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<document ID-CLB-Dataset="27209" ID-DOI="http://dx.doi.org/10.3897/phytokeys.107.26548" ID-GBIF-Dataset="b09b328e-de27-4071-8af6-eec29dc3b1df" ID-PMC="PMC6160804" ID-Pensoft-Pub="1314-2003-107-1" ID-Pensoft-UUID="686CFF85FFADFFEAC033443CF54BFFFC" ID-PubMed="30271249" ID-Zenodo-Dep="1346433" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-2003-107-1" ModsDocOrigin="PhytoKeys 107" ModsDocTitle="A revision of Dissochaeta (Melastomataceae, Dissochaeteae)" checkinTime="1555330651054" checkinUser="pensoft" docAuthor="Kartonegoro, Abdulrokhman, Veldkamp, Jan Frits, Hovenkamp, Peter &amp; Welzen, Peter van" docDate="2018" docId="55A38A86675790DB1AB00F0212683AD2" docLanguage="en" docName="PhytoKeys 107: 1-178" docOrigin="PhytoKeys 107" docSource="http://dx.doi.org/10.3897/phytokeys.107.26548" docTitle="Dissochaeta horrida Kartonegoro &amp; Veldkamp &amp; Hovenkamp &amp; Welzen 2018, comb. nov." docType="treatment" docVersion="8" id="686CFF85FFADFFEAC033443CF54BFFFC" lastPageNumber="1" masterDocId="686CFF85FFADFFEAC033443CF54BFFFC" masterDocTitle="A revision of Dissochaeta (Melastomataceae, Dissochaeteae)" masterLastPageNumber="178" masterPageNumber="1" pageNumber="1" updateTime="1732647791036" updateUser="ExternalLinkService">
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<mods:title id="416144097ADC39A646CEE20536BFDE98">A revision of Dissochaeta (Melastomataceae, Dissochaeteae)</mods:title>
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<mods:namePart id="AA139F68C8781744A0BF75E992106996">Kartonegoro, Abdulrokhman</mods:namePart>
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<mods:namePart id="7412D0BF20444B3C8C09D87FA06034A6">Veldkamp, Jan Frits</mods:namePart>
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<mods:namePart id="73C53C92F3096340590959E7C205578D">Hovenkamp, Peter</mods:namePart>
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<mods:namePart id="204801A1F8D14ED38145023A2F898EF1">Welzen, Peter van</mods:namePart>
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<mods:title id="C8268822152E0EC72DC0A1DD989121FF">PhytoKeys</mods:title>
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<mods:date id="459DDD99696F0DD6EB18FBFA82096402">2018</mods:date>
<mods:detail id="3B04E052AB06B0FB2F3B425D0ACBFD14" type="volume">
<mods:number id="AB1191EC28864DC6F25772CF7FC74107">107</mods:number>
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<mods:identifier id="00C296D13F70843CB5E0BB36B3B0A92B" type="DOI">http://dx.doi.org/10.3897/phytokeys.107.26548</mods:identifier>
<mods:identifier id="B3FDDF18088A36AA00AD5F954502F8CD" type="Pensoft-Pub">1314-2003-107-1</mods:identifier>
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<treatment id="55A38A86675790DB1AB00F0212683AD2" ID-GBIF-Taxon="156201501" LSID="urn:lsid:plazi:treatment:55A38A86675790DB1AB00F0212683AD2" httpUri="http://treatment.plazi.org/id/55A38A86675790DB1AB00F0212683AD2" lastPageNumber="1" pageId="0" pageNumber="1" scope_family="Melastomataceae">
<subSubSection id="9BDF992B3B11806D5C2AABC2541AEDE7" pageId="0" pageNumber="1" type="nomenclature">
<paragraph id="23D796582248465E945D737DB3230D62" pageId="0" pageNumber="1">
26.
<taxonomicName id="065A23603CD8A2018779C5885677BB1D" ID-CoL="8R8LR" LSID="5CCAF9A9-9801-5E2F-8BF8-AAC474483A61" authority="(Bakh. f.) Karton." authorityName="Kartonegoro &amp; Veldkamp &amp; Hovenkamp &amp; Welzen" authorityYear="2018" baseAuthorityName="Bakh. f." class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta horrida" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="horrida" status="comb. nov.">Dissochaeta horrida (Bakh.f.) Karton.</taxonomicName>
<taxonomicNameLabel id="622960DE5F92FBE23ADD2CE21820659C" pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
<figureCitation id="806D4FFC452904CC85516401B49964A3" captionStart="Figure 18" captionStartId="F35" captionText="Figure 18. Dissochaeta horrida a habit b branchlet c hypanthium d fruits. Photographs by A. Kartonegoro; vouchers: Kartonegoro 1073 (BO, L)." figureDoi="10.3897/phytokeys.107.26548.figure18" httpUri="https://binary.pensoft.net/fig/222129" pageId="0" pageNumber="1">Fig. 18</figureCitation>
<figureCitation id="90FB2B2115EB73A8B14D4EBC35AB000B" captionStart="Map 17" captionStartId="F34" captionText="Map 17. Distribution of D. hirsutoidea (●), D. horrida (■) and D. intermedia (▲)." figureDoi="10.3897/phytokeys.107.26548.map17" httpUri="https://binary.pensoft.net/fig/222128" pageId="0" pageNumber="1">, Map 17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="4AA7DB95A80FDFD181FFDE1EF25217E0" pageId="0" pageNumber="1" type="reference_group">
<paragraph id="5A6BD4D88C5B040688DDFF1420F1E679" pageId="0" pageNumber="1">
<taxonomicName id="02C8637DC03942367F567821242CAF38" authority="Korth." authorityName="J. F. Maxwell" authorityYear="1980" baseAuthorityName="Bakh. fil." class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" infraspecific-rank="var." kingdom="Plantae" lsidName="Dissochaeta rostrata var. horrida" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="variety" species="rostrata" variety="horrida">Dissochaeta rostrata Korth. var. horrida</taxonomicName>
(Bakh.
<emphasis id="0286321C6DAB9016F3902BB8A5E3A155" italics="true" pageId="0" pageNumber="1">f.</emphasis>
) J.F.Maxwell, Gard. Bull. Sing. 33: 320. 1980.
</paragraph>
</subSubSection>
<subSubSection id="8E657FEC24F4980953C61FAA1E4282E5" pageId="0" pageNumber="1" type="basionym">
<paragraph id="25A1E7EC808F282F5BDBCAB12879AD5B" pageId="0" pageNumber="1">Basionym.</paragraph>
<paragraph id="425734A658B8F8554189A911A633B869" pageId="0" pageNumber="1">
<taxonomicName id="57847A4C586890147182705893CC7FF0" authorityName="Bakh fil" authorityYear="1943" class="Magnoliopsida" family="Melastomataceae" genus="Macrolenes" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Macrolenes horrida" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="horrida">
<emphasis id="8ADE1C82ED82DE57813DA99EC64BCCB5" italics="true" pageId="0" pageNumber="1">Macrolenes horrida</emphasis>
</taxonomicName>
Bakh.
<emphasis id="FFC4E30DFE47BDEB19FE986FF42E5FC0" italics="true" pageId="0" pageNumber="1">f.</emphasis>
, Contr. Melastom.: 208. 1943.
</paragraph>
</subSubSection>
<subSubSection id="1CDCDE20FA3DFA834EF9A61794E486E1" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="3AAC78B8CFEC1ED04ED2BFF22C116577" pageId="0" pageNumber="1">Type.</paragraph>
<paragraph id="F81D047382B1BBCF49357901C2AAE296" pageId="0" pageNumber="1">
Indonesia. West Sumatra: Agam, Brani, 850 m elev., 22 Jun 1918, H.A.B.
<normalizedToken id="693C072BC6C4F99EB274473724BED5B4" originalValue="Bünnemeijer">Buennemeijer</normalizedToken>
3200 (holotype: L [L0537276]!; isotypes: BO [BO1751324, BO1751325]!).
</paragraph>
<caption id="ED8D7360874AA5AD05EFCB53DDE64807" ID-Zenodo-Dep="1346455" doi="10.3897/phytokeys.107.26548.figure18" httpUri="https://binary.pensoft.net/fig/222129" pageId="0" pageNumber="1" start="Figure 18" startId="F35">
<paragraph id="B2FADF78C7F562969F006F0628CD0BD7" pageId="0" pageNumber="1">
<emphasis id="14E898B5B18967859E1D945C87EBC440" bold="true" pageId="0" pageNumber="1">Figure 18.</emphasis>
<taxonomicName id="CE0F8ECDD74A68F6580FBEE47D1A3CA7" class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta horrida" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="horrida">
<emphasis id="DE1F7E3DC7069E6FC29EB8F442AB42F2" italics="true" pageId="0" pageNumber="1">Dissochaeta horrida</emphasis>
</taxonomicName>
<emphasis id="A3264371B8D90727253A7C4051C05F81" bold="true" pageId="0" pageNumber="1">a</emphasis>
habit
<emphasis id="50B3FDE3FEDF42AC1D46A9C6F4CC34D2" bold="true" pageId="0" pageNumber="1">b</emphasis>
branchlet
<emphasis id="E35F79737F2A2FC772EC7E9CAF8B48C1" bold="true" pageId="0" pageNumber="1">c</emphasis>
hypanthium
<emphasis id="8AADF598185962012A8DAFFD687391FC" bold="true" pageId="0" pageNumber="1">d</emphasis>
fruits. Photographs by A. Kartonegoro; vouchers: Kartonegoro 1073 (BO, L).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="DFF82F10A51204382D5DE84825CE29FA" pageId="0" pageNumber="1" type="description">
<paragraph id="677E52622983D4EF2D5D9F72D01365F8" pageId="0" pageNumber="1">Description.</paragraph>
<paragraph id="BAE5E8092B06EAA69DD34BF4DF8F2D5B" pageId="0" pageNumber="1">
Climbing up to 2 m in height. Branchlets terete, 4-5 mm in diameter, densely covered with brown-reddish pubescent hairs and with 4-5 mm long brown-reddish bristle hairs, young parts densely covered with red pubescent hairs and bristle hairs; nodes swollen, with interpetiolar ridge; internodes 7-8 cm long. Leaves: petioles terete, 10-15 mm long, densely covered with brown-reddish pubescent hairs and with 4-5 mm long brown-reddish bristle hairs; blades broadly ovate to suborbicular, 13-18
<normalizedToken id="E16182012D286E4B35B83F95495C4C3B" originalValue="×">x</normalizedToken>
6-10 cm, subcoriaceous, base emarginate, margin entire, apex acuminate, tip 0.5-1 cm long; nervation with 2 pairs of lateral nerves and 1 pair of intramarginal nerves; adaxially glabrous, dark green, abaxially densely brown-reddish pubescent. Inflorescences terminal, up to 60 cm long, many-flowered; main axis angular, densely covered with brown-reddish pubescent hairs and with 4-5 mm long brown bristle hairs; primary axes up to 34 cm long with 6 or 7 nodes, secondary axes up to 6 cm long with 2 or 3 nodes, tertiary axes up to 2 cm long with 1 or 2 nodes, quarternary axes when developed up to 1 cm long with 1 node; bracts linear, 7-9 mm long, densely covered with brown-reddish pubescent hairs and with 4-5 mm long brown-reddish bristle hairs; bracteoles subulate, ca. 2 mm long, densely covered with brown-reddish pubescent hairs and with 4-5 mm long brown-reddish bristle hairs; pedicels densely covered with brown pubescent hairs and with glandular tip bristle hairs, 6-7 mm long in central flowers, 2-3 mm long in lateral flowers. Hypanthium tubular, 7-8
<normalizedToken id="8B1AFF27C1C4D11E9CF30733D3BC6DE0" originalValue="×">x</normalizedToken>
ca. 3 mm, densely covered with brown-reddish pubescent hairs and with brown-reddish glandular-tipped bristle hairs; calyx lobes linear-lanceolate, 6-10 mm long, base truncate, apex acute, densely covered with brown-reddish pubescent hairs and with glandular-tipped bristle hairs; petal bud conical, 3-4 mm long, glabrous; mature petals oblong to suborbicular, 6-8
<normalizedToken id="81EE990FBB379236E752F5BDCB349483" originalValue="×">x</normalizedToken>
5-6 mm, purple. Stamens 8, subequal, filaments curved sideways; alternipetalous stamens with 6-8 mm long filaments, anthers slender, curved, sickle-shaped, thecae 12-13 mm long, pedoconnective with 1-3 mm long, basal crest minute, triangular, ca. 0.5 mm long, lateral appendages paired, filiform, 1.5-2 mm long; oppositipetalous stamens with 5 mm long filaments, bent at the end, anthers curved, S-shaped, thecae 9-10 mm long, basal crest consisting of minute auricles, ca. 0.3 mm long, lateral appendages paired, auricles, 0.5-1 mm long. Ovary
<normalizedToken id="E7C6FB97ADAA225C06BF3EEFACEBF7FC" originalValue="¾">3/4</normalizedToken>
of hypanthium in length, apex villous; style 15-20 mm long, glabrous; stigma minute; extra-ovarial chambers 8, extending to the base of the ovary. Fruits urceolate, ca. 10
<normalizedToken id="99985C19116268EF9CFA37E5AE2C136D" originalValue="×">x</normalizedToken>
5-6 mm, densely covered with brown-reddish pubescent hairs and with glandular-tipped bristle hairs; calyx lobe remnants persistent, lanceolate, 6-8 mm long, reflexed. Seeds ca. 0.7 mm long.
</paragraph>
</subSubSection>
<subSubSection id="9B0E6F1D423E8776773B7F2994FEA0AE" pageId="0" pageNumber="1" type="distribution">
<paragraph id="FAFA358CC936C2089813392DC9240617" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="8A77D9DAD4D77B6A2CF15F19F04E4150" pageId="0" pageNumber="1">Sumatra (West).</paragraph>
</subSubSection>
<subSubSection id="7ED80B1EB9B1C264852981F83FA57389" pageId="0" pageNumber="1" type="ecology">
<paragraph id="46CD7C31991FE937A1098B74115AA84F" pageId="0" pageNumber="1">Ecology and habitat.</paragraph>
<paragraph id="16443F0DB3B5D2D806AB252EE4A3B455" pageId="0" pageNumber="1">Lowland dipterocarp forest or open cliff area at 500-850 m elevation.</paragraph>
</subSubSection>
<subSubSection id="7766FD687E073C97AA57EFCD9688AB3D" pageId="0" pageNumber="1" type="notes">
<paragraph id="4DA3C37DB4C70942F77ADEC7B08B9151" pageId="0" pageNumber="1">Note.</paragraph>
<paragraph id="F7D55205ED33D2B847334DD7EA1F2E32" pageId="0" pageNumber="1">
<taxonomicName id="7E752C725ED346ED5173A52839F82575" class="Magnoliopsida" family="Melastomataceae" genus="Dissochaeta" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Dissochaeta horrida" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="horrida">
<emphasis id="6A3BA6A14DDFC997EF1CDC19C7507537" italics="true" pageId="0" pageNumber="1">Dissochaeta horrida</emphasis>
</taxonomicName>
is easy to recognise because most parts have reddish brown pubescent hairs and dense bristle hairs except the green, glabrous adaxially surface of the leaf blades. The shape of the calyx lobes (linear-lanceolate, 6-10 mm long) resembles
<taxonomicName id="328E81DD5F14734650DC304E814090BF" class="Magnoliopsida" family="Melastomataceae" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="D. floccosa" order="Myrtales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="floccosa">
<emphasis id="64864B4F0E8A5759FC8C6B40850199E8" italics="true" pageId="0" pageNumber="1">D. floccosa</emphasis>
</taxonomicName>
, also from Sumatra, but the indumentum of the latter is floccose rather than pubescent.
</paragraph>
</subSubSection>
<subSubSection id="531671944BDDE98E4E56843214CA1C4A" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="966B39EAC6022ED29C9DB0712A389C2B" pageId="0" pageNumber="1">Specimens examined.</paragraph>
<paragraph id="475C05AB3A39B8BCFC20A70A2B99B1EC" pageId="0" pageNumber="1">
<emphasis id="03336E52D249079B391DEB650E256171" bold="true" pageId="0" pageNumber="1">INDONESIA. Riau</emphasis>
: Indragiri, Taluk, 11 Jan 1956, W. Meijer 4271 (L).
<emphasis id="085192764ABFC7BC7B4EB39B7604DE1B" bold="true" pageId="0" pageNumber="1">West Sumatra</emphasis>
: Agam, Brani, 850 m, 22 Jun 1918, H.A.B.
<normalizedToken id="2C9F2A895D1A1ABCEE85B172A25B096C" originalValue="Bünnemeijer">Buennemeijer</normalizedToken>
3200 (BO, L); Lima Puluh Kota, Harau Valley, Sarasah Bonta, 500 m, 2 May 2001, Uce et al. 106 (ANDA);
<emphasis id="414C4784D1F29EE73F5D21F24CEA9024" italics="true" pageId="0" pageNumber="1">Ibid.</emphasis>
, 11 Sep 2017, A. Kartonegoro 1073 (BO, L).
</paragraph>
</subSubSection>
</treatment>
</document>
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