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<document ID-CLB-Dataset="229647" ID-DOI="http://dx.doi.org/10.3897/asp.81.e81961" ID-GBIF-Dataset="6ba6d2ca-5dad-4270-914f-1042327d503c" ID-Pensoft-Pub="1864-8312-81-79" ID-Pensoft-UUID="CA16EA0C5A33560FBC955981DFC9D502" ID-ZooBank="5C1FDC828F9F4869ADDD83FA96E507ED" ModsDocID="1864-8312-81-79" checkinTime="1674309819629" checkinUser="pensoft" docAuthor="Domahovski, Alexandre Cruz &amp; Cavichioli, Rodney Ramiro" docDate="2023" docId="55975EEF91FB5DAE81D55462BA44C00E" docLanguage="en" docName="ArthropodSystPhyl 81: 79-164" docOrigin="Arthropod Systematics &amp; amp; Phylogeny 81" docPubDate="2023-01-20" docSource="http://dx.doi.org/10.3897/asp.81.e81961" docTitle="Coarctana parvula Domahovski &amp; Cavichioli 2023, sp. nov." docType="treatment" docUuid="28F7FF73-3770-449C-90E5-AF6D321B0348" docUuidSource="ZooBank" docVersion="5" id="CA16EA0C5A33560FBC955981DFC9D502" lastPageNumber="79" masterDocId="CA16EA0C5A33560FBC955981DFC9D502" masterDocTitle="Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data" masterLastPageNumber="164" masterPageNumber="79" pageNumber="79" updateTime="1732669275789" updateUser="ExternalLinkService">
<mods:mods id="B9DEE6CC0EC5F9387A898B7513A8094D" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="ACB97F2E7749A310400A892AE64ABF92">Phylogenetic analysis and revision of the leafhopper genus Acuera DeLong &amp; Freytag (Hemiptera: Cicadellidae: Gyponini) based on morphological data</mods:title>
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<mods:name id="C895375F7782D70AFE9525AD917B126D" type="personal">
<mods:role id="E5FC55D9DDDED8F3930040B0CE8D9B0B">
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<mods:namePart id="B09C79F02298FE787757D7AD90163837">Domahovski, Alexandre Cruz</mods:namePart>
<mods:nameIdentifier id="E9F5503ACB597BD3E6C834249ABD9A7F" type="ORCID">https://orcid.org/0000-0003-4588-4236</mods:nameIdentifier>
<mods:affiliation id="5B865D3C51DD0123DFF7B46947851CD2">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
<mods:nameIdentifier id="AA7DC928E00E96E77F89569ACACD9E82" type="email">domahovskiac@yahoo.com.br</mods:nameIdentifier>
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<mods:namePart id="89AAF1216402E9902319752B8E9203BA">Cavichioli, Rodney Ramiro</mods:namePart>
<mods:nameIdentifier id="FB9F13D59938CFA8C1AA6F29E5555C74" type="ORCID">https://orcid.org/0000-0001-5996-0629</mods:nameIdentifier>
<mods:affiliation id="2F39416D4248F07BD30E551A719B45F7">Departamento de Zoologia, Setor de Ciencias Biologicas, Universidade Federal do Parana, Av. Cel. Francisco H. dos Santos, 100, Caixa Postal 19020, 81531 - 980, Curitiba, PR, Brasil</mods:affiliation>
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<mods:title id="3EFCE4E1A5CF97E1123CB99D657C84A4">Arthropod Systematics &amp; amp; Phylogeny</mods:title>
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<mods:part id="69253B9C2C1DF5E5AA87C2D593C20586">
<mods:date id="C21D0B0A31D606781874A39BBBEDD3E2">2023</mods:date>
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<mods:number id="D3D20F57CC07223623BBDF5BD1CED834">2023-01-20</mods:number>
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<treatment id="55975EEF91FB5DAE81D55462BA44C00E" ID-GBIF-Taxon="209013705" LSID="urn:lsid:zoobank.org:act:28F7FF73-3770-449C-90E5-AF6D321B0348" httpUri="http://treatment.plazi.org/id/55975EEF91FB5DAE81D55462BA44C00E" lastPageNumber="79" pageId="0" pageNumber="79" scope_family="Cicadellidae" scope_order="Hemiptera" scope_tribe="Gyponini">
<subSubSection id="370D8EEA9087D7C4E6B34D06ADE57429" pageId="0" pageNumber="79" type="nomenclature">
<paragraph id="5CE67F9252F4709FBB06C097620212E5" pageId="0" pageNumber="79">
<taxonomicName id="C788A9B636422EB4EB92CFFBD52F0BD9" LSID="https://zoobank.org/28F7FF73-3770-449C-90E5-AF6D321B0348" authority="Domahovski &amp; Cavichioli, 2023" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana parvula" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="parvula" status="sp. nov.">Coarctana parvula</taxonomicName>
<taxonomicNameLabel id="A5FDF69EE40EDF49B29264C9AD14970B" pageId="0" pageNumber="79">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="033F22DA0302FC2B3DAAB937B5E0CB01" pageId="0" pageNumber="79" type="description">
<paragraph id="FBEE739B3C1447B81F0FDB147F67B44A" pageId="0" pageNumber="79">
<figureCitation id="D93BE7745281C9BED8F276100173C9EF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">Figs 25</figureCitation>
<figureCitation id="9E9CC41892F2225E8738A9B2D88CFFC4" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">, 31M, N</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="F1D8556B1527AC593CF7434E8BD3665C" pageId="0" pageNumber="79" type="diagnosis">
<paragraph id="181A439E45A8FF71D359807882EEB16F" pageId="0" pageNumber="79">Diagnosis.</paragraph>
<paragraph id="91CB6783F46AF41609AB7B9F92F5B99C" pageId="0" pageNumber="79">
Pygofer (Fig.
<figureCitation id="CAE81C337DEB10F67DA7D91AC0787BE0" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25G</figureCitation>
) strongly tapered apically, subacute. Subgenital plate (Fig.
<figureCitation id="B3C49C9FCB2236DA6D29CDCC2592A1FF" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25H</figureCitation>
) with filiform setae near outer margin. Aedeagus (Fig.
<figureCitation id="5CD21DB5B5BFFDF806415F9490A9CFB5" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25L, M</figureCitation>
) apodemal process slender and tapered, external surface bearing a small process near mid-length; shaft with pair of subapical processes short, directed dorsally and pair of apical processes curved ventrally, with half length of the subapical processes.
</paragraph>
<caption id="751F8C022C1F86405E777F36317AD259" doi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79" start="Figure 25" startId="F25">
<paragraph id="869F0EDEBE8DB31604B9CD0A94E6DFA3" pageId="0" pageNumber="79">
<emphasis id="3BEB955997FD47E4CA55C54EE94070F6" bold="true" pageId="0" pageNumber="79">Figure 25.</emphasis>
<taxonomicName id="DAA7012A395BEA97A4898C35D9964FEC" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana parvula" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="271412BAA8F5048BFF5FDFA020518C37" italics="true" pageId="0" pageNumber="79">Coarctana parvula</emphasis>
</taxonomicName>
<emphasis id="F182CF0ABDF0CEBCFB256765A60AD735" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
, holotype male.
<emphasis id="F0B304CA5495359AE7EE0220BDDEC444" bold="true" pageId="0" pageNumber="79">A</emphasis>
head and thorax, dorsal view.
<emphasis id="72029C0C61FF180C98B0F99D6C8AB325" bold="true" pageId="0" pageNumber="79">B</emphasis>
head, ventral view.
<emphasis id="576648D1D0285A0B01A7F8E3DB648BD1" bold="true" pageId="0" pageNumber="79">C</emphasis>
head and thorax, lateral view.
<emphasis id="AAB7B99B58212B4946A48EDDEFCC2BD3" bold="true" pageId="0" pageNumber="79">D</emphasis>
forewing.
<emphasis id="D8B253FCCB65ADF0B18A27A27C50F4AB" bold="true" pageId="0" pageNumber="79">E</emphasis>
sternite VIII, ventral view.
<emphasis id="FDE5A96971BE4067D7BEC9F922861D46" bold="true" pageId="0" pageNumber="79">F</emphasis>
valve, ventral view.
<emphasis id="B629578EB50D7DF7A525E812220AEF05" bold="true" pageId="0" pageNumber="79">G</emphasis>
pygofer, valve and subgenital plate, lateral view.
<emphasis id="AD4B58D66C91DD8C75A5B7ACAD58DE28" bold="true" pageId="0" pageNumber="79">H</emphasis>
subgenital plate, ventral view.
<emphasis id="85F702EFF96AD0E7E02B1322016ADC99" bold="true" pageId="0" pageNumber="79">I</emphasis>
connective, dorsal view.
<emphasis id="8A2612750E8269FB58BBDA2169595D73" bold="true" pageId="0" pageNumber="79">J</emphasis>
style, dorsal view.
<emphasis id="FE8D2C066E20D75B591525F08B663EC1" bold="true" pageId="0" pageNumber="79">K</emphasis>
style, lateral view.
<emphasis id="E0CF2C751FB87BE051D26A97D9283644" bold="true" pageId="0" pageNumber="79">L</emphasis>
aedeagus, lateral view.
<emphasis id="4E7A7AE1A0CE00F1402CCB51C0C76D4D" bold="true" pageId="0" pageNumber="79">M</emphasis>
aedeagus, posterior view. Scale bars in mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="D10550D15BFA861E66BE7E0B4983AEED" pageId="0" pageNumber="79" type="description">
<paragraph id="516295B54166FBED51ED16EA19E3EB0F" pageId="0" pageNumber="79">Measurements.</paragraph>
<paragraph id="83D37C5AB44DCB0C02D5FA05825FD839" pageId="0" pageNumber="79">Total length: holotype male 8.8 mm; paratypes, males (n = 5) 8.5-8.8 mm.</paragraph>
</subSubSection>
<subSubSection id="33198A3421562450D5DBFEE40B532D12" pageId="0" pageNumber="79" type="description">
<paragraph id="F023119579DAB9257C4D866A318941DE" pageId="0" pageNumber="79">Description.</paragraph>
<paragraph id="51E3712E806827642AABE12C70B8E98F" pageId="0" pageNumber="79">
Head, in frontal view (Fig.
<figureCitation id="1EEA9C6A4B1616AFEF7DE97177C1F92A" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25B</figureCitation>
), frons 1.1
<normalizedToken id="C8F5F67EEEE509A8450A196C15D4F448" originalValue="×">x</normalizedToken>
wider than long; frontogenal suture distant from eye margin by approximately the width of scape; clypeus very wide, 1.1
<normalizedToken id="3027D04B6F7FBBB528DD72CC9495C15D" originalValue="×">x</normalizedToken>
wider than long; lateral margins straight and parallel near base and apically strongly convergent; apex straight. Head, in lateral view (Fig.
<figureCitation id="70DB898FA073E0BD00DB214F667C438D" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25C</figureCitation>
), crown-face transition thin, with 3 transverse carinae; clypeus slightly inflated. Profemur AV row with 5-6 setae; PV row with 2 setae. Protibia PD row with 5 setae and undifferentiated intercalary setae; PV row with 5 setae and undifferentiated intercalary setae. Metatibia PD, AD and AV rows with 24-26, 13 and 13-15 macrosetae respectively. Metatarsomere I elongated 4
<normalizedToken id="9713E9B37110577F40ADC73B590D003D" originalValue="×">x</normalizedToken>
longer than apical width; inner row of the ventral surface with 7-8 setae. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection id="200223A9DC30A151B1047AA34167C6E1" pageId="0" pageNumber="79" type="coloration">
<paragraph id="8B7747AED8C6B5A9A4AC2BE05B97683E" pageId="0" pageNumber="79">Coloration.</paragraph>
<paragraph id="20A52565F3BE06A7D7D999EC539EE150" pageId="0" pageNumber="79">
Head and thorax (Figs
<figureCitation id="DB3415F1A259F418F87F6C7E64617305" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25A, B</figureCitation>
,
<figureCitation id="54889B1E34B75ABBA03357CB37008790" captionStart="Figure 31" captionStartId="F31" captionText="Figure 31. Species of Coarctana gen. nov., dorsal (left) and lateral (right) views. A, B C. apena comb. nov., holotype male, AMNH. C, D C. asymmetrica sp. nov., holotype male, DZUP. E, F C. glabra sp. nov., holotype male, DZRJ. G, H C. nigromedia sp. nov., holotype male, DZUP. I, J C. occultata sp. nov., holotype male, DZUP. K, L C. oricula sp. nov., holotype male, DZUP. M, N C. parvula sp. nov., holotype male, DZUP." figureDoi="10.3897/asp.81.e81961.figure31" httpUri="https://binary.pensoft.net/fig/798184" pageId="0" pageNumber="79">31M, N</figureCitation>
) reddish-brown. Head (Fig.
<figureCitation id="E06CC3BF55C8F9FA00960364175CAD3F" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25A</figureCitation>
) with pair of small circular black spots, near posterior margin, behind ocellus. Face (Fig.
<figureCitation id="A29EA2C38EA5717FA863BB8E68CA3B05" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25B</figureCitation>
) yellowish without maculae. Pronotum (Fig.
<figureCitation id="7F42DFD0940B2A0B213747907D097EAE" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25A</figureCitation>
) with black punctures; pair of circular black spots, near anterior margin, behind eye; proepimeron (Fig.
<figureCitation id="467EA9886D27C038488354A27B938C0A" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25C</figureCitation>
), with narrow black band below pronotal carina. Forewing (Fig.
<figureCitation id="81C17AC992F3F31870376284DBC3A3A8" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25D</figureCitation>
) with black maculae on apex of anal veins, cross veins of discal and apical cells. Metatibia (Fig.
<figureCitation id="D771358CFFB1ABD5861B03F91089AED3" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25C</figureCitation>
) with cucullate bases of setae blacks.
</paragraph>
</subSubSection>
<subSubSection id="10130F30C66F6EB8BAE8929068947DF1" pageId="0" pageNumber="79" type="male terminalia">
<paragraph id="6618AB59328B694964962A87D22A4D82" pageId="0" pageNumber="79">Male terminalia.</paragraph>
<paragraph id="EFE58BB29F945C722167E0B8261B954A" pageId="0" pageNumber="79">
Sternite VIII (Fig.
<figureCitation id="A1C6AC333221D4C793ACA9CCD9884C75" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25E</figureCitation>
) 1.5
<normalizedToken id="CAC99BD222816A52C85A7035F48C8DD5" originalValue="×">x</normalizedToken>
wider than long; lateral margins parallel; posterior margin slightly rounded. Valve (Fig.
<figureCitation id="9BE7BCBF5A627E636854572E7A6A9CB8" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25F</figureCitation>
) 1.5
<normalizedToken id="5F00362066592112C525BA42092F9622" originalValue="×">x</normalizedToken>
wider than long; posterior margin tapered and rounded. Pygofer, in lateral view (Fig.
<figureCitation id="3671EA3C918BE503BCA86379270D75C2" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25G</figureCitation>
), 2
<normalizedToken id="66E06E2FDC3B31B37C7954F7F8D1C8DC" originalValue="×">x</normalizedToken>
longer than maximum height; basodorsal process thin and long, extending ventrally to almost half height of lateral lobe; anteroventral margin rounded; posterodorsal margin straight; posteroventral margin excavated; apex strongly tapered, subacute. Subgenital plate, in lateral view (Fig.
<figureCitation id="84779276135A9C4C61E9055A44F2917B" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25G</figureCitation>
), short, not reaching pygofer apex; in ventral view (Fig.
<figureCitation id="D44F8F6E6D609F5BEF90252217C740E9" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25H</figureCitation>
), elongated, 3.7
<normalizedToken id="534FA2031343BA98A347AD220C23A8B9" originalValue="×">x</normalizedToken>
longer than wide, maximum width near half length; dorsal surface with filiform setae near outer margin; lateral margins slightly rounded; apex rounded. Connective (Fig.
<figureCitation id="0A6FFFEC9DC9B21B0F795E7D7D6DC8B9" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25I</figureCitation>
) wider than long; anterior margin moderately excavated; dorsal keel and stem reduced. Style, in dorsal view (Fig.
<figureCitation id="9210946B23C7B1F8F1362D0C00352BBC" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25J</figureCitation>
), with outer lobe small and truncated; in lateral view (Fig.
<figureCitation id="050D70C0D3E40EE85C4FBFE1A4CE00E1" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25K</figureCitation>
), with blade slightly wider near base and tapered toward apex; dorsal and ventral margins slightly sinuate; ventral margin with apical third serrated; apex truncated, forming a small dorsal process. Aedeagus (Fig.
<figureCitation id="9AC7A4157AD518E86B2DB50855945977" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25L, M</figureCitation>
) preatrium not developed; dorsal apodeme with dorsal margin slightly excavated, lateral margins produced ventrolaterally; apodemal process slender and tapered toward apex, curved in right angle near base, external surface bearing small process near mid-length; apex acute, curved anterad; shaft cylindrical, very long and thin, strongly curved dorsally near base and slightly curved anterad at apical third; apex with pair of subapical processes short, directed dorsally and pair of apical processes curved ventrally, with half length of the subapical processes. Other characteristics as in generic description.
</paragraph>
</subSubSection>
<subSubSection id="E411120659A0696392F52BA79131CBBD" pageId="0" pageNumber="79" type="female">
<paragraph id="5869690D4E2473223E9B30660EBD6D10" pageId="0" pageNumber="79">Female.</paragraph>
<paragraph id="BEEF6B870B532EB6EDDB851C5DD9E63B" pageId="0" pageNumber="79">Unknown.</paragraph>
</subSubSection>
<subSubSection id="8BFB61F51CB7EC857AF929FA2575D3B9" pageId="0" pageNumber="79" type="materials_examined">
<paragraph id="E22D0652911290DFE467DBAD882752EB" pageId="0" pageNumber="79">Material examined.</paragraph>
<paragraph id="045F9990101EE1CCB9B2EF376154F496" pageId="0" pageNumber="79">
<materialsCitation id="76AFB1F4C802037C033D5B0B717AC879" ID-GBIF-Occurrence="4017211414" collectingDate="2012-08-23" collectorName="R. R. Cavichioli" country="Peru" elevation="985" latitude="-13.3550005" location="Estrada" longLatPrecision="21" longitude="-70.88944" municipality="Peru" specimenCount="1" stateProvince="Cuzco" typeStatus="Holotype">
<emphasis id="ABE11DA4CFDC19781562916B82994C2C" bold="true" pageId="0" pageNumber="79">
<typeStatus id="93ED9396D4EC57CBB436FF1EA084470E">Holotype</typeStatus>
male:
<collectingCountry id="A31CE530D43324C2A53EEB99A9344948" name="Peru">Peru</collectingCountry>
</emphasis>
: &quot;
<collectingMunicipality id="A68C1B90B855E435878DC4397F0DDD5E">Peru</collectingMunicipality>
,
<collectingRegion id="44069300F538891BF778A5D214970D80" country="Peru" name="Cusco">Cuzco</collectingRegion>
,
<location id="BB1F8DE72D1C60C4F80A7D0CC6300286" LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:BB1F8DE72D1C60C4F80A7D0CC6300286" country="Peru" latitude="-13.3550005" longLatPrecision="21" longitude="-70.88944" municipality="Peru" name="Estrada" stateProvince="Cuzco">Estrada</location>
30 C\
<locationDeviation id="7A17301019FF3C952DA30F511370EEDC" location="Quincemil">
<quantity id="AF0315D4EA8DB802757F95DD0AD48055" metricMagnitude="4" metricUnit="m" metricValue="2.6" unit="km" value="26.0">26km</quantity>
W Quincemil
</locationDeviation>
,
<collectingDate id="4795B54FA7BB2DA8E4494DF03B8562F0" value="2012-08-23">23.viii.2012</collectingDate>
, (light)\
<geoCoordinate id="DFE95C13C50337AFD7651D1E7D3AB29F" degrees="13" direction="south" minutes="21" orientation="latitude" precision="15" seconds="18" value="-13.3550005">13°21′18″S</geoCoordinate>
<geoCoordinate id="5E765FAF182104F62834E94540499435" degrees="70" direction="west" minutes="53" orientation="longitude" precision="15" seconds="22" value="-70.88944">70°53′22″W</geoCoordinate>
,\
<quantity id="6165D904021B91EE013E51122CCC86EA" metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">
<elevation id="9363257BEE25B62C535099DA5930EB9E" metricMagnitude="2" metricUnit="m" metricValue="9.85" unit="m" value="985.0">985m</elevation>
</quantity>
<collectorName id="074480E6CA0176BBD6270ED98A95F07B">R.R. Cavichioli</collectorName>
leg.&quot; (MUSM)
</materialsCitation>
.
<materialsCitation id="5AEE5EB02001D9EB5CDA7CF6211293EB" ID-GBIF-Occurrence="4017211329" collectingDate="2023-01-01" collectingDateMax="2023-12-31" collectingDateMin="2023-01-01" collectorName="R. R. Cavichioli" country="Peru" elevation="985" latitude="-13.3550005" location="Estrada" longLatPrecision="21" longitude="-70.88944" municipality="Peru" specimenCount="4" specimenCount-male="4" stateProvince="Cusco" typeStatus="Paratypes">
<emphasis id="C85897243183C5FBEFD16A1E46D16314" bold="true" pageId="0" pageNumber="79">
<typeStatus id="8B879D48A430D85B843185CE3851682A">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="6D0EB52070A07C736AF2010864584752" type="male">4♂</specimenCount>
, same data as holotype (DZUP)
</materialsCitation>
;
<materialsCitation id="9AEED217E175B29155D846CDE909AF6C" ID-GBIF-Occurrence="4017211377" collectingDate="2009-02-06" collectorName="ME Irwin, GA" country="Peru" county="Peru" elevation="1125" latitude="-5.689722" location="Taporoto Rd" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" specimenCount="1" specimenCount-male="1" stateProvince="Amazonas" typeStatus="paratype">
<specimenCount id="5D3E5317C0B07357EC699D812E657F61" type="male">1♂</specimenCount>
, &quot;
<collectingCounty id="5DB6A8D711C5AD96E4A5BC5BD5050AD3">Peru</collectingCounty>
: Dept.
<collectingRegion id="2982A89F895FEEC6D1A78E728D92EF57" country="Peru" name="Amazonas">Amazonas</collectingRegion>
,\ Distr.
<normalizedToken id="657C88144F4CDB36DE87C69937A58E2A" originalValue="Águas">Aguas</normalizedToken>
Verdes,\
<collectingMunicipality id="87B7F5578E6F2CA349D834B1613E9EA2">Bagua</collectingMunicipality>
\
<location id="A0B9D47CDA8F2938CF50A1AF908300F3" LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:A0B9D47CDA8F2938CF50A1AF908300F3" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Taporoto Rd" stateProvince="Amazonas">Taporoto Rd</location>
(5N)\ at
<location id="C19532E6EF3599AEA66C88961DCD0B5E" LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:C19532E6EF3599AEA66C88961DCD0B5E" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Km" stateProvince="Amazonas">Km</location>
403;
<location id="820782B2246A3D5B7B68690F2B3E76D7" LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:820782B2246A3D5B7B68690F2B3E76D7" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Malaise" stateProvince="Amazonas">Malaise</location>
\ 30.I-\
<collectingDate id="091B39F4C95B9355D58D89AA591ECD01" value="2009-02-06">6.II.2009</collectingDate>
;
<collectorName id="55552A7AE5E6D983F98085D0C32ABB78">ME Irwin, GA</collectorName>
\
<location id="720241BEFE040AF1B417F2F4D7CD8A32" LSID="urn:lsid:plazi:treatment:55975EEF91FB5DAE81D55462BA44C00E:720241BEFE040AF1B417F2F4D7CD8A32" country="Peru" county="Peru" latitude="-5.689722" longLatPrecision="21" longitude="-77.63694" municipality="Bagua" name="Amaya" stateProvince="Amazonas">Amaya</location>
,
<quantity id="727A95F6B20BFD3B67CA6812CBC46418" metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">
<elevation id="EE2154E49F79513C62C8340998774C7F" metricMagnitude="3" metricUnit="m" metricValue="1.125" unit="m" value="1125.0">1125m</elevation>
</quantity>
;\
<geoCoordinate id="EB8BFF17809B4D913CB5B0DC848063BD" degrees="05" direction="south" minutes="41" orientation="latitude" precision="15" seconds="23" value="-5.689722">05°41′23″S</geoCoordinate>
<geoCoordinate id="6C69C8FEFBE58D8A9A06B73835F9E237" degrees="77" direction="west" minutes="38" orientation="longitude" precision="15" seconds="13" value="-77.63694">77°38′13″W</geoCoordinate>
(DZRJ)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="1FDCBF6FFC5CF26F7B2CCFF8AF9B6004" pageId="0" pageNumber="79" type="etymology">
<paragraph id="6C1AB3920294626F88217DF0816210A7" pageId="0" pageNumber="79">Etymology.</paragraph>
<paragraph id="06A387FE33AC9796DFFBDAD0A3F7B9FA" pageId="0" pageNumber="79">The new species name refers to the aedeagus with apodemal processes bearing very small projections at the mid-length of external.</paragraph>
</subSubSection>
<subSubSection id="E7EDCA5E30C3F2E281BFB19076FEB5F8" pageId="0" pageNumber="79" type="remarks">
<paragraph id="7FDA4359B9E6A18491D7E6F65C2CB98B" pageId="0" pageNumber="79">Remarks.</paragraph>
<paragraph id="7FF649704268A4502CEFB95B0B270687" pageId="0" pageNumber="79">
<taxonomicName id="270B3B08CB404EB8F2A87DF9BF57E87B" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana parvula" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="parvula">
<emphasis id="A4DE1CA5ADA263AE3BD2C89C49D79C9C" italics="true" pageId="0" pageNumber="79">Coarctana parvula</emphasis>
</taxonomicName>
<emphasis id="F6D4254055F252065CCFD91818549280" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
(Fig.
<figureCitation id="E6D81E050AA6D9B5A16FF48E222FFA96" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Phylogenetic relationships of Acuera found in the implied weighting analysis for the two equally most parsimonious trees, K = 3.700 and 4.284, length = 514, consistency index = 0.362, retention index = 0.771. Symmetric resampling values (&lt;50) are marked next to branches; red stars represent the type species of the three subgenera of Acuera. Synapomorphies are marked with black circles, while homoplasies are marked with empty circles." figureDoi="10.3897/asp.81.e81961.figure7" httpUri="https://binary.pensoft.net/fig/798160" pageId="0" pageNumber="79">7</figureCitation>
) was strongly supported (SR = 92) as sister group to
<taxonomicName id="25C9CC2F7BF09331EE6B655DF3C7A2AE" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. apena" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="apena">
<emphasis id="7432DC5C23F6FEEE84482A7A82049AD3" italics="true" pageId="0" pageNumber="79">C. apena</emphasis>
</taxonomicName>
<emphasis id="123920392DE122D8BEEABE8693D56294" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
+
<taxonomicName id="39B2EC575778D065A60F277F8F2B0180" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. occultata" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="occultata">
<emphasis id="1A3D27032B7D475FFD85B6CBD343382F" italics="true" pageId="0" pageNumber="79">C. occultata</emphasis>
</taxonomicName>
<emphasis id="993BFE6D6481EC9E330E92B1A2679519" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
This clade was supported by three synapomorphies: 26:0, clypeus wider than long; 27:2, clypeus with lateral margins strongly convergent apically; 29:1, clypeus tumid in lateral view; and three homoplastic characters: 9:1, crown-face transition with three carinae; 31:1 absence of black irregular maculae near anterior margin of pronotum; and 94:1, pygofer with basodorsal processes strongly developed.
<taxonomicName id="4413EAB51ACEBE88B1F4AECFD000D7BE" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" genus="Coarctana" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Coarctana oricula" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="oricula">
<emphasis id="A2809BE309A806D3A5617DCC855761E1" italics="true" pageId="0" pageNumber="79">Coarctana oricula</emphasis>
</taxonomicName>
<emphasis id="965DCCB5B9496CA073BA52028B2995A3" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
can be differentiated from
<taxonomicName id="B3191549671AD57DC1E768715CF40551" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. apena" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="apena">
<emphasis id="1CC8F419D636ADC0873E5977502BEFF0" italics="true" pageId="0" pageNumber="79">C. apena</emphasis>
</taxonomicName>
<emphasis id="887B666AD030994EE02FF7525E19DCEA" bold="true" pageId="0" pageNumber="79">comb. nov.</emphasis>
and
<taxonomicName id="A7535F89F7412A5AC45DB7FCD3809B1B" authorityName="Domahovski &amp; Cavichioli" authorityYear="2023" class="Insecta" family="Cicadellidae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="C. occultata" order="Hemiptera" pageId="0" pageNumber="79" phylum="Arthropoda" rank="species" species="occultata">
<emphasis id="4E0738858C83EA7EB3BBC325D8A7E958" italics="true" pageId="0" pageNumber="79">C. occultata</emphasis>
</taxonomicName>
<emphasis id="A510F6F231FE3A3B0372E1BDEE4272C7" bold="true" pageId="0" pageNumber="79">sp. nov.</emphasis>
by the apodemal processes of aedeagus (Fig.
<figureCitation id="C608E938C948196FFD915B09A97C5E04" captionStart="Figure 25" captionStartId="F25" captionText="Figure 25. Coarctana parvula sp. nov., holotype male. A head and thorax, dorsal view. B head, ventral view. C head and thorax, lateral view. D forewing. E sternite VIII, ventral view. F valve, ventral view. G pygofer, valve and subgenital plate, lateral view. H subgenital plate, ventral view. I connective, dorsal view. J style, dorsal view. K style, lateral view. L aedeagus, lateral view. M aedeagus, posterior view. Scale bars in mm." figureDoi="10.3897/asp.81.e81961.figure25" httpUri="https://binary.pensoft.net/fig/798178" pageId="0" pageNumber="79">25L, M</figureCitation>
) with a small process near mid-length of external surface, and the shaft with a pair of subapical processes short, directed dorsally and a pair of apical processes curved ventrally.
</paragraph>
</subSubSection>
</treatment>
</document>
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