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<document id="128CAFB95191C7C5423A305C6C6F3FF7" ID-DOI="10.25221/fee.434.1" ID-ISSN="2713-2196" ID-Zenodo-Dep="7165976" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665252324355" checkinUser="felipe" docAuthor="Gorochov, A. V." docDate="2021" docId="5569A62BFFC90F7FFF04FDA8A6D9B3AA" docLanguage="en" docName="FarEastEntomol.434.1-25.pdf" docOrigin="Far Eastern Entomologist 434" docSource="http://dx.doi.org/10.25221/fee.434.1" docStyle="DocumentStyle:68641B8F4A21450A5F9A74164B730F50.2:FarEastEntomol.2017-.journal_article" docStyleId="68641B8F4A21450A5F9A74164B730F50" docStyleName="FarEastEntomol.2017-.journal_article" docStyleVersion="2" docTitle="Lipotactes (Lipotactes) alienus subsp. kubah Gorochov, 2021, subsp. n." docType="treatment" docUuid="AB6BD62C-7BA3-40B3-BA77-A94EDFCB46CC" docUuidSource="ZooBank" docVersion="3" lastPageNumber="8" masterDocId="A950DE53FFCF0F78FFE7FF81A749B23D" masterDocTitle="Taxonomy of the katydids (Orthoptera: Tettigoniidae) from East Asia and adjacent islands. Communication 14" masterLastPageNumber="25" masterPageNumber="1" pageNumber="7" updateTime="1699371420168" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="00244A7CCA2494B664A7F9098B4CD391">Taxonomy of the katydids (Orthoptera: Tettigoniidae) from East Asia and adjacent islands. Communication 14</mods:title>
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<mods:namePart id="C9B808CE30348269E08BBBF0146DE37B">Gorochov, A. V.</mods:namePart>
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<mods:date id="70EA9A3465062D51F0EFF4E44BDBF36F">2021</mods:date>
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<treatment id="5569A62BFFC90F7FFF04FDA8A6D9B3AA" LSID="urn:lsid:zoobank.org:act:AB6BD62C-7BA3-40B3-BA77-A94EDFCB46CC" httpUri="http://treatment.plazi.org/id/5569A62BFFC90F7FFF04FDA8A6D9B3AA" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
<subSubSection id="95DA44B6FFC90F7EFF04FDA8A30EB05D" pageId="6" pageNumber="7" type="nomenclature">
<paragraph id="DD7F173DFFC90F7EFF04FDA8A421B07C" blockId="6.[227,1095,552,643]" box="[227,872,552,577]" pageId="6" pageNumber="7">
<heading id="8637A051FFC90F7EFF04FDA8A421B07C" box="[227,872,552,577]" fontSize="9" level="2" pageId="6" pageNumber="7" reason="2">
<emphasis id="EFB4CB2FFFC90F7EFF04FDA8A421B07C" bold="true" box="[227,872,552,577]" pageId="6" pageNumber="7">
<taxonomicName id="1AC06CBEFFC90F7EFF04FDA8A5C9B07D" ID-CoL="8QJVM" authorityName="Gorochov" authorityYear="2021" box="[227,640,552,577]" class="Insecta" family="Tettigoniidae" genus="Lipotactes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="subSpecies" species="alienus" subGenus="Lipotactes" subSpecies="kubah">
<emphasis id="EFB4CB2FFFC90F7EFF04FDA8A61DB07C" bold="true" box="[227,340,553,577]" italics="true" pageId="6" pageNumber="7">Lipotactes</emphasis>
(
<emphasis id="EFB4CB2FFFC90F7EFE84FDA8A69DB07C" bold="true" box="[355,468,553,577]" italics="true" pageId="6" pageNumber="7">Lipotactes</emphasis>
)
<emphasis id="EFB4CB2FFFC90F7EFE04FDA9A5C9B07D" bold="true" box="[483,640,552,576]" italics="true" pageId="6" pageNumber="7">alienus kubah</emphasis>
</taxonomicName>
Gorochov,
<taxonomicNameLabel id="F4877654FFC90F7EFCE3FDA8A421B07C" box="[772,872,553,577]" pageId="6" pageNumber="7" rank="subSpecies">subsp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FDCBA30EB05D" blockId="6.[227,1095,552,643]" box="[227,1095,586,608]" pageId="6" pageNumber="7">
http://zoobank.org/NomenclaturalActs/
<uuid id="A9662DE8FFC90F7EFD82FDCBA30EB05D" box="[613,1095,586,608]" pageId="6" pageNumber="7">AB6BD62C-7BA3-40B3-BA77-A94EDFCB46CC</uuid>
</paragraph>
</subSubSection>
<subSubSection id="95DA44B6FFC90F7EFF04FDEBA6F1B0BE" box="[227,440,618,643]" pageId="6" pageNumber="7" type="description">
<paragraph id="DD7F173DFFC90F7EFF04FDEBA6F1B0BE" blockId="6.[227,1095,552,643]" box="[227,440,618,643]" pageId="6" pageNumber="7">
<figureCitation id="45FB0BB8FFC90F7EFF04FDEBA608B0BE" box="[227,321,618,643]" captionStart="Figs 1–25" captionStartId="4.[265,306,1145,1167]" captionTargetBox="[228,1119,528,1122]" captionTargetId="figure-116@4.[227,1122,527,1125]" captionTargetPageId="4" captionText="Figs 1–25. Lipotactes spp., schematically: 1–3 – L. (Lipotactes) alienus lambir subsp. n.;" figureDoi="http://doi.org/10.5281/zenodo.7165978" httpUri="https://zenodo.org/record/7165978/files/figure.png" pageId="6" pageNumber="7">Figs 7, 8</figureCitation>
,
<figureCitation id="45FB0BB8FFC90F7EFEAAFDEAA6DBB0BF" box="[333,402,618,643]" captionStart="Figs 26–40" captionStartId="8.[265,306,1527,1549]" captionTargetBox="[262,1083,229,1488]" captionTargetId="figure-2@8.[261,1092,226,1497]" captionTargetPageId="8" captionText="Figs 26–40. Lipotactes spp.: 26–29 – L. (Lipotactes) alienus lambir subsp. n.; 30–33 – L. (L.)" figureDoi="http://doi.org/10.5281/zenodo.7165980" httpUri="https://zenodo.org/record/7165980/files/figure.png" pageId="6" pageNumber="7">34, 35</figureCitation>
,
<figureCitation id="45FB0BB8FFC90F7EFE7AFDEAA6F1B0BE" box="[413,440,619,643]" captionStart="Figs 77–87" captionStartId="21.[265,306,1299,1321]" captionTargetBox="[232,1119,263,1288]" captionTargetId="figure-0@21.[227,1121,257,1293]" captionTargetPageId="21" captionText="Figs 77–87. Lipotactes spp.: 77 – L. (Lipotactes) alienus lambir subsp. n.; 78 – L. (L.) a." figureDoi="http://doi.org/10.5281/zenodo.7165986" httpUri="https://zenodo.org/record/7165986/files/figure.png" pageId="6" pageNumber="7">79</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="95DA44B6FFC90F7EFEEEFD24A429B0CA" pageId="6" pageNumber="7" type="materials_examined">
<paragraph id="DD7F173DFFC90F7EFEEEFD24A32EB080" blockId="6.[227,1127,676,1575]" box="[265,1127,676,701]" pageId="6" pageNumber="7">
<materialsCitation id="6DA81D60FFC90F7EFEEEFD24A32EB080" box="[265,1127,676,701]" collectorName="Borneo I." country="Malaysia" location="Sarawak State" pageId="6" pageNumber="7" specimenCount="1" specimenCount-male="1" stateProvince="Sarawak" typeStatus="holotype">
MATERIAL.
<typeStatus id="027BA99FFFC90F7EFE47FD24A548B080" box="[416,513,677,701]" pageId="6" pageNumber="7" type="holotype">Holotype</typeStatus>
–
<specimenCount id="CBC6DCB4FFC90F7EFDFCFD24A578B081" box="[539,561,677,700]" pageId="6" pageNumber="7" type="male">♂</specimenCount>
,
<emphasis id="EFB4CB2FFFC90F7EFDDBFD25A5EBB081" bold="true" box="[572,674,676,700]" pageId="6" pageNumber="7">
<collectingCountry id="A5D757ADFFC90F7EFDDBFD25A5EBB081" box="[572,674,676,700]" name="Malaysia" pageId="6" pageNumber="7">Malaysia</collectingCountry>
</emphasis>
:
<collectorName id="703572EBFFC90F7EFD57FD24A45AB081" box="[688,787,677,700]" pageId="6" pageNumber="7">Borneo I.</collectorName>
,
<collectingRegion id="1F04D9DFFFC90F7EFCF8FD24A4FEB080" box="[799,951,677,701]" country="Malaysia" name="Sarawak" pageId="6" pageNumber="7">Sarawak State</collectingRegion>
, Kubah National
</materialsCitation>
</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FD43A32FB0E7" blockId="6.[227,1127,676,1575]" box="[227,1126,706,730]" pageId="6" pageNumber="7">
Park on Matang Mt near Kuching City,
<quantity id="1A38BAD8FFC90F7EFD42FD43A46CB0E4" box="[677,805,706,730]" metricMagnitude="2" metricUnit="m" metricValue="3.5" metricValueMax="5.0" metricValueMin="2.0" pageId="6" pageNumber="7" unit="m" value="350.0" valueMax="500.0" valueMin="200.0">200–500 m</quantity>
, primary forest, 27.XI–1.XII
</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FD5EA429B0CA" blockId="6.[227,1127,676,1575]" box="[227,864,735,759]" pageId="6" pageNumber="7">2016, A. Gorochov, M. Berezin, E. Tkatsheva, I. Kamskov.</paragraph>
</subSubSection>
<subSubSection id="95DA44B6FFC90F7FFEEEFD7DA5A7B366" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="description">
<paragraph id="DD7F173DFFC90F7EFEEEFD7DA32EB10C" blockId="6.[227,1127,676,1575]" pageId="6" pageNumber="7">
DESCRIPTION.
<emphasis id="EFB4CB2FFFC90F7EFE23FD7DA6B2B129" box="[452,507,764,788]" italics="true" pageId="6" pageNumber="7">Male</emphasis>
(
<typeStatus id="027BA99FFFC90F7EFDEDFD7DA524B129" box="[522,621,764,788]" pageId="6" pageNumber="7" type="holotype">holotype</typeStatus>
). Size, coloration and structure of body similar to those of
<emphasis id="EFB4CB2FFFC90F7EFEB9FC9BA63DB10C" box="[350,372,794,817]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
(
<emphasis id="EFB4CB2FFFC90F7EFE63FC9BA6D3B10C" box="[388,410,794,817]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
)
<emphasis id="EFB4CB2FFFC90F7EFE4CFC9BA544B10C" box="[427,525,793,817]" italics="true" pageId="6" pageNumber="7">a. lambir</emphasis>
and
<emphasis id="EFB4CB2FFFC90F7EFDA3FC9BA446B10C" box="[580,783,793,817]" italics="true" pageId="6" pageNumber="7">L. (L.) a. similajau</emphasis>
but with following differences:
</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FCB7A32FB1B4" blockId="6.[227,1127,676,1575]" pageId="6" pageNumber="7">head with rostrum and dorsum almost uniformly yellowish; pronotum yellowish with brown anterior margin of disc and interrupted sinuate transverse line on disc near this margin as well as a pair of short oblique lines slightly behind previous (sinuate)</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FC0FA32EB1C0" blockId="6.[227,1127,676,1575]" pageId="6" pageNumber="7">
line; fore and middle femora uniformly yellowish; hind femur also uniformly yellowish but with brown apex and a pair of dark brown longitudinal spots on inner and outer surfaces very near this apex (almost as in
<emphasis id="EFB4CB2FFFC90F7EFCD0FC48A48AB1DD" box="[823,963,968,992]" italics="true" pageId="6" pageNumber="7">
L. a.
<taxonomicName id="1AC06CBEFFC90F7EFC91FC49A48AB1DD" authorityName="Brunner-Wattenwyl" authorityYear="1898" box="[886,963,968,992]" class="Insecta" family="Tettigoniidae" genus="Lipotactes" kingdom="Animalia" order="Orthoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="alienus">alienus</taxonomicName>
</emphasis>
); rest of body yellowish with more or less light brown majority of abdominal tergites and sternites
</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FB83A32EB627" blockId="6.[227,1127,676,1575]" box="[227,1127,1026,1050]" pageId="6" pageNumber="7">(in living condition, latter sternites possibly lighter), cercal and paraproctal apices,</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FB9EA32EB691" blockId="6.[227,1127,676,1575]" pageId="6" pageNumber="7">
cercal proximedial hooks, median part of epiproct, and styles of genital plate, as well as with brown dorsum of last tergite (
<figureCitation id="45FB0BB8FFC90F7EFD4CFBBCA460B669" box="[683,809,1084,1109]" captionStart="Figs 26–40" captionStartId="8.[265,306,1527,1549]" captionTargetBox="[262,1083,229,1488]" captionTargetId="figure-2@8.[261,1092,226,1497]" captionTargetPageId="8" captionText="Figs 26–40. Lipotactes spp.: 26–29 – L. (Lipotactes) alienus lambir subsp. n.; 30–33 – L. (L.)" figureDoi="http://doi.org/10.5281/zenodo.7165980" httpUri="https://zenodo.org/record/7165980/files/figure.png" pageId="6" pageNumber="7">Figs 34, 35</figureCitation>
); tegmina with slightly larger loop-like cell behind stridulatory vein and wider apical area as well as somewhat longer (moderately short) stridulatory vein of left tegmen (
<figureCitation id="45FB0BB8FFC90F7EFCBDFBF6A4E2B6B2" box="[858,939,1143,1167]" captionStart="Figs 77–87" captionStartId="21.[265,306,1299,1321]" captionTargetBox="[232,1119,263,1288]" captionTargetId="figure-0@21.[227,1121,257,1293]" captionTargetPageId="21" captionText="Figs 77–87. Lipotactes spp.: 77 – L. (Lipotactes) alienus lambir subsp. n.; 78 – L. (L.) a." figureDoi="http://doi.org/10.5281/zenodo.7165986" httpUri="https://zenodo.org/record/7165986/files/figure.png" pageId="6" pageNumber="7">Fig. 79</figureCitation>
); last tergite with posteromedian notch as wide as in
<emphasis id="EFB4CB2FFFC90F7EFDB5FB14A521B691" box="[594,616,1173,1196]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
(
<emphasis id="EFB4CB2FFFC90F7EFD90FB14A5C4B691" box="[631,653,1173,1196]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
)
<emphasis id="EFB4CB2FFFC90F7EFD7AFB14A5B4B691" box="[669,765,1172,1196]" italics="true" pageId="6" pageNumber="7">a. lambir</emphasis>
and almost as deep as in
<emphasis id="EFB4CB2FFFC90F7EFBEFFB14A357B691" box="[1032,1054,1173,1196]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
(
<emphasis id="EFB4CB2FFFC90F7EFBC9FB14A30DB691" box="[1070,1092,1173,1196]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
)
<emphasis id="EFB4CB2FFFC90F7EFBB4FB14A32EB691" box="[1107,1127,1173,1196]" italics="true" pageId="6" pageNumber="7">a.</emphasis>
</paragraph>
<paragraph id="DD7F173DFFC90F7EFF04FB30A46BB703" blockId="6.[227,1127,676,1575]" pageId="6" pageNumber="7">
<emphasis id="EFB4CB2FFFC90F7EFF04FB30A60DB6F4" box="[227,324,1201,1225]" italics="true" pageId="6" pageNumber="7">similajau</emphasis>
; proximedial cercal hook with ventral branch as long as in these subspecies but having barely widened (not narrowed) apical part, and with dorsal lobule intermediate between such lobules of above-mentioned taxa in shape but almost pressed to ventral cercal branch (
<figureCitation id="45FB0BB8FFC90F7EFE10FA89A515B71C" box="[503,604,1288,1313]" captionStart="Figs 1–25" captionStartId="4.[265,306,1145,1167]" captionTargetBox="[228,1119,528,1122]" captionTargetId="figure-116@4.[227,1122,527,1125]" captionTargetPageId="4" captionText="Figs 1–25. Lipotactes spp., schematically: 1–3 – L. (Lipotactes) alienus lambir subsp. n.;" figureDoi="http://doi.org/10.5281/zenodo.7165978" httpUri="https://zenodo.org/record/7165978/files/figure.png" pageId="6" pageNumber="7">Figs 7, 8</figureCitation>
); posteromedian notch of genital plate between styles even deeper than in
<emphasis id="EFB4CB2FFFC90F7EFE1AFAA7A55AB700" box="[509,531,1318,1341]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
(
<emphasis id="EFB4CB2FFFC90F7EFDC5FAA7A571B700" box="[546,568,1318,1341]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
)
<emphasis id="EFB4CB2FFFC90F7EFDA0FAA7A58CB700" box="[583,709,1317,1341]" italics="true" pageId="6" pageNumber="7">a. similajau</emphasis>
(
<figureCitation id="45FB0BB8FFC90F7EFD33FAA4A45FB703" box="[724,790,1317,1342]" captionStart="Figs 1–25" captionStartId="4.[265,306,1145,1167]" captionTargetBox="[228,1119,528,1122]" captionTargetId="figure-116@4.[227,1122,527,1125]" captionTargetPageId="4" captionText="Figs 1–25. Lipotactes spp., schematically: 1–3 – L. (Lipotactes) alienus lambir subsp. n.;" figureDoi="http://doi.org/10.5281/zenodo.7165978" httpUri="https://zenodo.org/record/7165978/files/figure.png" pageId="6" pageNumber="7">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="DD7F173DFFC90F7EFEEEFAC2A683B766" blockId="6.[227,1127,676,1575]" box="[265,458,1347,1371]" pageId="6" pageNumber="7">
<emphasis id="EFB4CB2FFFC90F7EFEEEFAC2A610B766" box="[265,345,1347,1371]" italics="true" pageId="6" pageNumber="7">Female</emphasis>
unknown.
</paragraph>
<paragraph id="DD7F173DFFC90F7EFEEEFAE1A595B7A8" blockId="6.[227,1127,676,1575]" pageId="6" pageNumber="7">MEASUREMENTS. Length (in mm). Body 11; pronotum 4.2; tegminal parts visible behind pronotum 0.6; hind femora 11.7.</paragraph>
<paragraph id="DD7F173DFFC90F7FFEEEFA1BA5A7B366" blockId="6.[227,1127,676,1575]" lastBlockId="7.[227,1127,233,407]" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7">
COMPARISON. The new subspecies differs from the nominotypical one in the male tegmina having the apical area wider, and in the male cercal proximedial hook with a widened apex of its ventral branch and with the dorsal lobule almost pressed to the latter branch. From all the other subspecies of this species,
<emphasis id="EFB4CB2FFFC90F7EFC5BFA73A49BB434" box="[956,978,1522,1545]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
(
<emphasis id="EFB4CB2FFFC90F7EFC05FA73A4B1B434" box="[994,1016,1522,1545]" italics="true" pageId="6" pageNumber="7">L.</emphasis>
)
<emphasis id="EFB4CB2FFFC90F7EFBEFFA73A32FB434" box="[1032,1126,1521,1545]" italics="true" pageId="6" pageNumber="7">a. kubah</emphasis>
is distinguished by the same characters of the male tegmina and cerci, as well as by the stridulatory vein of the left tegmen longer, the loop-like tegminal cell larger, the body coloration lighter and more uniform, the male last tergite with the posteromedian notch wider or deeper, and the male genital plate with the posteromedian notch deeper (see
<figureCitation id="45FB0BB8FFC80F7FFDA6FEC2A5A8B366" box="[577,737,323,347]" captionStart="Figs 1–25" captionStartId="4.[265,306,1145,1167]" captionTargetBox="[228,1119,528,1122]" captionTargetId="figure-116@4.[227,1122,527,1125]" captionTargetPageId="4" captionText="Figs 1–25. Lipotactes spp., schematically: 1–3 – L. (Lipotactes) alienus lambir subsp. n.;" figureDoi="http://doi.org/10.5281/zenodo.7165978" httpUri="https://zenodo.org/record/7165978/files/figure.png" pageId="7" pageNumber="8">Figs 1, 4 and 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="95DA44B6FFC80F7FFEEEFEE0A6D9B3AA" pageId="7" pageNumber="8" type="etymology">
<paragraph id="DD7F173DFFC80F7FFEEEFEE0A6D9B3AA" blockId="7.[227,1127,233,407]" pageId="7" pageNumber="8">ETYMOLOGY. The new subspecies is named after Kubah National Park where it was collected.</paragraph>
</subSubSection>
</treatment>
</document>
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