNG-MOLL- 51319); H-J; K; L; M; N. Five syntypes of N. meridionalis from Sicily (ZMB, without coll. no.); O - R. Topotype of N. guadianensis collected in the Guadiana River near Mertola, Portugal incorporated into the phylogeny (UGSB 22084); S - V. Specimen from Soller, Balearic Islands used in the phylogeny (UGSB 19162). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure6" httpUri="https://binary.pensoft.net/fig/376455" pageId="0" pageNumber="25">6</figureCitation>
|
||
-
|
||
<figureCitation id="BEF9C2DA8C7A95BDE862E3FA57A214D6" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">10</figureCitation>
|
||
). Furthermore, the lack of a pseudo-apophysis differentiates the new species from
|
||
<taxonomicName id="856636764B4D858DC86E22284406A3C9" lsidName="T. altenai" pageId="0" pageNumber="25" rank="species" species="altenai">
|
||
<emphasis id="B5978636DFEB217B134DE70E720710B1" italics="true" pageId="0" pageNumber="25">T. altenai</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5976955648B76B4DC30DFBD9FB264619" lsidName="T. anatolicus" pageId="0" pageNumber="25" rank="species" species="anatolicus">
|
||
<emphasis id="2E92356D69A8742424F970680A5D4A1F" italics="true" pageId="0" pageNumber="25">T. anatolicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="F109D43074A873D6F193C3FFF33F6BEC" lsidName="T. baeticus" pageId="0" pageNumber="25" rank="species" species="baeticus">
|
||
<emphasis id="2B9CAA16D2434CA9C77F9F0551C6A8BF" italics="true" pageId="0" pageNumber="25">T. baeticus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4BD4144FBC50112C7CD9F17208071BED" lsidName="T. gurur" pageId="0" pageNumber="25" rank="species" species="gurur">
|
||
<emphasis id="F3AA186D5C3B68C7CEF442274891AF20" italics="true" pageId="0" pageNumber="25">T. gurur</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="66A52AA153B145E83D1880F14E23CB70" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="960465A63F4655DF523C3C232560FD22" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="2B149A81F99F660FB2BC928E555112EA" lsidName="T. macri" pageId="0" pageNumber="25" rank="species" species="macri">
|
||
<emphasis id="9DBBFB78B6189891E3A7C48D3D09172E" italics="true" pageId="0" pageNumber="25">T. macri</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation id="661428A8151BA9547A7A076383795275" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Theodoxus altenai Schuett, 1965. A - C. Holotype from Lake Kirkgoez, Doesemealti, Antalya, Turkey (RNL V. 56 / 1, NMNL). D - G. Topotype (UGSB 24167) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure4" httpUri="https://binary.pensoft.net/fig/376453" pageId="0" pageNumber="25">4</figureCitation>
|
||
-
|
||
<figureCitation id="2037BD8464C24CEDD03604122384276A" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Theodoxus baeticus (Lamarck, 1822). A - D. Specimen from Cuquillo, Granada, Andalusia, Spain (UGSB 22089); E - H. Specimen (T. meridionalis - morphotype) collected in the Palancia River, Navajas, Spain (UGSB 22090); I - L. Specimen (T. elongatulus - morphotype) from Arrancada, Pombal, Portugal (UGSB 22080); M - P. Specimen (T. velascoi - morphotype) from Montanejos, Castellon, Spain (UGSB 22082). Theodoxus cf. valentinus (Graells, 1846). Q - S. Specimen conforming to T. valentinus from the Verd River, Masalaves, Spain (UGSB 21787); T. Operculum of a specimen conforming to T. valentinus from the same locality as Q-S (UGSB 22087). All specimens, except Q-S, were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure8" httpUri="https://binary.pensoft.net/fig/376457" pageId="0" pageNumber="25">8</figureCitation>
|
||
,
|
||
<figureCitation id="DFDEFAE3640DF5561EC5487F59C0D64F" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Theodoxus gurur Sands & Gloeer sp. nov. A - D. Holotype from the Suyunbasi spring complex, Ayrancilar, Izmir, Turkey stored in NMNL (RMNH. MOL. 342197); E - G. Paratype from same locality and storage facility (RMNH. MOL. 342199). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure13" httpUri="https://binary.pensoft.net/fig/376462" pageId="0" pageNumber="25">13</figureCitation>
|
||
,
|
||
<figureCitation id="266829841584085CDBF4099F0C06CC1A" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">15</figureCitation>
|
||
,
|
||
<figureCitation id="90038E916A3D46138D86A13125936628" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">17</figureCitation>
|
||
,
|
||
<figureCitation id="5C51983D599A87EF14CE886F69069917" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Theodoxus macri (G. B. Sowerby II, 1849). A - D. Specimen collected close to Harbiye Falls, Harbiye, Antakya, Turkey (UGSB 24179); E - H. Specimen from Lake Yenisehir, Reyhanli, Turkey (UGSB 24180); I, J. Specimen from Lake Balik, Adalar, Turkey (UGSB 24181). All specimens (A-J) were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure18" httpUri="https://binary.pensoft.net/fig/376467" pageId="0" pageNumber="25">18</figureCitation>
|
||
). Additionally, the presence of a weak callus on the top right edge of the operculum base in
|
||
<taxonomicName id="5B0B0B6351B3C284F9FE77A769B12554" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="BDD32C0051B3D7982B8A4C64AD6DB53A" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. helps to differentiate this species from
|
||
<taxonomicName id="33AFDA31E2D2C21AA0A719E89354A9B6" lsidName="T. gurur" pageId="0" pageNumber="25" rank="species" species="gurur">
|
||
<emphasis id="10057CD62F419118C3199924CB906B51" italics="true" pageId="0" pageNumber="25">T. gurur</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="5387A74B784E5C25D33751EF203C5A43" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="0AF340C6CB5FD7307BED5D4437C7E1F2" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, which lack a callus (Figs
|
||
<figureCitation id="1FB4FB884C3737A482E94ECCC61D4C23" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Theodoxus gurur Sands & Gloeer sp. nov. A - D. Holotype from the Suyunbasi spring complex, Ayrancilar, Izmir, Turkey stored in NMNL (RMNH. MOL. 342197); E - G. Paratype from same locality and storage facility (RMNH. MOL. 342199). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure13" httpUri="https://binary.pensoft.net/fig/376462" pageId="0" pageNumber="25">13</figureCitation>
|
||
,
|
||
<figureCitation id="32C5E23187566F28958C38BD4168A520" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Theodoxus jordani (G. B. Sowerby I, 1836). A - D. Specimen from the Avakas Gorge, Peyia, Cyprus (UGSB 18321) incorporated into the phylogeny (Fig. 2); E - H. Specimen collected in the Ceyhan River, Eski Misis, Adana, Turkey (UGSB 20777); I - L. Specimen from the Orontes River, Antakya, Hatay, Turkey (UGSB 24175) used in the phylogeny (Fig. 2); M - O. Specimen (T. euphraticus - morphotype) from the Seyed Hosein Park spring, near the Bishapur, Fars, Iran (UGSB 22233); P - R. Topotype of N. jordani from the River Jordan, Israel (UGSB 24177) incorporated into the phylogeny (Fig. 2); S, T. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21681); U. Specimen (T. niloticus - morphotype) collected in the Nile River, Egypt (UGSB 24178) used in the phylogeny (Fig. 2); V. Specimen (T. euphraticus - morphotype) from the Karun River, Ahvaz, Iran (UGSB 21682). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure15" httpUri="https://binary.pensoft.net/fig/376464" pageId="0" pageNumber="25">15</figureCitation>
|
||
,
|
||
<figureCitation id="A5A27D62B4EE4364501ABA7B6BD1C76B" captionStart="Figure 17" captionStartId="F17" captionText="Figure 17. Theodoxus jordani (G. B. Sowerby I, 1836). A - C. Syntype of N. meridionalis var. mesopotamica from near Diyarbakir, Turkey (ZMZ 528914); D, E. Specimen (T. mesopotamicus - morphotype) collected in Sanliurfa, Turkey (UGSB 23426) and used in the phylogeny (Fig. 2); F - H; K - M. Two syntypes of N. cinctella collected at the source of River Chabur near Ras al-Ayn, Syria (ZMB / Moll 21732); I, J. Topotype of N. cinctella from the same locality (UGSB 18762). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure17" httpUri="https://binary.pensoft.net/fig/376466" pageId="0" pageNumber="25">17</figureCitation>
|
||
,
|
||
<figureCitation id="96083D3E10A9E7EF5353983C53A5093D" captionStart="Figure 18" captionStartId="F18" captionText="Figure 18. Theodoxus macri (G. B. Sowerby II, 1849). A - D. Specimen collected close to Harbiye Falls, Harbiye, Antakya, Turkey (UGSB 24179); E - H. Specimen from Lake Yenisehir, Reyhanli, Turkey (UGSB 24180); I, J. Specimen from Lake Balik, Adalar, Turkey (UGSB 24181). All specimens (A-J) were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure18" httpUri="https://binary.pensoft.net/fig/376467" pageId="0" pageNumber="25">18</figureCitation>
|
||
,
|
||
<figureCitation id="35EA793D67BC58C1426CF9F4D5710E4C" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24</figureCitation>
|
||
), as well as from
|
||
<taxonomicName id="071317A022AB6170985850FDA55622C0" lsidName="T. anatolicus" pageId="0" pageNumber="25" rank="species" species="anatolicus">
|
||
<emphasis id="E4C9A7D38956CC7CA36E58B92626CA89" italics="true" pageId="0" pageNumber="25">T. anatolicus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="35A9D54C71CCE0FC89FF14FEEBA9A63F" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="DF577ED6A37B56A938DBA49159B946D3" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="6677D8FA2FE536E914A2CFA40330BD6B" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="42C500E5373654C06B622B2C99F9C160" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="63C8D826E6189E595FEDAFF73C57F395" lsidName="T. pallidus" pageId="0" pageNumber="25" rank="species" species="pallidus">
|
||
<emphasis id="232FF0F7DBBE7AB7946B0890A6F7009A" italics="true" pageId="0" pageNumber="25">T. pallidus</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="AB937DBB09D7E74B1F3B0C0C6E76D219" lsidName="T. wilkei" pageId="0" pageNumber="25" rank="species" species="wilkei">
|
||
<emphasis id="52AB3E97409126C14C9313CC8B9D62EF" italics="true" pageId="0" pageNumber="25">T. wilkei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., which have stronger calluses (Figs
|
||
<figureCitation id="2C75763478F753A877F4C9E3DF4F5CCA" captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Theodoxus anatolicus (Recluz, 1841). A - D. Specimen collected at Isikli, Denizli, Turkey (UGSB 24168) incorporated into the phylogeny (Fig. 2); E - G. Lectotype of N. anatolica from Izmir, Turkey (MHNG-MOLL- 15028). H, I. Paralectotype of N. anatolica from Izmir, Turkey (MHNG-MOLL- 15028); J, K. Paralectotype of N. anatolica from Chios, Greece (MNHN-IM- 2000 - 32519); L, M. Syntype of N. belladonna from Izmir, Turkey (ZMZ 528908). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure5" httpUri="https://binary.pensoft.net/fig/376454" pageId="0" pageNumber="25">5</figureCitation>
|
||
,
|
||
<figureCitation id="AEF0D99F76A69A331621FF2D0BB16396" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen collected close to Oued Laabid, Morocco (UGSB 18106) (Fig. 2); E - H. Specimen from the Danube River, Woerth an der Donau, Germany (UGSB 24173); I - L. Specimen (T. euxinus - morphotype) collected in the Papuc Creek, Kiyikoey, Turkey (UGSB 24172) (Fig. 2); M - P. Specimen (T. euxinus - morphotype) from Ovidiopol, Ukraine (UGSB 18124); Q - S. Specimen (T. heldreichi - morphotype) collected in Lake Egirdir, Turkey (UGSB 20361); T, U. Specimen (T. danasteri - morphotype) collected in Krasna Kosa, Ukraine (UGSB 18417) (Fig. 2); V. Opercula of a T. danasteri - morphotype from Lake Skadar, Montenegro (UGSB 24171). Specimens A-D, I-L and T-U were used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure9" httpUri="https://binary.pensoft.net/fig/376458" pageId="0" pageNumber="25">9</figureCitation>
|
||
,
|
||
<figureCitation id="ED8785113324016943F7C7D51FA9EC9F" captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Theodoxus fluviatilis (Linnaeus, 1758). A - D. Specimen (conforming to T. sarmaticus) from the Dnieper-Bug estuary, Ukraine (UGSB 18328); E - H. Specimen (T. subthermalis - morphotype) from Novy Afon, Abkhazia, Georgia (UGSB 18402); I - K. Syntype of N. saulcyi from near the Penteli Monastery, Greece (MHNG-MOLL- 111736); L. Specimen (T. saulcyi - morphotype) from Stilos, Crete, Greece (UGSB 20519) incorporated into the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure10" httpUri="https://binary.pensoft.net/fig/376459" pageId="0" pageNumber="25">10</figureCitation>
|
||
,
|
||
<figureCitation id="F62550F66CE46037CDBC13EEF0107A70" captionStart="Figure 19" captionStartId="F19" captionText="Figure 19. Theodoxus major Issel, 1865. A, B. Specimen (T. pallasi - morphotype) collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20712); C, D. Specimen (T. pallasi - morphotype) collected in the Aral Sea, Kazakhstan in 1979 (UGSB 19129); E - H. Specimen (T. pallasi - morphotype) from the Shahpol River, Aliabad Askar Khan, Iran (UGSB 18091); I - K. Specimen of T. major collected at the outflow of Lake Akna, Aknalich, Armenia (UGSB 20482); L - N. Specimen (T. astrachanicus - morphotype) collected in Utlyukskij Liman, Ukraine (UGSB 18130); O, P. Specimen of T. major collected in the Hrazdan River at the inflow of the Yerevan Reservoir, Armenia (UGSB 20496); Q - T. Specimen of T. major from Zoeram, North Khorasan, Iran (UGSB 21661); U, V. Specimen of T. schultzii collected in the Caspian Sea, offshore from Aktau, Kazakhstan (UGSB 20714); W - Y. Lectotype of T. pallasi from Caspian Sea, Dagestan, Russia (ZIN 54547 / 63); Za, Zb. Syntype of N. schultzii from the Caspian Sea (ZIN, no. 5 in systematic catalogue). Specimens E-K and O-T used in the phylogeny (Fig. 2). Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure19" httpUri="https://binary.pensoft.net/fig/376468" pageId="0" pageNumber="25">19</figureCitation>
|
||
,
|
||
<figureCitation id="E1281E633081C029E380F5B680917B8B" captionStart="Figure 20" captionStartId="F20" captionText="Figure 20. Theodoxus pallidus (Dunker, 1861). A - D. Specimen from Haji Abad, Fars, Iran (UGSB 22227); E - H. Syntype of N. pallida from southern Iran (ZMB 108.792); I, J; K, L. Two syntypes of N. pallida from southern Iran (ZMB 108.793); M - P. Specimen (T. doriae - morphotype) from a spring near Harat, Yazd, Iran (UGSB 21706); Q, R; S, T; U, V; W - Y. Four syntypes of N. doriae from the thermal springs of Kerman, Iran (MSNG, without coll. no.). Specimens A-D and M-P were used in the phylogeny. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure20" httpUri="https://binary.pensoft.net/fig/376469" pageId="0" pageNumber="25">20</figureCitation>
|
||
,
|
||
<figureCitation id="B2845D3FF0DDF0954E0A6506CDDADC28" captionStart="Figure 24" captionStartId="F24" captionText="Figure 24. Theodoxus wesselinghi Sands & Gloeer sp. nov. A - D. Holotype collected in the Sakarya River, Caykoey, Bilecik, Turkey (RMNH. MOL. 342200); E - G. Paratype from the same location as the holotype (RMNH. MOL. 342202); H - J. Paratype from Fele, Isparta province, Turkey (RMNH. MOL. 342204); K - M. Paratype from Eflatun Pinari, Konya province, Turkey (RMNH. MOL. 342206). All photographed material (A-M) is stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure24" httpUri="https://binary.pensoft.net/fig/376473" pageId="0" pageNumber="25">24</figureCitation>
|
||
,
|
||
<figureCitation id="760AE766B1C2D5A669C838BCC40D7D91" captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Theodoxus wilkei Sands & Gloeer sp. nov. A - D. Holotype (RMNH. MOL. 342208); E - G. Paratype (RMNH. MOL. 342210); H - J. Paratype (RMNH. MOL. 342211). All photographed material is from the Cifteler Spring, Cifteler, Eskisehir, Turkey and stored in NMNL. Scale bars: 1 mm." figureDoi="10.3897/zse.96.48312.figure27" httpUri="https://binary.pensoft.net/fig/376476" pageId="0" pageNumber="25">27</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="E88EE596BB0A7E0362934D90F5913189" pageId="0" pageNumber="25">
|
||
Based on the available data for
|
||
<taxonomicName id="CFF9E516316922A17CBFB1CA7C51E6A3" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="C3EB82334890CB4BF34E94EE48A56D86" italics="true" pageId="0" pageNumber="25">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
ralulae,
|
||
<taxonomicName id="18A5686C5ED385BBE1EA79E438308EF7" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="F88752D90457200DA2B502E810552739" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be distinguished by a more globular R-central face from
|
||
<taxonomicName id="E33037E2BCCB782610AF2A8F9CDA7BFF" lsidName="T. gurur" pageId="0" pageNumber="25" rank="species" species="gurur">
|
||
<emphasis id="6D224B22FD5696F4D5014CB9A0CE5124" italics="true" pageId="0" pageNumber="25">T. gurur</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="378808BB304D9E92E9117C6AAFEF27A1" lsidName="T. wilkei" pageId="0" pageNumber="25" rank="species" species="wilkei">
|
||
<emphasis id="1E6E449E4BECCFE14B886799F9121694" italics="true" pageId="0" pageNumber="25">T. wilkei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov.,
|
||
<taxonomicName id="8B0022B74D0218BBCEF26D612E06F7B4" lsidName="T. fluviatilis" pageId="0" pageNumber="25" rank="species" species="fluviatilis">
|
||
<emphasis id="DA70E049629AFBA4E59B419B6A400F38" italics="true" pageId="0" pageNumber="25">T. fluviatilis</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="8673688096EB0F66A3F5B9B8679D65E6" lsidName="T. jordani" pageId="0" pageNumber="25" rank="species" species="jordani">
|
||
<emphasis id="2B0C9DFD24C3CAC902E024FAAAE0A6DA" italics="true" pageId="0" pageNumber="25">T. jordani</emphasis>
|
||
</taxonomicName>
|
||
, where it is more rectangular or triangulate (see
|
||
<bibRefCitation id="FCFD5D7BD3E7BEB86760FFC21642261A" DOI="https://doi.org/10.2307/4063879" author="Baker, HB" journalOrPublisher="Proceedings of the Academy of Natural Sciences of Philadelphia" pageId="0" pageNumber="25" pagination="117 - 178" refId="B18" refString="Baker, HB, 1923. Notes on the radula of the Neritidae. Proceedings of the Academy of Natural Sciences of Philadelphia 75: 117 - 178, DOI: https://doi.org/10.2307/4063879" title="Notes on the radula of the Neritidae." url="https://doi.org/10.2307/4063879" volume="75" year="1923">Baker 1923</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="D9732FEE15E34EC35BCEAEF94ABEA5B0" author="Zettler, ML" journalOrPublisher="Mollusca" pageId="0" pageNumber="25" pagination="13 - 72" refId="B166" refString="Zettler, ML, 2008. Zur Taxonomie und Verbreitung der Gattung Theodoxus Montfort, 1810 in Deutschland. Darstellung historischer und rezenter Daten einschliesslich einer Bibliografie. Mollusca 26 (1): 13 - 72" title="Zur Taxonomie und Verbreitung der Gattung Theodoxus Montfort, 1810 in Deutschland. Darstellung historischer und rezenter Daten einschliesslich einer Bibliografie." volume="26" year="2008">Zettler 2008</bibRefCitation>
|
||
; Figs
|
||
<figureCitation id="0E5E329AA078BF49D65B8EBA2873B528" captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Radula of Theodoxus gurur Sands & Gloeer sp. nov. paratypes (UGSB 20689). A. Portion of the radula showing full sets of teeth; B. Magnified view of the central and lateral teeth; C. Magnified view of the lateral and marginal teeth; D. Magnified view of the first row of marginal teeth; E. Merging point between the first and second rows of marginal teeth; F. Magnified view of the faces of inner marginal teeth belonging to the first row. Scale bars: 100 μm (A-D), 20 μm (E-F)." figureDoi="10.3897/zse.96.48312.figure14" httpUri="https://binary.pensoft.net/fig/376463" pageId="0" pageNumber="25">14B</figureCitation>
|
||
,
|
||
<figureCitation id="2FC11CB7BED28752B58B605FCDB5619A" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">25B</figureCitation>
|
||
,
|
||
<figureCitation id="A7C62C71C5EC0FCA909DC87B44E7A029" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Radula of Theodoxus wilkei Sands & Gloeer sp. nov. paratypes (UGSB 20687). A. Portion of the radula showing full sets of teeth; B. Magnified view of the central and lateral teeth; C. Magnified view of the lateral and first and second rows of marginal teeth; D. Magnified view of the lateral teeth; E. The first row of marginal teeth; F. Magnified view of the faces of inner marginal teeth belonging to the first row. Scale bars: 100 μm (A-D), 20 μm (E-F)." figureDoi="10.3897/zse.96.48312.figure28" httpUri="https://binary.pensoft.net/fig/376477" pageId="0" pageNumber="25">28B</figureCitation>
|
||
). Furthermore, the smooth upper edge of the E-lateral can be used to distinguish this species from
|
||
<taxonomicName id="51AD5F74ABAFB630C8F0178D00797FDA" lsidName="T. wilkei" pageId="0" pageNumber="25" rank="species" species="wilkei">
|
||
<emphasis id="8E820755D21F15C32197BBDAE5A2CF51" italics="true" pageId="0" pageNumber="25">T. wilkei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="A7E5BD23C6AE0C51DCF2D213E413CA08" lsidName="T. major" pageId="0" pageNumber="25" rank="species" species="major">
|
||
<emphasis id="603DE7CBB9B29A728E1D3D29E2510FCA" italics="true" pageId="0" pageNumber="25">T. major</emphasis>
|
||
</taxonomicName>
|
||
, which generally have E-laterals with serrated edges (see
|
||
<bibRefCitation id="8F9054776F0FC3526F3807B57B435DD7" DOI="https://doi.org/10.1127/arch.moll/146/213-226" author="Anistratenko, VV" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="25" pagination="213 - 226" refId="B17" refString="Anistratenko, VV, Zettler, ML, Anistratenko, OY, 2017. On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region. Archiv fuer Molluskenkunde 146 (2): 213 - 226, DOI: https://doi.org/10.1127/arch.moll/146/213-226" title="On the taxonomic relationship between Theodoxus pallasi and T. astrachanicus (Gastropoda: Neritidae) from the Ponto-Caspian region." url="https://doi.org/10.1127/arch.moll/146/213-226" volume="146" year="2017">Anistratenko et al. 2017</bibRefCitation>
|
||
; Figs
|
||
<figureCitation id="ADCC79EC21C0DAE6F44D7FC07A6DD729" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Radula of Theodoxus wesselinghi Sands & Gloeer sp. nov. paratypes. A. Portion of the radula showing full sets of teeth (UGSB 20688); B. Magnified view of the central teeth (UGSB 20688); C. Magnified view of the lateral and marginal teeth (UGSB 20686); D. Magnified view of the first and second rows of marginal teeth (UGSB 20688); E. Magnified view of the first row of marginal teeth (UGSB 20686); F. Magnified view of the faces of inner marginal teeth belonging to the first row (UGSB 20685). Scale bars: 100 μm (A-E), 20 μm (F)." figureDoi="10.3897/zse.96.48312.figure25" httpUri="https://binary.pensoft.net/fig/376474" pageId="0" pageNumber="25">25C</figureCitation>
|
||
,
|
||
<figureCitation id="9CF839D7A831007FF723B567D1EB7B94" captionStart="Figure 28" captionStartId="F28" captionText="Figure 28. Radula of Theodoxus wilkei Sands & Gloeer sp. nov. paratypes (UGSB 20687). A. Portion of the radula showing full sets of teeth; B. Magnified view of the central and lateral teeth; C. Magnified view of the lateral and first and second rows of marginal teeth; D. Magnified view of the lateral teeth; E. The first row of marginal teeth; F. Magnified view of the faces of inner marginal teeth belonging to the first row. Scale bars: 100 μm (A-D), 20 μm (E-F)." figureDoi="10.3897/zse.96.48312.figure28" httpUri="https://binary.pensoft.net/fig/376477" pageId="0" pageNumber="25">28C, D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="ACCE82E7A96349B8C45444B42B1319D0" pageId="0" pageNumber="25" type="remarks">
|
||
<paragraph id="BAEDE6D8808B95B4A0172F3B690733E9" pageId="0" pageNumber="25">Remarks.</paragraph>
|
||
<paragraph id="FA9F5984C65E84D8260367349219D8B0" pageId="0" pageNumber="25">
|
||
<taxonomicName id="CBB4B2BB7C999B67BE672B5170ACAAB8" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi">
|
||
<emphasis id="16F1122D9DB688596778F941CBD38630" italics="true" pageId="0" pageNumber="25">Theodoxus wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. forms part of a larger clade that includes
|
||
<taxonomicName id="56C020E252FA4FC6563800F835119346" lsidName="T. syriacus" pageId="0" pageNumber="25" rank="species" species="syriacus">
|
||
<emphasis id="97A01DFABF70C0C882EE5965D2DCE30D" italics="true" pageId="0" pageNumber="25">T. syriacus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="04FAAD7FE8246104148F890477D93E50" lsidName="T. wilkei" pageId="0" pageNumber="25" rank="species" species="wilkei">
|
||
<emphasis id="0644CEAF504F69BE130D7C6C8F7BAB55" italics="true" pageId="0" pageNumber="25">T. wilkei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., where it shares a closer sister-species relationship with
|
||
<taxonomicName id="CBCF74E903E916FAEB2C7BF35C796D50" lsidName="T. wilkei" pageId="0" pageNumber="25" rank="species" species="wilkei">
|
||
<emphasis id="78AB2976D44CAD9861B6586E48CB0633" italics="true" pageId="0" pageNumber="25">T. wilkei</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (
|
||
<bibRefCitation id="3EE7AE01D06AADF60DF57C6A62B8694A" DOI="https://doi.org/10.1111/jbi.13590" author="Sands, AF" journalOrPublisher="Journal of Biogeography" pageId="0" pageNumber="25" pagination="1274 - 1286" refId="B138" refString="Sands, AF, Sereda, SV, Stelbrink, B, Neubauer, TA, Lazarev, S, Wilke, T, Albrecht, C, 2019a. Contributions of biogeographical functions to species accumulation may change over time in refugial regions. Journal of Biogeography 46 (6): 1274 - 1286, DOI: https://doi.org/10.1111/jbi.13590" title="Contributions of biogeographical functions to species accumulation may change over time in refugial regions." url="https://doi.org/10.1111/jbi.13590" volume="46" year="2019 a">Sands et al. 2019a</bibRefCitation>
|
||
; Fig.
|
||
<figureCitation id="EF0305B7788C954011D4D40F8AA24E21" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
). The three species likely diverged from one another in quick succession over the Pliocene-Pleistocene transition (Fig.
|
||
<figureCitation id="4F838739A6CF917449A03FB9F278BE63" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Dated phylogeny of the genus Theodoxus constructed with BEAST, based on existing GenBank and newly incorporated data for COI, 16 S and ATPα (Suppl. material 1: Table S 1). Morphospecies where associations are queried are highlighted by a question mark after the species name (see the " Systematic checklist " for more information). Small green (PP ≥ 0.95) and red (PP <0.95) dots on nodes indicate the relative posterior probability support of divergence events. Node labels (1 - 32) correspond to divergence dates and HPD intervals in millions of years ago (Mya) for selected nodes (see Suppl. material 1: Table S 3). Dashed dark blue and light blue lines represent the minimum interspecific and maximum intraspecific variation determined by Sands et al. (2019 a) to aid Theodoxus spp. delimitation (see see Methods, Phylogenetics and species delimitation and that study for more information on the delimitation methods)." figureDoi="10.3897/zse.96.48312.figure2" httpUri="https://binary.pensoft.net/fig/376451" pageId="0" pageNumber="25">2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="0078B8EC70BCA45E2D35CC7A92B48040" pageId="0" pageNumber="25" type="distribution">
|
||
<paragraph id="AFFE6A4DB3B88599B1D456B395199E33" pageId="0" pageNumber="25">Distribution.</paragraph>
|
||
<paragraph id="F1A463CB6A087E4FD467B71467A2F5B9" pageId="0" pageNumber="25">
|
||
Known so far only from the four localities in central-west Anatolia (Figs
|
||
<figureCitation id="BD2FCEFBD52C06477428EA35A6BA1463" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Occurrence map for Theodoxus spp. in Asia. All points conform to specimens used in the phylogenetic analyses (Suppl. material 1: Table S 1), except for T. gloeri for which no molecular data are available and only the type and paratype localities are indicated. Species are partitioned across four identical maps (A - D) to limit overlap of points as far as possible. Barring T. gloeri, closer related species are grouped together (Fig. 2)." figureDoi="10.3897/zse.96.48312.figure3" httpUri="https://binary.pensoft.net/fig/376452" pageId="0" pageNumber="25">3D</figureCitation>
|
||
,
|
||
<figureCitation id="C6DDA26607FCE7886101A8984F58E41B" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23A-H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="379FBB7569533F38D92CF8585BD90CF2" pageId="0" pageNumber="25" type="ecology">
|
||
<paragraph id="E7FB58595E4A2B96D1F07CE1A8CBBFE3" pageId="0" pageNumber="25">Ecology.</paragraph>
|
||
<paragraph id="248752DD694258134F8BAC5BC0F6D34B" pageId="0" pageNumber="25">
|
||
<taxonomicName id="24CF3B76136B4160C976D58F2014CB75" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi">
|
||
<emphasis id="2F34D485B142E203C0959DBAFA62B704" italics="true" pageId="0" pageNumber="25">Theodoxus wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. can be found in both springs (
|
||
<normalizedToken id="57CB011D3581609C5FF9B65990CE241C" originalValue="Balıkdamı">Balikdami</normalizedToken>
|
||
Wetland, Eflatun
|
||
<normalizedToken id="DD25DCB79A1E3E1BBD06F2429BAA7A73" originalValue="Pınarı">Pinari</normalizedToken>
|
||
, Fele; Fig.
|
||
<figureCitation id="B8747600C69C8DD1582EE1DB514CD670" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23C-H</figureCitation>
|
||
) and streams (Sakarya River,
|
||
<normalizedToken id="E57F8F8236CA0636D8F0EAFA4AFE0CE0" originalValue="Çayköy">Caykoey</normalizedToken>
|
||
; Fig.
|
||
<figureCitation id="B9A05BF96F3FAF479A9E03209A3AF20C" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23A, B</figureCitation>
|
||
) with clear water. The occurrence of macrophytes appears negligible to the species as it occurs in localities both with (Fig.
|
||
<figureCitation id="69C2FF9A31422EF32B56FDDB9A2C0E96" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23E-H</figureCitation>
|
||
) and without (Fig.
|
||
<figureCitation id="9195379965A84566E43D5DD074EC5749" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23A-D</figureCitation>
|
||
) aquatic plant growth. The floors of all localities are made up of coarse grained sand and large and small rocks and stones that
|
||
<taxonomicName id="FBDF2E2162253EAD20D770CCED33B4E1" lsidName="T. wesselinghi" pageId="0" pageNumber="25" rank="species" species="wesselinghi">
|
||
<emphasis id="500F73D92414BC0EDB44549E5CE8C7DB" italics="true" pageId="0" pageNumber="25">T. wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is often attached to (personal observation M.E.G.; Fig.
|
||
<figureCitation id="E75C8EEBE7710684DF179B772B3EE9FE" captionStart="Figure 23" captionStartId="F23" captionText="Figure 23. Type locality of Theodoxus wesselinghi Sands & Gloeer sp. nov. A, B. Sakarya River, Caykoey, Bilecik, Turkey, 40.0439 ° N, 30.452 ° E. Further paratype localities: C, D. Unnamed roadside spring in Fele, Isparta province, Turkey, 38.00358 ° N, 31.47217 ° E; E, F. Eflatun Pinari, near Sadikhaci, Konya, Turkey, 37.8256 ° N, 31.6748 ° E; G, H. Balikdami Wetland spring, Eskisehir, Turkey, 39.15277 ° N, 31.61562 ° E." figureDoi="10.3897/zse.96.48312.figure23" httpUri="https://binary.pensoft.net/fig/376472" pageId="0" pageNumber="25">23H</figureCitation>
|
||
).
|
||
<taxonomicName id="782D842DA906B4F6C1702092EAF131BC" authorityName="Sands & Gloeer" authorityYear="2020" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theodoxus wesselinghi" order="Cycloneritida" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="wesselinghi">
|
||
<emphasis id="C6C8D74F2F8802AF2768A0B8FA7FC275" italics="true" pageId="0" pageNumber="25">Theodoxus wesselinghi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. co-occurs with
|
||
<taxonomicName id="629DF674490525B1734D6CF685989B78" baseAuthorityName="Gray" baseAuthorityYear="1843" class="Gastropoda" family="Tateidae" genus="Potamopyrgus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamopyrgus antipodarum" order="Littorinimorpha" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="antipodarum">
|
||
<emphasis id="9DB832F4D7EBABA67C845BDA6F58671A" italics="true" pageId="0" pageNumber="25">Potamopyrgus antipodarum</emphasis>
|
||
</taxonomicName>
|
||
(Gray, 1843) and
|
||
<taxonomicName id="4F454E87951A113056E3830F27D5C49E" class="Gastropoda" family="Hydrobiidae" genus="Isparta" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Isparta felei" order="Littorinimorpha" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="felei">
|
||
<emphasis id="BFE1215220EDBC024E17E3600AD5F569" italics="true" pageId="0" pageNumber="25">Isparta felei</emphasis>
|
||
</taxonomicName>
|
||
<normalizedToken id="B4B306C75DDAB8369E65DD5C47325F47" originalValue="Yıldırım">Yildirim</normalizedToken>
|
||
, Koca,
|
||
<normalizedToken id="FA581BE17FF9EB6633D40A6AD3DD69A8" originalValue="Gürlek">Guerlek</normalizedToken>
|
||
&
|
||
<normalizedToken id="FB03FB401B5EE5CA2C507E7070965D94" originalValue="Glöer">Gloeer</normalizedToken>
|
||
,
|
||
<quantity id="0CA33378D7992E3F9DA38EE20E753550" metricMagnitude="1" metricUnit="m" metricValue="5.125719999999999" unit="in" value="2018.0">2018 in</quantity>
|
||
Fele spring,
|
||
<taxonomicName id="1584C0007C2CD87B4B68582CEDBA951D" authorityName="Radoman" authorityYear="1973" class="Gastropoda" family="Hydrobiidae" genus="Falsipyrgula" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Falsipyrgula" order="Littorinimorpha" pageId="0" pageNumber="25" phylum="Mollusca" rank="genus">
|
||
<emphasis id="FCF5034DC649321E579A1F199D0DF3C2" italics="true" pageId="0" pageNumber="25">Falsipyrgula</emphasis>
|
||
</taxonomicName>
|
||
sp. in Eflatun
|
||
<normalizedToken id="E6841C7D4A4742EBF7AD2E14792B5C68" originalValue="Pınarı">Pinari</normalizedToken>
|
||
, and
|
||
<taxonomicName id="CAE2E33253D6271F60E748E33F378EBF" class="Gastropoda" family="Hydrobiidae" genus="Pseudamnicola" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudamnicola natolica" order="Littorinimorpha" pageId="0" pageNumber="25" phylum="Mollusca" rank="species" species="natolica">
|
||
<emphasis id="2A9860F926D67C9A617EA5D7ACA35C9D" italics="true" pageId="0" pageNumber="25">Pseudamnicola natolica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<normalizedToken id="D19EA61DF97D316BBD5C22935817B481" originalValue="Küster">Kuester</normalizedToken>
|
||
, 1852) (and possibly also
|
||
<taxonomicName id="1C7BCA81288FE2CD8E06951144CBD0B6" lsidName="T. gloeri" pageId="0" pageNumber="25" rank="species" species="gloeri">
|
||
<emphasis id="88E26B058060AD9CA8704839F3955EC4" italics="true" pageId="0" pageNumber="25">T. gloeri</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="D024D435B781BBED6334C8DEB68096E2" author="Odabasi, DA" journalOrPublisher="Ecologica Montenegrina" pageId="0" pageNumber="25" pagination="327 - 333" refId="B115" refString="Odabasi, DA, Arslan, N, 2015. Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey). Ecologica Montenegrina 2 (4): 327 - 333" title="Description of a new subterranean nerite: Theodoxus gloeri n. sp. with some data on the freshwater gastropod fauna of Balikdami Wetland (Sakarya River, Turkey)." volume="2" year="2015">
|
||
<normalizedToken id="65E21EBA65F0C71EEDE6A97F986A08EA" originalValue="Odabaşı">Odabasi</normalizedToken>
|
||
and Arslan 2015
|
||
</bibRefCitation>
|
||
) in the
|
||
<normalizedToken id="3EABFCA69E984CDFBF0CDFBCE671FC45" originalValue="Balıkdamı">Balikdami</normalizedToken>
|
||
Wetland spring (personal observation M.E.G.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |