153 lines
14 KiB
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153 lines
14 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.656.8914" ID-GBIF-Dataset="51a5b313-2a8b-4bde-8d9d-06bd051fcb42" ID-PMC="PMC5345363" ID-Pensoft-Pub="1313-2970-656-85" ID-PubMed="28331403" ID-ZBK="89234B7B2F01446EB8C077C29CE106AE" ModsDocAuthor="" ModsDocDate="2017" ModsDocID="1313-2970-656-85" ModsDocOrigin="ZooKeys 656" ModsDocTitle="The genus Alaolacon Candèze, a senior synonym of the genus Eumoeus Candèze (Coleoptera, Elateridae, Agrypninae)" checkinTime="1487100179832" checkinUser="pensoft" docAuthor="Arimoto, Koichi & Arimoto, Hisayuki" docDate="2017" docId="55E90B738DDF29699A12FBBFD72D3D07" docLanguage="en" docName="ZooKeys 656: 85-110" docOrigin="ZooKeys 656" docSource="http://dx.doi.org/10.3897/zookeys.656.8914" docTitle="Alaolacon fujiokai Arimoto & Arimoto, 2017, sp. n." docType="treatment" docUuid="FF52714A-2F8B-413C-9777-C96D283C3465" docUuidSource="ZooBank" docVersion="5" lastPageNumber="95" masterDocId="901EFFE5FFA1FFA53456FFDC5C68FFFC" masterDocTitle="The genus Alaolacon Candeze, a senior synonym of the genus Eumoeus Candeze (Coleoptera, Elateridae, Agrypninae)" masterLastPageNumber="110" masterPageNumber="85" pageNumber="94" updateTime="1668164237870" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>The genus Alaolacon Candeze, a senior synonym of the genus Eumoeus Candeze (Coleoptera, Elateridae, Agrypninae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Arimoto, Koichi</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Arimoto, Hisayuki</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2017</mods:date>
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<mods:detail type="volume">
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<mods:number>656</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>85</mods:start>
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<mods:end>110</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.656.8914</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.656.8914</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-656-85</mods:identifier>
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<mods:identifier type="ZBK">89234B7B2F01446EB8C077C29CE106AE</mods:identifier>
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<mods:identifier type="ZooBank">89234B7B2F01446EB8C077C29CE106AE</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127855044" LSID="urn:lsid:zoobank.org:act:FF52714A-2F8B-413C-9777-C96D283C3465" httpUri="http://treatment.plazi.org/id/55E90B738DDF29699A12FBBFD72D3D07" lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94">
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<subSubSection pageId="9" pageNumber="94" type="nomenclature">
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<paragraph pageId="9" pageNumber="94">
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<taxonomicName LSID="http://zoobank.org/FF52714A-2F8B-413C-9777-C96D283C3465" class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon fujiokai" order="Coleoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="fujiokai">Alaolacon fujiokai</taxonomicName>
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<taxonomicNameLabel pageId="9" pageNumber="94">sp. n.</taxonomicNameLabel>
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Figures 20-21, 22-29, 30-35, 36-37
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</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="etymology">
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<paragraph pageId="9" pageNumber="94">Etymology.</paragraph>
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<paragraph pageId="9" pageNumber="94">The name of this species honors Mr. Masahiro Fujioka for providing the material.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="type material">
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<paragraph pageId="9" pageNumber="94">Type material.</paragraph>
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<paragraph pageId="9" pageNumber="94">Holotype. Male, Tawau, East Malaysia (Sabah), Malaysia, V 2014 [ELKU].</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="diagnosis">
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<paragraph pageId="9" pageNumber="94">Diagnosis.</paragraph>
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<paragraph pageId="9" pageNumber="94">Body black, elytra blue and with metallic luster, legs black; setae black on dorsal side and white on ventral side; frontal depression deep; eye small; prothorax almost as long as wide, widest posteriorly; pronotum with anterior angles bisinuate and rounded, medina longitudinal depression deep, extending from before pronotal anterior margin to base, punctate; prosternal process inclined strongly behind procoxae; anterior angles of hypomeron acute; scutellum concave laterally, widest near posterior 1/3; hind wings with cross vein between veins MP4 and CuA2 located anterad to contact point between veins MP3 and MP4, without wedge cell; median lobe of male aedeagus stout.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="description">
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<paragraph pageId="9" pageNumber="94">Measurements.</paragraph>
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<paragraph pageId="9" pageNumber="94">BL: 18.9, BW: 6.11, MIE: 2.08, MAE: 3.05, OI: 68, PL: 5.95, PML: 5.23; PW: 5.91, PI: 101, EL: 12.1, EW: 6.11, EI: 197.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="description">
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<paragraph pageId="9" pageNumber="94">Description of male.</paragraph>
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<paragraph pageId="9" pageNumber="94">Body (Figs 20, 21) shiny, elytra with metallic luster. Color. Black except for elytra black-blue; mouth-parts brown-black, but mandible black, galea and lacinia orange; apical edge of tarsal segment V and tarsal claws red-brown; pregenital segments and aedeagus black-brown. Hairs. Dorsal surface covered with black flatted setae; ventral surface with white flatted setae; legs with intermixed black and white setae; mouth-parts and pronotal anterior margin near eyes with yellow-brown setae; filiform setae of antennomeres III-XII brown and long.</paragraph>
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<caption pageId="9" pageNumber="94">
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<paragraph pageId="9" pageNumber="94">
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Figures 20, 21.
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon fujiokai" order="Coleoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="fujiokai">Alaolacon fujiokai</taxonomicName>
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sp. n., male, holotype. 20 habitus, dorsal view 21 ditto, ventral view.
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</paragraph>
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</caption>
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<paragraph pageId="9" pageNumber="94">
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Head (Fig. 22). Frontal depression deep. Eyes small. Antennomere I long; antennomere II short, dish-shaped; antennomeres III-XI flabellation strong; antennomere XII elongate (Fig. 23). Apical maxillary palpomere 1.8
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide (Fig. 24) (Mandibles chipped in apical parts.)
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</paragraph>
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<caption pageId="9" pageNumber="94">
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<paragraph pageId="9" pageNumber="94">
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Figures 22-29.
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon fujiokai" order="Coleoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="fujiokai">Alaolacon fujiokai</taxonomicName>
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sp. n., male, holotype. 22 head and pronotum, anterolateral view 23 right antenna, dorsal view 24 mouth-parts 25 prosternal process, lateral view 26 anterior angle of hypomeron (arrow: mesal edge carinate anterolaterad) 27 posterior part of hypomeron and mesothorax, ventral view (arrows: posterior margin with two angles) 28 scutellum 29 right hind wing (arrow: cross vein between veins MP4 and CuA2 located anterad to contact point between veins MP3 and MP4).
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</paragraph>
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</caption>
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<paragraph pageId="9" pageNumber="94">
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Prothorax almost as long as wide, widest posteriorly; sides rounded anteriorly, liner posteriorly. Pronotum with anterior angles bisinuate and rounded; median longitudinal depression deep, extending from before pronotal anterior margin to base, punctate. Prosternal spine inclined strongly (at 18 degrees) behind procoxae (Fig. 25). Hypomeron with anterior angles acute (Fig. 26). Scutellum (Fig. 28) concave laterally, 1.2
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, widest near posterior 1/3. Hind wings with cross vein between veins MP4 and CuA2 apparent, not completely connected with CuA2, located anterad to contact point between veins MP3 and MP4 (Fig. 29: arrow); wedge cell absent. Elytra with sides almost parallel on basal half; intervals with small and coarse punctures.
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</paragraph>
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<paragraph pageId="9" pageNumber="94">
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Abdomen. Ventrite V 0.67
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<normalizedToken originalValue="×">x</normalizedToken>
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times as long as wide. Tergite VIII (Fig. 30) 0.72
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<normalizedToken originalValue="×">x</normalizedToken>
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as long as wide, colorless basal area. Sternite VIII (Fig. 31) with darker W-shaped band; median notch shallow and truncate transversally. Tergite IX (Fig. 32) with median notch shallow and rounded. Sternite IX (Fig. 33) narrowed abruptly on posterior half to apex. Aedeagus (Figs 34, 35). Median lobe stout; basal struts 0.35
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<normalizedToken originalValue="×">x</normalizedToken>
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total length of median lobe. Parameres with dense and long setae. Basal piece 0.29
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<normalizedToken originalValue="×">x</normalizedToken>
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total length of aedeagus.
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</paragraph>
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<caption pageId="9" pageNumber="94">
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<paragraph pageId="9" pageNumber="94">
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Figures 30-35.
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon fujiokai" order="Coleoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="fujiokai">Alaolacon fujiokai</taxonomicName>
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sp. n., male, holotype. 30 tergite VIII 31 sternite VIII 32 tergites
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<normalizedToken originalValue="IX–X">IX-X</normalizedToken>
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33 sternites
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<normalizedToken originalValue="IX–X">IX-X</normalizedToken>
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34 aedeagus, dorsal view 35 ditto, ventral view.
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</paragraph>
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</caption>
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<caption pageId="9" pageNumber="94">
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<paragraph pageId="9" pageNumber="94">
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Figures 36, 37.
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon fujiokai" order="Coleoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="fujiokai">Alaolacon fujiokai</taxonomicName>
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sp. n., male, holotype, setae of elytra. 36 median part 37 apical part.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="female">
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<paragraph pageId="9" pageNumber="94">Female.</paragraph>
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<paragraph pageId="9" pageNumber="94">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="9" pageNumber="94" type="distribution">
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<paragraph pageId="9" pageNumber="94">Distribution.</paragraph>
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<paragraph pageId="9" pageNumber="94">Malaysia: Sabah: Tawau.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94" type="remarks">
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<paragraph pageId="9" pageNumber="94">Remarks.</paragraph>
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<paragraph lastPageId="10" lastPageNumber="95" pageId="9" pageNumber="94">
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This species is distinct by black body, blue elytra with metallic luster, black setae on dorsal side, white setae on ventral side and strong antennomeres III-XI flabellation. It is similar to
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon candezei" order="Coleoptera" pageId="9" pageNumber="94" phylum="Arthropoda" rank="species" species="candezei">Alaolacon candezei</taxonomicName>
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Fleutiaux, 1928 in having a black body, blue elytra with metallic luster, pronotum anterior angles bisinuate and rounded, and
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<pageBreakToken pageId="10" pageNumber="95" start="start">scutellum</pageBreakToken>
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concave laterally, except for drastic sexual differences of antennae, but is distinguished by the following contrasting characters (
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon candezei" order="Coleoptera" pageId="10" pageNumber="95" phylum="Arthropoda" rank="species" species="candezei">Alaolacon candezei</taxonomicName>
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in parentheses): legs black (Fig. 21) (legs red, Fig. 2); setae black on dorsal side and white on ventral side (Fig. 20) (all setae white, Fig. 1); frontal depression deep (Fig. 22) (frontal depression moderate, Fig. 4); pronotal median longitudinal depression extending from before pronotal anterior margin to base (Fig. 22) (pronotal median longitudinal depression not reaching anterior margin or base, Fig. 4); anterior angles of hypomeron acute (Fig. 26) (anterior angles of hypomeron rounded, Fig. 8); prosternal spin inclined strongly behind procoxae (Fig. 25) (prosternal spine inclined weakly behind procoxae, Fig. 7); scutellum widest near posterior 1/3 (Fig. 28) (scutellum widest near posterior 2/5, Fig. 10); hind wings with cross vein between veins MP4 and CuA2 located anterad to contact point between veins MP3 and MP4 (Fig. 29: arrow) (hind wings with cross vein between veins MP4 and CuA2 located at contact point between veins MP3 and MP4, Fig. 11: arrow).
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</paragraph>
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<paragraph pageId="10" pageNumber="95">
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon fujiokai" order="Coleoptera" pageId="10" pageNumber="95" phylum="Arthropoda" rank="species" species="fujiokai">Alaolacon fujiokai</taxonomicName>
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and
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<taxonomicName class="Insecta" family="Elateridae" genus="Alaolacon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Alaolacon candezei" order="Coleoptera" pageId="10" pageNumber="95" phylum="Arthropoda" rank="species" species="candezei">Alaolacon candezei</taxonomicName>
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are similar species from the same island, but we recognized they are different species by the setal color and the hind wing venation. We believe that setal complementary color difference probably is not caused by sexual dimorphism because such dimorphism has not previously been observed in species of the
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<taxonomicName lsidName="" pageId="10" pageNumber="95" rank="subfamily" subfamily="Agrypninae">Agrypninae</taxonomicName>
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. We also believe that differences in hind wing venation are unlikely to be caused by sexual dimorphism because such dimorphism has not previously been observed in species with flying females.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |