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282 lines
58 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.881.36744" ID-GBIF-Dataset="3cef5ca2-433f-45f7-a32b-087b5f1946a4" ID-PMC="PMC6813179" ID-Pensoft-Pub="1313-2970-881-135" ID-Pensoft-UUID="16F69FEA95835AE5BF6B9C34F83E49D7" ID-PubMed="31662614" ID-ZooBank="B058CA156A3D41A19ADFD00384234D8E" ModsDocID="1313-2970-881-135" checkinTime="1571384733377" checkinUser="pensoft" docAuthor="Mvogo Ndongo, Pierre A., von Rintelen, Thomas & Cumberlidge, Neil" docDate="2019" docId="55AE8B61BEBF5CFC925B97D5C7A7C0B5" docLanguage="en" docName="ZooKeys 881: 135-164" docOrigin="ZooKeys 881" docSource="http://dx.doi.org/10.3897/zookeys.881.36744" docTitle="Louisea yabassi Ndongo, Rintelen & Cumberlidge, 2019, sp. nov." docType="treatment" docUuid="FA6DE8AD-B833-415C-95C7-47287F3C6158" docUuidSource="ZooBank" docVersion="6" id="16F69FEA95835AE5BF6B9C34F83E49D7" lastPageNumber="135" masterDocId="16F69FEA95835AE5BF6B9C34F83E49D7" masterDocTitle="Taxonomic revision of the endemic Cameroonian freshwater crab genus Louisea Cumberlidge, 1994 (Crustacea, Decapoda, Brachyura, Potamonautidae), with descriptions of two new species from Nkongsamba and Yabassi" masterLastPageNumber="164" masterPageNumber="135" pageNumber="135" updateTime="1668167896540" updateUser="ExternalLinkService">
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<mods:title>Taxonomic revision of the endemic Cameroonian freshwater crab genus Louisea Cumberlidge, 1994 (Crustacea, Decapoda, Brachyura, Potamonautidae), with descriptions of two new species from Nkongsamba and Yabassi</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Mvogo Ndongo, Pierre A.</mods:namePart>
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<mods:namePart>von Rintelen, Thomas</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Cumberlidge, Neil</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
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<mods:detail type="volume">
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<mods:number>881</mods:number>
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<mods:start>135</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.881.36744</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.881.36744</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-881-135</mods:identifier>
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<treatment ID-GBIF-Taxon="159754740" LSID="urn:lsid:zoobank.org:act:FA6DE8AD-B833-415C-95C7-47287F3C6158" httpUri="http://treatment.plazi.org/id/55AE8B61BEBF5CFC925B97D5C7A7C0B5" lastPageNumber="135" pageId="0" pageNumber="135">
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<taxonomicName LSID="55AE8B61-BEBF-5CFC-925B-97D5C7A7C0B5" class="Malacostraca" family="Potamonautidae" genus="Louisea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Louisea yabassi" order="Decapoda" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yabassi">Louisea yabassi</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="135">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Largest adult male (CW 17.5 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 26930) (missing right cheliped) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 1 ', www. mapress. com / j / zt) c largest adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 5, and right cheliped and p 2, p 3) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 8 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure2" httpUri="https://binary.pensoft.net/fig/347890" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure2">Figs 2c</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) (missing left p 4) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 4 and right p 2) c subadult male, paratype (CW 13.8 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 4, and right cheliped and p 2 - p 5) d second largest adult male (CW 18.38 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 10 mm (b), 8.42 mm (c), 11.63 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure3" httpUri="https://binary.pensoft.net/fig/347891" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure3">3c</figureCitation>
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,
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">4d</figureCitation>
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,
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">5c</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">6c</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7c</figureCitation>
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<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">8e, f</figureCitation>
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<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Four species of Louisea endemic to southwestern Cameroon, right cheliped merus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 d', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure9" httpUri="https://binary.pensoft.net/fig/347897" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure9">9c</figureCitation>
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,
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<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Four species of Louisea endemic to southwestern Cameroon, right cheliped carpus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 3 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure10" httpUri="https://binary.pensoft.net/fig/347898" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure10">10c</figureCitation>
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,
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">11c</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">12c</figureCitation>
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,
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Four species of Louisea endemic to southwestern Cameroon, G 2 a second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure13" httpUri="https://binary.pensoft.net/fig/347901" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure13">13c</figureCitation>
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<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Four species of Louisea endemic to southwestern Cameroon, left third maxilliped. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 4 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure14" httpUri="https://binary.pensoft.net/fig/347902" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure14">14c</figureCitation>
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<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Four species of Louisea endemic to southwestern Cameroon, ventral view of right mandible. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (a, c), 2 mm (b, d)." figureDoi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure15">15c</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="135" type="materials_examined">
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<paragraph pageId="0" pageNumber="135">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="135">CAMEROON. Holotype: adult male (CW 18.11 mm, CL 12.78 mm, CH 8.30 mm, FW 6.29 mm; CW/FW 2.88, CL/FW 2.03, CH/FW 1.32, FW/CL 0.49, FW/CW 0.34), Yabassi, 10 September 1909, coll. Riggenbach (ZMB Crust. 21575). Paratype: subadult male (CW 13.82 mm, CL 10.61 mm, CH 6.25 mm, FW 5.01 mm; CW/FW 2.75, CL/FW 2.11, CH/FW 1.24, FW/CL 0.47, FW/CW 0.36), same data as holotype (ZMB Crust. 21575).</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="135" type="diagnosis">
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<paragraph pageId="0" pageNumber="135">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="135">
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Carapace smooth, urogastric groove faint; postfrontal crest distinct, prominent, complete, meeting anterolateral margin behind intermediate tooth (
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Four species of Louisea endemic to southwestern Cameroon, whole animal, dorsal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) (missing left p 4) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 4 and right p 2) c subadult male, paratype (CW 13.8 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) (missing left p 2 - p 4, and right cheliped and p 2 - p 5) d second largest adult male (CW 18.38 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 10 mm (b), 8.42 mm (c), 11.63 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure3" httpUri="https://binary.pensoft.net/fig/347891" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure3">Fig. 3c</figureCitation>
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); exorbital, intermediate teeth large, triangular; epibranchial tooth undetectable (
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">Figs 4c</figureCitation>
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,
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<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7c</figureCitation>
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). Vertical sulcus on carapace branchiostegal wall curving backward to meet anterolateral margin at epibranchial tooth (
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6c</figureCitation>
|
||
). Mandibular palp bi-segmented; terminal segment (TS) bilobed, with large distinct anterior lobe 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
terminal segment length (
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Four species of Louisea endemic to southwestern Cameroon, ventral view of right mandible. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (a, c), 2 mm (b, d)." figureDoi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure15">Fig. 15c</figureCitation>
|
||
). Third maxilliped ischium with distinct vertical groove (
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Four species of Louisea endemic to southwestern Cameroon, left third maxilliped. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 4 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure14" httpUri="https://binary.pensoft.net/fig/347902" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure14">Fig. 14c</figureCitation>
|
||
). Episternal sulci s4/e4, s5/e5, s6/e6 faint or missing, s7/e7 complete (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5c</figureCitation>
|
||
). Major cheliped dactylus highly arched enclosing oval interspace when closed, with five large teeth (one small distal, two large medial, two small proximal) (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8e</figureCitation>
|
||
); propodus of major cheliped with two large proximal teeth, large medial tooth, small distal tooth (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8e</figureCitation>
|
||
); cheliped carpus inner margin with long, broad distal tooth, relatively narrow, subequal proximal tooth (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Four species of Louisea endemic to southwestern Cameroon, right cheliped carpus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 3 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure10" httpUri="https://binary.pensoft.net/fig/347898" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure10">Fig. 10c</figureCitation>
|
||
). G1TA short (TA/SS 0.22), directed outwards at 45° angle to longitudinal axis of G1SS, with distinct longitudinal groove proximally distinctly broad, abruptly narrow, slim and tube-like at distal two-thirds (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Figs 11c</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">12c</figureCitation>
|
||
). G1SS tapering slightly from broad basal margin to relatively wide distal margin (0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS basal margin); dorsal face with broad dorsal membrane (maximum width 0.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS length) at TA/SS junction (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Fig. 11c</figureCitation>
|
||
). G2TA long (TA/SS 0.44), flagellum-like, almost as long as G2SS (
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Four species of Louisea endemic to southwestern Cameroon, G 2 a second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure13" httpUri="https://binary.pensoft.net/fig/347901" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure13">Fig. 13c</figureCitation>
|
||
). Mature at CW 19 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="description">
|
||
<paragraph pageId="0" pageNumber="135">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Carapace ovoid, flat (CH/FW 1.28,
|
||
<emphasis italics="true" pageId="0" pageNumber="135">N</emphasis>
|
||
= 2), wide (CW/FW 2.8), smooth, urogastric groove distinct; front wide (FW/CW 0.35,
|
||
<emphasis italics="true" pageId="0" pageNumber="135">N</emphasis>
|
||
= 2), deflexed, anterior margin straight; postfrontal crest distinct, prominent, completely crossing carapace, meeting anterolateral margin of carapace behind intermediate tooth (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">Fig. 4d</figureCitation>
|
||
); exorbital, intermediate teeth large, triangular; epibranchial tooth undetectable (
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Four species of Louisea endemic to southwestern Cameroon, carapace, dorsal view. a Largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) d adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575). Scale bars: 9.1 mm (a), 7.25 mm (b), 7.90 mm (c), 8.62 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure4" httpUri="https://binary.pensoft.net/fig/347892" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure4">Figs 4c</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Four species of Louisea endemic to Southwestern Cameroon, carapace, frontal view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 7.20 mm (a), 6.60 mm (b), 8.06 mm (c), 8.70 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure7" httpUri="https://binary.pensoft.net/fig/347895" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure7">7c</figureCitation>
|
||
). Carapace branchiostegal sidewall with vertical, longitudinal sutures dividing it into three regions (suborbital, subhepatic, pterygostomial) (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6c</figureCitation>
|
||
); longitudinal suture beginning at respiratory opening, curving backward across sidewall dividing suborbital- and subhepatic regions from pterygostomial region (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6c</figureCitation>
|
||
); vertical sulcus on carapace branchiostegal wall curving backward to meet anterolateral margin at epibranchial tooth (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6c</figureCitation>
|
||
), dividing suborbital from subhepatic regions (
|
||
<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Four species of Louisea endemic to southwestern Cameroon, whole animal, ventral view. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b second largest adult male (CW 14.8 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 11.63 mm (a), 9.61 mm (b), 9.50 mm (c), 11.60 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure6" httpUri="https://binary.pensoft.net/fig/347894" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure6">Fig. 6c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Mandibular palp bi-segmented; terminal segment (TS) bilobed, with large distinct anterior lobe 0.6
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
terminal segment length (
|
||
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Four species of Louisea endemic to southwestern Cameroon, ventral view of right mandible. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (a, c), 2 mm (b, d)." figureDoi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure15">Fig. 15c</figureCitation>
|
||
). Third maxilliped exopod completely lacking flagellum; ischium with distinct vertical groove (
|
||
<figureCitation captionStart="Figure 14" captionStartId="F14" captionText="Figure 14. Four species of Louisea endemic to southwestern Cameroon, left third maxilliped. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 4 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure14" httpUri="https://binary.pensoft.net/fig/347902" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure14">Fig. 14c</figureCitation>
|
||
). Sternal sulcus s2/s3 prominent, completely crossing sternum; s3/s4 incomplete, reduced to 2 short lateral notches (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5c</figureCitation>
|
||
). Episternal sulci s4/e4, s5/e5, s6/e6 faint or missing, s7/e7 complete (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Male chelipeds unequal, right chelipeds larger than left cheliped; fingers slim, elongated. Right (major) cheliped dactylus relatively stout, straight, with five large teeth (one small distal, two large medial, two small proximal); fixed finger (pollex) with five large teeth (one distal, four proximal) (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8e</figureCitation>
|
||
); dactylus and propodus of left cheliped straight, with small teeth on occluding margin (
|
||
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Four species of Louisea endemic to Southwestern Cameroon, frontal view of right and left chela. a, b Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) c, d largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 a, b', www. mapress. com / j / zt) e, f adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) g, h largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c, e, g), 2 mm (b, d, f), 10 mm (h)." figureDoi="10.3897/zookeys.881.36744.figure8" httpUri="https://binary.pensoft.net/fig/347896" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure8">Fig. 8f</figureCitation>
|
||
); cheliped carpus inner margin with two pointed teeth, distal tooth large with pointed tip, proximal tooth smaller (
|
||
<figureCitation captionStart="Figure 10" captionStartId="F10" captionText="Figure 10. Four species of Louisea endemic to southwestern Cameroon, right cheliped carpus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 3 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure10" httpUri="https://binary.pensoft.net/fig/347898" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure10">Fig. 10c</figureCitation>
|
||
); medial inferior margin of cheliped merus with large jagged distal tooth followed by numerous distinct smaller teeth (
|
||
<figureCitation captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Four species of Louisea endemic to southwestern Cameroon, right cheliped merus. a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 3 d', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 5 mm (a, c), 2 mm (b), 10 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure9" httpUri="https://binary.pensoft.net/fig/347897" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure9">Fig. 9c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Male pleon triangular, sides not indented with small setae; telson (a7) rounded at distal margin (
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Four species of Louisea endemic to southwestern Cameroon, thoracic sternites (s 1 - s 8) and pleonal segments (a 4 - a 7). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (missing left p 5) (' Mvogo Ndongo et al. 2018: fig. 2 c', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 12.42 mm (a), 9 mm (b), 8.30 mm (c), 17 mm (d)." figureDoi="10.3897/zookeys.881.36744.figure5" httpUri="https://binary.pensoft.net/fig/347893" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure5">Fig. 5c</figureCitation>
|
||
). G1TA short (TA/SS 0.22), directed outwards at 45° angle to longitudinal axis of G1SS, with distinct longitudinal groove on ventral face, proximally distinctly broad, abruptly narrow, slim and tube-like at distal two-thirds (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Figs 11c</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">12c</figureCitation>
|
||
). G1SS tapering slightly from broad basal margin to relatively wide distal margin (0.5
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS basal margin); ventral face of with raised triangular flap extending halfway across segment forming roof of chamber for G2, flap tapering diagonally from broad base to narrow point at SS/TA junction (
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Four species of Louisea endemic to southwestern Cameroon, right G 1 ventral view (a-c), left G 1 ventral view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 b') c adult male, holotype (CW 18.1 mm) of L. yabassi from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure12" httpUri="https://binary.pensoft.net/fig/347900" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure12">Fig. 12c</figureCitation>
|
||
); dorsal face with broad dorsal membrane (maximum width 0.1
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
SS length) at TA/SS junction (
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Four species of Louisea endemic to southwestern Cameroon, right G 1 dorsal view (a-c), left G 1 dorsal view (d). a Second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) (' Mvogo Ndongo et al. 2018: fig. 5 a', www. mapress. com / j / zt) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a, c, d), 1 mm (b)." figureDoi="10.3897/zookeys.881.36744.figure11" httpUri="https://binary.pensoft.net/fig/347899" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure11">Fig. 11c</figureCitation>
|
||
). G2TA long (TA/SS 0.44), flagellum-like, almost as long as G2SS (
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Four species of Louisea endemic to southwestern Cameroon, G 2 a second largest adult male (CW 16.1 mm) of L. edeaensis from Lake Ossa wetland complex (ZMB Crust. 30335) b largest adult male (CW 16.2 mm) of L. balssi from Man's Crater Lake Manengouba (ZMB Crust. 30319) c adult male, holotype (CW 18.1 mm) of L. yabassi sp. nov. from Yabassi (ZMB Crust. 21575) d largest adult male, holotype (CW 20.0 mm) of L. nkongsamba sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 2 mm (a-d)." figureDoi="10.3897/zookeys.881.36744.figure13" httpUri="https://binary.pensoft.net/fig/347901" pageId="0" pageNumber="135" tableDoi="10.3897/zookeys.881.36744.figure13">Fig. 13c</figureCitation>
|
||
). Mature at CW 19 mm.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="color in life">
|
||
<paragraph pageId="0" pageNumber="135">Color in life.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">Unknown.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="type locality">
|
||
<paragraph pageId="0" pageNumber="135">Type locality.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">Yabassi, littoral region of Cameroon.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="etymology">
|
||
<paragraph pageId="0" pageNumber="135">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">The new species is named for Yabassi in south-western Cameroon, where it was collected in 1909. The species epithet is a noun in apposition.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="habitat">
|
||
<paragraph pageId="0" pageNumber="135">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
<taxonomicName class="Malacostraca" family="Potamonautidae" genus="Louisea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Louisea yabassi" order="Decapoda" pageId="0" pageNumber="135" phylum="Arthropoda" rank="species" species="yabassi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">Louisea yabassi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. is known only from Yabassi, a humid area of the coastal rain forest of southwestern Cameroon. It is possible that this species is also present in the Ebo forest near Yabassi, which is one of the largest remaining tracts of lowland and submontane rainforest in the area.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="135" type="remarks">
|
||
<paragraph pageId="0" pageNumber="135">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="135">
|
||
Characters of the carapace and chelipeds of adult male specimens from Yabassi assigned to
|
||
<taxonomicName lsidName="L. edeaensis" pageId="0" pageNumber="135" rank="species" species="edeaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. edeaensis</emphasis>
|
||
</taxonomicName>
|
||
by
|
||
<bibRefCitation author="Cumberlidge, N" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="135" pagination="122 - 131" refId="B8" refString="Cumberlidge, N, 1994. Louisea, a new genus of freshwater crab (Brachyura: Potamoidea: Potamonautidae) for Globonautesmacropusedeaensis Bott, 1969 from Cameroon. Proceedings of the Biological Society of Washington 107: 122 - 131" title="Louisea, a new genus of freshwater crab (Brachyura: Potamoidea: Potamonautidae) for Globonautesmacropusedeaensis Bott, 1969 from Cameroon." volume="107" year="1994">Cumberlidge (1994)</bibRefCitation>
|
||
and by
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4231.2.9" author="Mvogo Ndongo, PA" journalOrPublisher="Zootaxa" pageId="0" pageNumber="135" pagination="273 - 280" refId="B26" refString="Mvogo Ndongo, PA, von Rintelen, T, Schubart, CD, Albrecht, C, Tamesse, JL, Cumberlidge, N, 2017a. New data on the taxonomy, ecology, and conservation of the rediscovered Louiseaedeaensis (Bott, 1969) (Brachyura: Potamoidea: Potamonautidae), an endangered freshwater crab from Cameroon. Zootaxa 4231: 273 - 280" title="New data on the taxonomy, ecology, and conservation of the rediscovered Louiseaedeaensis (Bott, 1969) (Brachyura: Potamoidea: Potamonautidae), an endangered freshwater crab from Cameroon." url="https://doi.org/10.11646/zootaxa.4231.2.9" volume="4231" year="2017 a">Mvogo Ndongo et al. (2017a)</bibRefCitation>
|
||
proved to be inconsistent with the holotype from Edea. Re-examination of these specimens supported the hypothesis that specimens from Edea and those from Yabassi belong to two different species:
|
||
<taxonomicName lsidName="L. edeaensis" pageId="0" pageNumber="135" rank="species" species="edeaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. edeaensis</emphasis>
|
||
</taxonomicName>
|
||
(Edea, Yaounde, and Lake Ossa) and
|
||
<taxonomicName lsidName="L. yabassi" pageId="0" pageNumber="135" rank="species" species="yabassi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. yabassi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. (Yabassi). The differences between
|
||
<taxonomicName lsidName="L. yabassi" pageId="0" pageNumber="135" rank="species" species="yabassi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. yabassi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and its congeners are discussed below under general remarks.
|
||
</paragraph>
|
||
<caption doi="10.3897/zookeys.881.36744.figure15" httpUri="https://binary.pensoft.net/fig/347903" pageId="0" pageNumber="135" start="Figure 15" startId="F15">
|
||
<paragraph pageId="0" pageNumber="135">
|
||
<emphasis bold="true" pageId="0" pageNumber="135">Figure 15.</emphasis>
|
||
Four species of
|
||
<taxonomicName class="Malacostraca" family="Potamonautidae" genus="Louisea" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Louisea" order="Decapoda" pageId="0" pageNumber="135" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">Louisea</emphasis>
|
||
</taxonomicName>
|
||
endemic to southwestern Cameroon, ventral view of right mandible.
|
||
<emphasis bold="true" pageId="0" pageNumber="135">a</emphasis>
|
||
Second largest adult male (CW 16.1 mm) of
|
||
<taxonomicName lsidName="L. edeaensis" pageId="0" pageNumber="135" rank="species" species="edeaensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. edeaensis</emphasis>
|
||
</taxonomicName>
|
||
from Lake Ossa wetland complex (ZMB Crust. 30335)
|
||
<emphasis bold="true" pageId="0" pageNumber="135">b</emphasis>
|
||
largest adult male (CW 16.2 mm) of
|
||
<taxonomicName lsidName="L. balssi" pageId="0" pageNumber="135" rank="species" species="balssi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. balssi</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<normalizedToken originalValue="Man’s">Man's</normalizedToken>
|
||
Crater Lake Manengouba (ZMB Crust. 30319)
|
||
<emphasis bold="true" pageId="0" pageNumber="135">c</emphasis>
|
||
adult male, holotype (CW 18.1 mm) of
|
||
<taxonomicName lsidName="L. yabassi" pageId="0" pageNumber="135" rank="species" species="yabassi">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. yabassi</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Yabassi (ZMB Crust. 21575)
|
||
<emphasis bold="true" pageId="0" pageNumber="135">d</emphasis>
|
||
largest adult male, holotype (CW 20.0 mm) of
|
||
<taxonomicName lsidName="L. nkongsamba" pageId="0" pageNumber="135" rank="species" species="nkongsamba">
|
||
<emphasis italics="true" pageId="0" pageNumber="135">L. nkongsamba</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. from Mt. Nlonako (ZMB Crust. 31618). Scale bars: 1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="135">a, c</emphasis>
|
||
), 2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="135">b, d</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |