treatments-xml/data/55/6B/8B/556B8BDE855659439061281CD7CCB1FF.xml

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<mods:title id="B67C32E9088565BD86C89CC28EFE32FA">Molecular phylogenetic and morphological studies reveal increased species diversity in the millipede genus Skleroprotopus Attems, 1901 in China (Julida: Mongoliulidae)</mods:title>
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<mods:namePart id="F2CB6339AB5EB934C4AA271EF880A4C6">Chen, Rong</mods:namePart>
<mods:affiliation id="ECD7CE63E4FA9079E2F55CCEA5B86663">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, China</mods:affiliation>
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<mods:namePart id="47C0A444837E80B724BAF3617E6925F8">Zhao, Yi</mods:namePart>
<mods:affiliation id="F02A58C915D1F4C86F83F55AFAEFE76C">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, China &amp; Guangdong Academy of Forestry, 233 Guangshan 1 st Road, Guangzhou 510520, China</mods:affiliation>
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<mods:namePart id="BC2BDF2EFB2405EAF070987DA384DEA6">Golovatch, Sergei</mods:namePart>
<mods:affiliation id="65EDB7BA326EBA85FC42B9C9DCD6F505">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071, Russia</mods:affiliation>
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<mods:namePart id="F5CFC05975DDA087A835DA44B2FE0B75">Liu, Wei-Xin</mods:namePart>
<mods:affiliation id="DB8F24E37158A911E4CE40CD3E38E08B">College of Plant Protection, South China Agricultural University, 483 Wushan Road, Guangzhou 510642, China</mods:affiliation>
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<mods:date id="814A7068AFF7A7F871DF86D11EC1CBCB">2024</mods:date>
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<treatment id="556B8BDE855659439061281CD7CCB1FF" ID-DOI="http://doi.org/10.5281/zenodo.14513077" ID-Zenodo-Dep="14513077" ID-arpha="556B8BDE-8556-5943-9061-281CD7CCB1FF" ID-zoobank="https://zoobank.org/84BD5B7A-81AB-472D-8FF9-82A756B27920" LSID="urn:lsid:plazi:treatment:556B8BDE855659439061281CD7CCB1FF" LSID-ZBK="urn:lsid:zoobank.org:act:84BD5B7A-81AB-472D-8FF9-82A756B27920" httpUri="http://treatment.plazi.org/id/556B8BDE855659439061281CD7CCB1FF">
<subSubSection id="A16599E2A0DEE2C933E75D28ADBE6C07" type="nomenclature">
<paragraph id="6E28204B7C6CEABCCABEE775CA83D2DC">
<heading id="D3C04E1333ED2B4AC5AB642FDE6A64FF" reason="label">3.3. 8.</heading>
<taxonomicName id="EBE70E0133F9A0C2EF9967F6AC974A57" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="genjudi" status="sp. nov.">
<emphasis id="1A4FBD552A0D8E9A58DCFFDCF310A145" italics="true">Skleroprotopus genjudi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="55B410DACC2C63788B603312592AC2D7" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="2A9CA9A47B4B2A6316231E0A380B9C06" type="description">
<paragraph id="53DC7700C1D16092CB78140F6C2F5F8A">
<figureCitation id="ED820EF86FBDC015F8F66E2A884F8094" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov." figureDoi="10.3897/asp.82.e136751.figure2" httpUri="https://binary.pensoft.net/fig/1201329">Figures 2 D</figureCitation>
, 
<figureCitation id="F67F321B9F1961A4CD210A6DF4C28EEF" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="SECID0E6NBI" type="material">
<paragraph id="F7D04D6011966166EBA843A603B55070">
<heading id="57B9E17AAFB452C5183CD81B1AA90E61" reason="title">Material examined.</heading>
</paragraph>
<paragraph id="EB759FFCF19B8FAB70AAAB586B71CF47">
<materialsCitation id="72B12C2AD739028AF81BD418C6B1E2A0" collectingDate="2014-11-09" collectionCode="SCAU" country="China" county="Chongyi County" elevation="400" latitude="25.452509" location="Niedu Town" longLatPrecision="1" longitude="114.083694" municipality="Ganzhou City" specimenCount="1" stateProvince="Jiangxi" typeStatus="holotype">
<typeStatus id="159D4AC0E9F0BF0CD95A27876629ADDB" type="holotype">Holotype</typeStatus>
male (
<named-content id="7935F098F77B00E1D705F8AEB30D70F0" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="0CB3ECCDB7CDE18A358DB14921F23517">SCAU</collectionCode>
</named-content>
), 
<collectingCountry id="7EC6E79DABB20AF3D7223F859FBFCEEB" name="China">China</collectingCountry>
, 
<collectingRegion id="EC55A241D6902E98C3804AB3A32E96D7" country="China" name="Jiangxi">Jiangxi Province</collectingRegion>
, 
<collectingMunicipality id="D7EC2EA280F2A71310294E3B5E56E072">Ganzhou City</collectingMunicipality>
, 
<collectingCounty id="AFD6378CC4A952F5EE490BF7D02D05D2">Chongyi County</collectingCounty>
, 
<location id="6B25A6C138578FB38E9197D040B23C66" LSID="urn:lsid:plazi:treatment:556B8BDE855659439061281CD7CCB1FF:6B25A6C138578FB38E9197D040B23C66" country="China" county="Chongyi County" latitude="25.452509" longLatPrecision="1" longitude="114.083694" municipality="Ganzhou City" name="Niedu Town" stateProvince="Jiangxi">Niedu Town</location>
, 
<location id="52C05BE4DA7EB0BADB450B32E9CAA8FB" LSID="urn:lsid:plazi:treatment:556B8BDE855659439061281CD7CCB1FF:52C05BE4DA7EB0BADB450B32E9CAA8FB" country="China" county="Chongyi County" latitude="25.452509" longLatPrecision="1" longitude="114.083694" municipality="Ganzhou City" name="Cave Luohanyan" stateProvince="Jiangxi">Cave Luohanyan</location>
, 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMOBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[114.083689,25.452508]}">
<geoCoordinate id="340BC3D6F4C2BF306B85A13ABA591008" degrees="25" direction="north" minutes="27" orientation="latitude" precision="1" seconds="9.03" value="25.452509">25 ° 27 ′ 9.03 ′′ N</geoCoordinate>
<geoCoordinate id="6DFD3D278616CBAABEC27716EAE5326F" degrees="114" direction="east" minutes="05" orientation="longitude" precision="1" seconds="1.28" value="114.083694">114 ° 05 ′ 1.28 ′′ E</geoCoordinate>
</named-content>
, 
<quantity id="37DCFAE4F30DBD2A37223A56885155B1" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">
<elevation id="9C1398F409542DBC66D94D52EEE06722" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</elevation>
</quantity>
, 
<date id="8BF6A0722287EB407639F0E7FE14910A" value="2014-11-09">
<collectingDate id="330FD46B3ED294DFA013D45FC78952ED" value="2014-11-09">9. XI. 2014</collectingDate>
</date>
, fifteen graduate students from the class of 2013 leg
</materialsCitation>
. 
<typeStatus id="C6DD652F5B3D7C60EC173B920A503E20" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="951F046A7B18ADDB80BCF332AE39359A" count="28" type="male">28 males</specimenCount>
, 
<specimenCount id="DB70D7E9240BCA0606CC5831FB00010A" count="12" type="female">12 females</specimenCount>
(
<named-content id="6E7F5B7C602FBA9C2B287F7BD35FBE4D" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="9E14D7B05FC52DF4FD6E63AE79607357">SCAU</collectionCode>
</named-content>
), same data as for 
<typeStatus id="BFE5ABEF35F435FBA634DBA114BE55B3" type="holotype">holotype</typeStatus>
; 
<specimenCount id="BE72BD54320ABE61A086A434048B328F" count="6" type="male">6 males</specimenCount>
, 
<specimenCount id="AFDAED6160D6F03643BCD2C0227A720F" count="6" type="female">6 females</specimenCount>
(
<named-content id="82A79E3125BE340DDB9180B8805EE3D3" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="175247AD8488236CD539A434A573C6C6">SCAU</collectionCode>
</named-content>
), same cave, 
<date id="4F73B13C04A6A4B5F4559EAFD5CBD2C2" value="2011-01-12">12. I. 2011</date>
, Tian Mingyi, Liu Weixin, Zheng Yuan and Sun Feifei leg. Other material examined: 
<specimenCount id="85FFC32DFCCBE597AE3F35B7ACEF1057" count="4" type="male">4 males</specimenCount>
, 
<specimenCount id="76E675CA62F2FAAB36E7AAE6AD948772" count="5" type="female">5 females</specimenCount>
(
<named-content id="FC9B77D5012590B5097D3407280FC2F8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="B2698B7A9E0A62106E86D77DD08E882C">SCAU</collectionCode>
</named-content>
), same town, Cave Miao Dong, 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EAPBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[114.085214,25.467117]}">
<geoCoordinate id="A402CB6910FB5239C1B6A7BC6FD954C0" degrees="25" direction="north" minutes="28" orientation="latitude" precision="1" seconds="1.62" value="25.467117">25 ° 28 ′ 1.62 ′′ N</geoCoordinate>
<geoCoordinate id="97CA7D0CB8B421EE56447C8D0A519C0E" degrees="114" direction="east" minutes="05" orientation="longitude" precision="1" seconds="6.77" value="114.08521">114 ° 05 ′ 6.77 ′′ E</geoCoordinate>
</named-content>
, 
<quantity id="37F4F542183F9C727F6E8919E4D22A53" metricMagnitude="2" metricUnit="m" metricValue="3.5" unit="m" value="350.0">350 m</quantity>
, 
<date id="ACCB2EEAF8EB28CAD830571658CF78BC" value="2011-01-13">13. I. 2011</date>
, Tian Mingyi, Liu Weixin, Zheng Yuan and Sun Feifei leg.; 
<specimenCount id="6A0505E6729F67F725B001E7913F2F0C" count="3" type="male">3 males</specimenCount>
(
<named-content id="DE46E872F0FED3458CFB42D1AB9B84E8" content-type="dwc:institutional_code" xlink_href="http://grbio.org/institution/south-china-agricultural-university" xlink_title="Zoological Collection of the South China Agricultural University">
<collectionCode id="A8CD60FFFD0B108A19D4DCBCCC015162">SCAU</collectionCode>
</named-content>
), same town, Cave Lianhua Dong, 
<named-content content-type="dwc:verbatimCoordinates" id="NCID0EMPBI" specific-use="{&quot;type&quot;:&quot;Point&quot;,&quot;coordinates&quot;:[114.093611,25.469444]}">
<geoCoordinate id="63D7D5A7ADF12DD10CC5CACEAC6F67D7" degrees="25" direction="north" minutes="28" orientation="latitude" precision="15" seconds="10" value="25.469444">25 ° 28 ′ 10 ′′ N</geoCoordinate>
, 
<geoCoordinate id="EF787E5F8DD26E62F1CA060FE794B265" degrees="114" direction="east" minutes="05" orientation="longitude" precision="15" seconds="37" value="114.09361">114 ° 05 ′ 37 ′′ E</geoCoordinate>
</named-content>
, 
<quantity id="485A3291E8C8FA7D9ECA7B6D6DCDEAF1" metricMagnitude="2" metricUnit="m" metricValue="4.0" unit="m" value="400.0">400 m</quantity>
, 
<date id="50A0531B8BF71F536BC88FDA8433759D" value="2017-04-16">16. IV. 2017</date>
, fifteen graduate students from the class of 2016 leg.
</paragraph>
</subSubSection>
<subSubSection id="SECID0ERPBI" type="diagnosis">
<paragraph id="75A9E065D60F76FC4BDA9FD436C504C4">
<heading id="7604C28CA224E3D618A667B91224DA9F" reason="title">Diagnosis.</heading>
</paragraph>
<paragraph id="5891842291DC832E74F047FC8659AB87">
Differs from congeners mostly based on the following combination of characters: (1) penis much shorter than coxae 2 (Fig. 
<figureCitation id="4EB0091DC9B99DF485D4CEE5F4DD0967" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 E</figureCitation>
); (2) male leg 7 with an extremely long coxal process, strongly bent anteriad, telopodite rather small, 2 or 3 - segmented (Fig. 
<figureCitation id="AA441F3472375566198730E1B39ED1EE" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 F</figureCitation>
); (3) anterior gonopod (Fig. 
<figureCitation id="273B3F0DBF89FF6CD61807A67C3382DA" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 G, H</figureCitation>
) with a coxal process of medium length and carrying a much lower, irregularly dentate, membranous lobe, and a very long and slender flagellum, distally branched. — In addition, this new species differs from all other species analyzed in a&gt; 8.2 % uncorrected 
<emphasis id="823A6BA68B60D85AE378D3CA97380CF1" italics="true">p</emphasis>
- distance of the 
<abbrev id="ABBRID0EFQBI" xlink_title="Protein-coding gene sequences">COI</abbrev>
barcoding gene.
</paragraph>
<caption id="D7504693385D9F425EC1974E16EE7CA5" ID-DOI="10.3897/asp.82.e136751.figure11" ID-arpha="E8F1B761-AB4E-5666-B4FC-6FD3B938EA4C" httpUri="https://binary.pensoft.net/fig/1201338" startId="F11">
<paragraph id="A9E4F4849ABCDBE5DDD5D2724174F210">
<label id="FDFE3F2A7F937F35F7C2B6F1067ED817">Figure 11.</label>
</paragraph>
<paragraph id="69B21DEF041226F2E35274AA897C3AAB">
<taxonomicName id="A9B7C6EDD6778722DCF33CE27FF48570" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="genjudi" status="sp. nov.">
<emphasis id="083C422FD6DE398C19F1323C6BFA317B" italics="true">Skleroprotopus genjudi</emphasis>
</taxonomicName>
<emphasis id="594F3C52ADBBB126B63C8767EEF6C4EF" bold="true">
<taxonomicNameLabel id="889EF1AB0CF1B2CBC510B15052E6404E" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
. paratypes from Cave Luohanyan. 
<emphasis id="C5292505EA118681121C3E650A0D3BE0" bold="true">A</emphasis>
, 
<emphasis id="3881A948B77481D1BA31AA76F22A4ABA" bold="true">D</emphasis>
– 
<emphasis id="B55968BE378DDC880EFAE67BE66CF0E5" bold="true">I</emphasis>
male paratype; 
<emphasis id="9A8EAC0D20FECD49A17893E16BE51494" bold="true">B</emphasis>
, 
<emphasis id="B55956026307D3981BC03A20C149B699" bold="true">C</emphasis>
female paratype. 
<emphasis id="B376C14F6802669031D0D26CD40328DB" bold="true">A</emphasis>
, 
<emphasis id="6AA5BB3F269300A889299B5467EFC27E" bold="true">B</emphasis>
gnathochilarium; 
<emphasis id="F422CFFC512E1DA2D6E204A367187B91" bold="true">C</emphasis>
right vulva, mesal view; 
<emphasis id="23EB77D90EA2977FD8DD4CCFE1CD138F" bold="true">D</emphasis>
legs 1, anterior view; 
<emphasis id="749E5A104BEAA8A98F3E7E9825120D24" bold="true">E</emphasis>
legs 2 with penis, posterior view; 
<emphasis id="E48F703CB6E3E77AE05EDE27D2F7766C" bold="true">F</emphasis>
legs 7, anterior view; 
<emphasis id="2792B79002C2DF5C1E1D5C6CF8ADE050" bold="true">G</emphasis>
anterior gonopods, anterior view; 
<emphasis id="D71B8810D421EC49A8C8408EA05EF54E" bold="true">H</emphasis>
anterior gonopods, posterior view; 
<emphasis id="CC3F95C3E5C9120EECBB9613B660A579" bold="true">I</emphasis>
posterior gonopods, anterior view. 
<emphasis id="E618395058ED8179EEDE8237115615B4" bold="true">Abbreviations</emphasis>
: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. 
<emphasis id="D159B2C17CBF1589CFF197419154B426" bold="true">Scale bars</emphasis>
: 0.2 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="SECID0EDSBI" type="etymology">
<paragraph id="C68F696E10D5419821F60AC02CF701FA">
<heading id="786F0AF394DF910CBCD365ABFE05D71E" reason="title">Etymology.</heading>
</paragraph>
<paragraph id="EB470F24B7633BD59E17D3FE7063056A">The species name is primarily derived from its collection site, which is often used as a place for our graduate students to conduct internships on cave biodiversity. “ Genjudi ” here refers to the main operational base, in Chinese is “ 根据地 ”; a noun in apposition.</paragraph>
</subSubSection>
<subSubSection id="SECID0EISBI" type="description">
<paragraph id="9650269DAF691DFB1FD113F9F25CE250">
<heading id="0101B8A7A7960D19BC8C89B4798D83A8" reason="title">Description.</heading>
</paragraph>
<paragraph id="6EE106BAF5D21DA7965E767747A042C7">
Length of both sexes ca 30.0–42.0 mm, 
<quantity id="457AFEA2C1F20A765078890AEEB74A1A" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.1" metricValueMin="1.7" unit="mm" value="1.9" valueMax="2.1" valueMin="1.7">1.7–2.1 mm</quantity>
in diameter, body with 51–63 podous + 1–2 apodous rings + telson. Natural coloration (Fig. 
<figureCitation id="D9BDB0A2CECAA2D403F17D735A5362A2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Pictures of live animals, photos by Prof. Mingyi Tian from SCAU. A Skleroprotopus change sp. nov.; B Skleroprotopus megistus sp. nov.; C Skleroprotopus longissimus sp. nov.; D Skleroprotopus genjudi sp. nov.; E Skleroprotopus tiankeng sp. nov.; F Skleroprotopus laiyuanensis sp. nov.; G Skleroprotopus incisodentatus sp. nov." figureDoi="10.3897/asp.82.e136751.figure2" httpUri="https://binary.pensoft.net/fig/1201329">2 D</figureCitation>
) uniformly grayish yellow. Antennae and legs light yellow. Eye patches grey to blackish, subtriangular, arranged in 4–7 irregular rows, altogether about 30–40 ommatidia per eye patch.
</paragraph>
<paragraph id="EB347B43D6AD7BB1B96F71B30AC0FCB2">
Head capsule smooth and hairless, eyes slightly bulged, epicranial suture well-defined. Labral margin with 1 (2) + 1 (2) supra-labral and 11 + 11 labral setae. Antennae long and slender, reaching behind to middle of ring 4 when stretched dorsally. In length, antennomeres 3&gt; 2&gt; 4&gt; 5&gt; 6&gt; 1&gt; 7. Antennomeres 5 and 6 each with distal corolla of sensilla basiconica. Antennomere 7 with 4 sensory cones. Mandibular stipes enlarged, with two small lobes in males, rounded in females. Gnathochilarium with at least 7 setae on each lamella lingualis (
<emphasis id="4C6A31A9C2D48B1F331541AC9D5AE69E" bold="true">ll</emphasis>
); promentum (
<emphasis id="D3FFD45CBA4F0AFFF513A8B8BDF14416" bold="true">pr</emphasis>
) lance-shaped at base, swollen anteriorly, short and rod-like in males (Fig. 
<figureCitation id="7FBFFF8C492159B13DB76F6AE2194EA4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 A</figureCitation>
), vs a drop-shaped 
<emphasis id="FADECA87186F3B35693667C81256A49F" bold="true">pr</emphasis>
in females (Fig. 
<figureCitation id="1DF104EE4EF75BC453BE5F13475962D4" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 B</figureCitation>
).
</paragraph>
<paragraph id="0CE2E0805A1823A023AA21E08E687364">Collum with 4–6 lower striae laterally, but lowest 2 striae not reaching the anterior margin. Prozona with 2–6 irregular subtransverse striae and metazona with 16–24 longitudinal striae laterally. Suture dividing pro- and metazona very narrow, a regular comb. Ozopores starting with ring 6, lying mid-laterally on sides of metatergites.</paragraph>
<paragraph id="41084E6737DF30F36CA57CA01357DC18">Epiproct slightly protruding caudad, posterior margin with 3 + 3 setae. Paraprocts convex, with 1 + 1 setae. Hypoproct eye-shaped, with 1 + 1 setae.</paragraph>
<paragraph id="34C717F9D343A90407DECA1770747BF6">
Legs long and slender, about 1.2 × as long as midbody height. Male leg-pair 1 (Fig. 
<figureCitation id="B3647B0D8D02DE2F7854B89D749A972D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 D</figureCitation>
) hypertrophied, 5 - segmented, strongly curved anteriad; coxa (
<emphasis id="23EBF8982B8BE790FB247A9704012C87" bold="true">cx</emphasis>
) and telopoditomeres 1 and 2 (
<emphasis id="E40644947AAA67A9E74C559467E3B37A" bold="true">t 1</emphasis>
, 
<emphasis id="CCF3C4FB9E8FF3A9EBD89217D82CF623" bold="true">t 2</emphasis>
) with several long setae. Telopoditomere 3 (
<emphasis id="7A680F75AB004580943E3D51332E9A2D" bold="true">t 3</emphasis>
) longest, about as long as all other telopoditomeres combined. Telopoditomere 4 (
<emphasis id="E78FC3BDAC6FD648AD62EF97898FE36A" bold="true">t 4</emphasis>
) irregularly shaped, with two vague emarginations, one lateral, the other dorsal. Male leg-pair 2 (Fig. 
<figureCitation id="39594E5B5E52985770C7031ABC4CC32D" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 E</figureCitation>
) reduced in size and slender; penis (
<emphasis id="486854E522E6A91A33F7102DEA1555CF" bold="true">pn</emphasis>
) shorter than coxae (
<emphasis id="F256C18B6D7F939EF2FAC54EBB591242" bold="true">cx</emphasis>
), distal part with five long setae. Male leg-pair 7 (Fig. 
<figureCitation id="C50A4F1E9A9E5CC24B4EDC05512E2005" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 F</figureCitation>
) strongly modified, coxal process (
<emphasis id="A601E7BBB53D4B07BB52321BA1D3BAC0" bold="true">cp</emphasis>
) extremely long, flattened dorsoventrally, only slightly attenuated distad, strongly bent anteriad, densely setose nearly throughout at both mesal and lateral margins; telopodite (
<emphasis id="6B5B9245C04FE7190392221A238023D9" bold="true">te</emphasis>
) remains poorly visible laterally near base of 
<emphasis id="9A4B231490B0CAD5802D0284D7E36330" bold="true">cp</emphasis>
, vestigial, consisting of 2 or 3 very small telopoditomeres.
</paragraph>
<paragraph id="3921E45CBD00309FD5EB62D9422141CF">
Anterior gonopods (Fig. 
<figureCitation id="9B4A044B25478BDCA5CE8C7E4EE69281" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 G, H</figureCitation>
) with a coxal process (
<emphasis id="50F0412D5F7E1CAE784B2040FAEB718D" bold="true">cp</emphasis>
) of medium length, laterally bearing a long row of short villi, posteriorly carrying with a lower membranous lobe (
<emphasis id="38324FEE70A72CDBEE3AC5D860323AEC" bold="true">ml</emphasis>
), outer margin of 
<emphasis id="1B6A15EFD17F258306C6E43780371382" bold="true">ml</emphasis>
irregularly dentate, somewhat broadened and reflexed in proximal half. Flagellum (
<emphasis id="008A7FDA828B24122E62CA16DABA920C" bold="true">f</emphasis>
) very long and slender, distally serrate and branched. Telopodite (
<emphasis id="502E3CED6BA6E11109F788EB162F01E5" bold="true">te</emphasis>
) about 2 / 3 as long as 
<emphasis id="2F5EECD2EE595B70D95A59272BCB3F10" bold="true">cp</emphasis>
, with long setae both apically and mesally, laterobasally with a minute remnant (
<emphasis id="D869F023C589B9E142AB70E4BC499BA8" bold="true">r</emphasis>
) of a second podomere.
</paragraph>
<paragraph id="874A50C933DD4F5A171A237B279CA7E1">
Posterior gonopods (Fig. 
<figureCitation id="7836C7E9E44450A7ADBB2B008D77ADB8" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 I</figureCitation>
) erect, with bristles medially, distally branched, anterior branch smooth and slender, with a pointed tip, posterior one densely setose.
</paragraph>
<paragraph id="E183D8612500D889DC0F5A252A0EECD6">
Vulva (Fig. 
<figureCitation id="6906267E94D5FE49CD8115C24F46EFF2" captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Skleroprotopus genjudi sp. nov. . paratypes from Cave Luohanyan. A, D – I male paratype; B, C female paratype. A, B gnathochilarium; C right vulva, mesal view; D legs 1, anterior view; E legs 2 with penis, posterior view; F legs 7, anterior view; G anterior gonopods, anterior view; H anterior gonopods, posterior view; I posterior gonopods, anterior view. Abbreviations: cp, coxal process; cx, coxa; f, flagellum; ll, lamella lingualis; ml, membranous lobe; op, operculum; pn, penis; pr, promentum; r, remnant; t 1 – t 4, telopoditomeres 1–4; te, telopodite. Scale bars: 0.2 mm." figureDoi="10.3897/asp.82.e136751.figure11" httpUri="https://binary.pensoft.net/fig/1201338">11 C</figureCitation>
) as usual, with two parallel rows of short setae on both anterior and posterior surfaces, operculum (
<emphasis id="7417161385AEA3ED09F972173E2E812D" bold="true">op</emphasis>
) with several long setae distally.
</paragraph>
</subSubSection>
<subSubSection id="SECID0EIVBI" type="remarks">
<paragraph id="017926682F56F896D5E9C795819A0579">
<heading id="CA01661EB2F14E013841486569068A2A" reason="title">Remarks.</heading>
</paragraph>
<paragraph id="0FB297BFEEFFD71D8863CCBDADF4051F">
This new species was found in several caves at Niedu Town, Chouyi City, 
<collectingRegion id="D879C7DC4E45B699DB45E3F67C363D00" country="China" name="Jiangxi">Jiangxi</collectingRegion>
. Based on the unpigmented body, as well as long legs and antennae, 
<taxonomicName id="A6EA56C51309D5C2BC0C7A8E4F07D630" authorityName="Chen &amp; Zhao &amp; Golovatch &amp; Liu" authorityYear="2024" class="Diplopoda" family="Mongoliulidae" genus="Skleroprotopus" kingdom="Animalia" order="Julida" phylum="Arthropoda" rank="species" species="genjudi" status="sp. nov.">
<emphasis id="2C6EA59568A886F0AF30B45316C83A06" italics="true">S. genjudi</emphasis>
</taxonomicName>
<emphasis id="F4B55606E36BC520090522FF903D64B3" bold="true">
<taxonomicNameLabel id="13BBF60F6F39BB153DFB31B5474C4400" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is considered a troglobite.
</paragraph>
</subSubSection>
</treatment>
</document>