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<document id="447CD0967859F5C7F3C93FB2F70EBD07" ID-DOI="10.5281/zenodo.5169575" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169575" ID-ZooBank="89FC75EA-2324-4361-B818-FBA7B7682A00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628278715662" checkinUser="felipe" docAuthor="Larochelle, Larivière, Marie-Claude, Larochelle &amp; Larivière" docDate="2017" docId="556787D3027FFFBD3FCAFC77FD58FC27" docLanguage="en" docName="InsectaMundi.0594.1-110.pdf" docOrigin="Insecta Mundi 2017 (594)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Oopterus laeviventris" docType="treatment" docVersion="4" lastPageNumber="32" masterDocId="A95EFFAB025FFF9C3F0AFFF6FFFFFF9D" masterDocTitle="Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="110" masterPageNumber="1" pageNumber="31" updateTime="1699039365375" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="F29C8B5761A79483464AD4C86DB683EF">Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="2C950B3C7F56B7FADD357975271CB570">Larochelle</mods:namePart>
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<mods:namePart id="A62764CCC36ED393E7FC3D7B2D51EC76">Larivière, Marie-Claude</mods:namePart>
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<mods:namePart id="49239031451EFDC4699423FA3308673D">Larochelle</mods:namePart>
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<mods:namePart id="E119564493A11A68710A051752E980EF">Larivière</mods:namePart>
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<paragraph id="DD7136C5027FFFBC3FCAFC77FD5CFC04" blockId="32.[192,675,897,954]" box="[192,675,897,921]" pageId="32" pageNumber="31">
<heading id="863981A9027FFFBC3FCAFC77FD5CFC04" bold="true" box="[192,675,897,921]" fontSize="10" level="2" pageId="32" pageNumber="31" reason="3">
<taxonomicName id="1ACE4D46027FFFBC3FCAFC77FD5CFC04" ID-CoL="49SWS" authority="(Sharp, 1883)" baseAuthorityName="Sharp" baseAuthorityYear="1883" box="[192,675,897,921]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="laeviventris">
<emphasis id="EFBAEAD7027FFFBC3FCAFC77FD5CFC04" bold="true" box="[192,675,897,921]" pageId="32" pageNumber="31">
<emphasis id="EFBAEAD7027FFFBC3FCAFC77FE19FC04" bold="true" box="[192,486,897,921]" italics="true" pageId="32" pageNumber="31">Oopterus laeviventris</emphasis>
(
<bibRefCitation id="B95F4B34027FFFBC3EFDFC77FD65FC04" author="Sharp, D." box="[503,666,897,921]" pageId="32" pageNumber="31" pagination="66 - 68" refId="ref56483" refString="Sharp, D. 1883. Some new species and genera of Coleoptera from New Zealand. Entomologist's Monthly Magazine 20: 23 - 27, 66 - 68." type="journal article" year="1883">Sharp, 1883</bibRefCitation>
)
</emphasis>
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</heading>
</paragraph>
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<subSubSection id="95D4654E027FFFBC3FCAFC54FE83FC27" box="[192,380,930,954]" pageId="32" pageNumber="31" type="description">
<paragraph id="DD7136C5027FFFBC3FCAFC54FE83FC27" blockId="32.[192,675,897,954]" box="[192,380,930,954]" pageId="32" pageNumber="31">
<figureCitation id="45F52A40027FFFBC3FCAFC54FEE8FC27" box="[192,279,930,954]" captionStart="Figures 3134" captionStartId="84.[192,288,1832,1854]" captionTargetBox="[214,1427,197,1815]" captionTargetId="figure-46@84.[192,1435,192,1818]" captionTargetPageId="84" captionText="Figures 3134. Dorsal habitus. 31) Oopterus fulvipes. 32) O. sobrinus. 33) O. collaris. 34) O. laeviventris. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5169589" httpUri="https://zenodo.org/record/5169589/files/figure.png" pageId="32" pageNumber="31">Fig. 34</figureCitation>
,
<figureCitation id="45F52A40027FFFBC3E2EFC54FEBBFC27" box="[292,324,930,954]" captionStart="Figures 7481" captionStartId="94.[192,288,1832,1854]" captionTargetBox="[254,1414,252,1806]" captionTargetId="figure-59@94.[192,1440,192,1819]" captionTargetPageId="94" captionText="Figures 7481. Aedeagi, lateral. 74) Oopterus discoideus new species. 75) O. marrisi new species. 76) O. lewisi. 77) O. patulus. 78) O. fulvipes. 79) O. sobrinus. 80) O. collaris. 81) O. laeviventris." figureDoi="http://doi.org/10.5281/zenodo.5169615" httpUri="https://zenodo.org/record/5169615/files/figure.png" pageId="32" pageNumber="31">81</figureCitation>
,
<figureCitation id="45F52A40027FFFBC3E5AFC54FE83FC27" box="[336,380,930,954]" captionStart="Figures 130135" captionStartId="103.[192,288,1829,1851]" captionTargetBox="[192,1412,192,1818]" captionTargetId="figure-48@103.[192,1412,192,1818]" captionTargetPageId="103" captionText="Figures 130135. Species distribution maps. 130) Oopterus laeviventris. 131) O. lewisi. 132) O. marrineri. 133) O. marrisi new species. 134) O. mohiensis new species. 135) O. monticola new species." figureDoi="http://doi.org/10.5281/zenodo.5169635" httpUri="https://zenodo.org/record/5169635/files/figure.png" pageId="32" pageNumber="31">130</figureCitation>
</paragraph>
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<subSubSection id="95D4654E027FFFBC3FCAFC14FD6CFB04" pageId="32" pageNumber="31" type="reference_group">
<paragraph id="DD7136C5027FFFBC3FCAFC14FDE7FBC7" blockId="32.[192,1440,994,1178]" pageId="32" pageNumber="31">
<treatmentCitationGroup id="FDDE11EB027FFFBC3FCAFC14FDE7FBC7" pageId="32" pageNumber="31">
<treatmentCitation id="5C6F10D4027FFFBC3FCAFC14FD42FC67" author="Sharp, D." box="[192,701,994,1018]" page="24" pageId="32" pageNumber="31" year="1883">
<taxonomicName id="1ACE4D46027FFFBC3FCAFC14FD42FC67" ID-CoL="7BKT6" authority="Sharp, 1883: 24" authorityName="Sharp" authorityPageNumber="24" authorityYear="1883" box="[192,701,994,1018]" class="Insecta" family="Carabidae" genus="Tarastethus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="laeviventris">
<emphasis id="EFBAEAD7027FFFBC3FCAFC14FE13FC67" box="[192,492,994,1018]" italics="true" pageId="32" pageNumber="31">Tarastethus laeviventris</emphasis>
<materialsCitation id="6DA63C98027FFFBC3EFEFC14FD42FC67" box="[500,701,994,1018]" collectionCode="BMNH, BR, T" country="New Zealand" location="Greymouth" pageId="32" pageNumber="31" specimenCount="1" typeStatus="holotype">
<bibRefCitation id="B95F4B34027FFFBC3EFEFC14FD42FC67" author="Sharp, D." box="[500,701,994,1018]" pageId="32" pageNumber="31" pagination="66 - 68" refId="ref56483" refString="Sharp, D. 1883. Some new species and genera of Coleoptera from New Zealand. Entomologist's Monthly Magazine 20: 23 - 27, 66 - 68." type="journal article" year="1883">Sharp, 1883: 24</bibRefCitation>
</materialsCitation>
</taxonomicName>
</treatmentCitation>
.
<typeStatus id="02758867027FFFBC3DC0FC14FCC2FC67" box="[714,829,994,1018]" pageId="32" pageNumber="31" type="holotype">Holotype</typeStatus>
: male (BMNH) labeled: “
<taxonomicName id="1ACE4D46027FFFBC3B7FFC14FA5FFC67" ID-CoL="7BKT6" authorityName="Sharp" authorityYear="1883" box="[1141,1440,994,1018]" class="Insecta" family="Carabidae" genus="Tarastethus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="laeviventris">
<emphasis id="EFBAEAD7027FFFBC3B7FFC14FA5FFC67" box="[1141,1440,994,1018]" italics="true" pageId="32" pageNumber="31">Tarastethus laeviventris</emphasis>
</taxonomicName>
[male symbol] type D.S. [BR] Greymouth Helms. (hand-written) / Type H.T. (circular red-bordered label; typed) /
<location id="D811601E027FFFBC3E90FBD4FDD9FBA7" LSID="urn:lsid:plazi:treatment:556787D3027FFFBD3FCAFC77FD58FC27:D811601E027FFFBC3E90FBD4FDD9FBA7" box="[410,550,1058,1082]" country="New Zealand" name="Greymouth" pageId="32" pageNumber="31">Greymouth</location>
<collectingCountry id="A5D97655027FFFBC3D27FBD4FD31FBA7" box="[557,718,1058,1082]" name="New Zealand" pageId="32" pageNumber="31">New Zealand</collectingCountry>
Helms. (white label with red horizontal line; typed) / Sharp Coll. 1905-313. (typed).”
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</paragraph>
<paragraph id="DD7136C5027FFFBC3FCAFB94FD3AFBE7" blockId="32.[192,1440,994,1178]" box="[192,709,1122,1146]" pageId="32" pageNumber="31">
<treatmentCitationGroup id="FDDE11EB027FFFBC3FCAFB94FD3AFBE7" box="[192,709,1122,1146]" pageId="32" pageNumber="31">
<treatmentCitation id="5C6F10D4027FFFBC3FCAFB94FD3EFBE7" author="Britton, E. B." box="[192,705,1122,1146]" page="477" pageId="32" pageNumber="31" year="1940">
<taxonomicName id="1ACE4D46027FFFBC3FCAFB94FD3EFBE7" ID-CoL="43Y3F" authority=": Britton 1940: 477" authorityName="Britton" authorityPageNumber="477" authorityYear="1940" box="[192,705,1122,1146]" class="Insecta" family="Carabidae" genus="Molopsida" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="laeviventris">
<emphasis id="EFBAEAD7027FFFBC3FCAFB94FE28FBE7" box="[192,471,1122,1146]" italics="true" pageId="32" pageNumber="31">Molopsida laeviventris</emphasis>
:
<bibRefCitation id="B95F4B34027FFFBC3EEFFB94FD3EFBE7" author="Britton, E. B." box="[485,705,1122,1146]" pageId="32" pageNumber="31" pagination="473 - 508" refId="ref54644" refString="Britton, E. B. 1940. The Carabidae (Coleoptera) of New Zealand. Part 1, Pterostichini. Transactions of the Royal Society of New Zealand 69: 473 - 508 + pl. 68 - 75." type="journal article" year="1940">Britton 1940: 477</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="DD7136C5027FFFBC3FCAFB74FD6CFB04" blockId="32.[192,1440,994,1178]" box="[192,659,1153,1178]" pageId="32" pageNumber="31">
<treatmentCitationGroup id="FDDE11EB027FFFBC3FCAFB74FD6CFB04" box="[192,659,1153,1178]" pageId="32" pageNumber="31">
<treatmentCitation id="5C6F10D4027FFFBC3FCAFB74FD70FB04" author="Johns, P. M." box="[192,655,1153,1178]" page="57" pageId="32" pageNumber="31" year="1980">
<taxonomicName id="1ACE4D46027FFFBC3FCAFB74FD70FB04" ID-CoL="49SWS" authority=": Johns 1980: 57" authorityName=": Johns" authorityPageNumber="57" authorityYear="1980" box="[192,655,1153,1178]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="31" phylum="Arthropoda" rank="species" species="laeviventris">
<emphasis id="EFBAEAD7027FFFBC3FCAFB74FE3AFB07" box="[192,453,1154,1178]" italics="true" pageId="32" pageNumber="31">Oopterus laeviventris</emphasis>
:
<bibRefCitation id="B95F4B34027FFFBC3ED9FB74FD70FB04" author="Johns, P. M." box="[467,655,1153,1178]" pageId="32" pageNumber="31" pagination="55 - 67" refId="ref55828" refString="Johns, P. M. 1980. Ground beetles (Carabidae) of Arthur's Pass National Park. Mauri Ora 8: 55 - 67." type="journal article" year="1980">Johns 1980: 57</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027FFFBD3FCAFB37FE7CFEE4" lastPageId="33" lastPageNumber="32" pageId="32" pageNumber="31" type="description">
<paragraph id="DD7136C5027FFFBD3FCAFB37FE7CFEE4" blockId="32.[192,1441,1217,1914]" lastBlockId="33.[192,1440,194,378]" lastPageId="33" lastPageNumber="32" pageId="32" pageNumber="31">
<emphasis id="EFBAEAD7027FFFBC3FCAFB37FE9CFB44" bold="true" box="[192,355,1217,1241]" pageId="32" pageNumber="31">Description</emphasis>
. Body length
<quantity id="1A369B20027FFFBC3D18FB37FD58FB44" box="[530,679,1217,1241]" metricMagnitude="-3" metricUnit="m" metricValue="4.699999999999999" metricValueMax="5.3" metricValueMin="4.1" pageId="32" pageNumber="31" unit="mm" value="4.699999999999999" valueMax="5.3" valueMin="4.1">4.15.3 mm</quantity>
. Head, pronotum, elytra, and abdomen mostly piceous black; apex of head reddish; lateral margins, suture, and apex of elytra reddish; abdomen reddish laterally and apically; antennae, palpi, and legs pale yellow; base of femora piceous black. Microsculpture absent. Iridescence absent. Very shiny, without metallic luster.
<emphasis id="EFBAEAD7027FFFBC3C76FAD4FC3AFAA7" bold="true" box="[892,965,1314,1338]" pageId="32" pageNumber="31">Head</emphasis>
. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, moderately long: segment 1 (scape) very long, about 2.5x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora not inflated. Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, about as long as ligula.
<emphasis id="EFBAEAD7027FFFBC3E65FA34FE2CFA47" bold="true" box="[367,467,1474,1498]" pageId="32" pageNumber="31">Thorax</emphasis>
. Pronotum strongly convex, coarsely punctate across base, unwrinkled, strongly transverse, slightly cordate, widest about middle; apex subtruncate; anterolateral angles poorly developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, slightly sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (anteriorly); posterolateral angles obtuse; posterolateral carinae blunt, very long (about as long as adjoining foveae); laterobasal foveae well defined, coarsely punctate, very deep and wide, subrectangular, not prolonged forward, double; posterior bead absent; base emarginate, wider than pronotal apex, about as wide as elytral base.
<emphasis id="EFBAEAD7027FFFBC3E51F954FE64F927" bold="true" box="[347,411,1698,1722]" pageId="32" pageNumber="31">Legs</emphasis>
. Moderately long.
<emphasis id="EFBAEAD7027FFFBC3D73F954FD2FF927" bold="true" box="[633,720,1698,1722]" pageId="32" pageNumber="31">Elytra</emphasis>
. Strongly convex, ovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders rounded. Sides strongly rounded. Scutellar striole present. Striae mostly replaced by rows of coarse punctures; stria 1 complete, very deep; rows 25 moderately deep, obsolete apically; rows 67 obsolete basally; interval 3 with three setiferous punctures, close to row 3. Recurrent stria long, sharp, directed apically toward row 5. Subapical seta present. Intervals slightly convex. Sutural apices obtuse.
<emphasis id="EFBAEAD7027FFFBC3DAAF8B7FCDCF8C4" bold="true" box="[672,803,1857,1881]" pageId="32" pageNumber="31">Abdomen</emphasis>
. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae.
<emphasis id="EFBAEAD7027FFFBC3CF6F897FB7EF8E4" bold="true" box="[1020,1153,1889,1913]" pageId="32" pageNumber="31">Aedeagus</emphasis>
. Lateral view (
<figureCitation id="45F52A40027FFFBC3A31F894FA6DF8E7" box="[1339,1426,1890,1914]" captionStart="Figures 7481" captionStartId="94.[192,288,1832,1854]" captionTargetBox="[254,1414,252,1806]" captionTargetId="figure-59@94.[192,1440,192,1819]" captionTargetPageId="94" captionText="Figures 7481. Aedeagi, lateral. 74) Oopterus discoideus new species. 75) O. marrisi new species. 76) O. lewisi. 77) O. patulus. 78) O. fulvipes. 79) O. sobrinus. 80) O. collaris. 81) O. laeviventris." figureDoi="http://doi.org/10.5281/zenodo.5169615" httpUri="https://zenodo.org/record/5169615/files/figure.png" pageId="32" pageNumber="31">Fig. 81</figureCitation>
): moderately arcuate, moderately widened in apical half; base strongly biconvex dorsally; middle slightly concave dorsally, mostly straight ventrally, with dorsal membranous area very wide and moderately long; apex subtriangular, slightly concave dorsally, straight ventrally, with extreme tip wide and long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with five or six apical setae.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027EFFBD3FCAFE57FBD4FE64" pageId="33" pageNumber="32" type="materials_examined">
<paragraph id="DD7136C5027EFFBD3FCAFE57FAC8FE27" blockId="33.[192,1335,417,442]" box="[192,1335,417,442]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFE57FE3AFE24" bold="true" box="[192,453,417,441]" pageId="33" pageNumber="32">Material examined</emphasis>
.
<specimenCount id="CBC8FD4C027EFFBD3EDEFE57FD78FE27" box="[468,647,417,442]" pageId="33" pageNumber="32" type="generic">157 specimens</specimenCount>
(
<collectionCode id="BBDFAE00027EFFBD3D9CFE54FD12FE27" box="[662,749,418,442]" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="33" pageNumber="32" type="Museum">AMNZ</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3DF2FE54FCACFE24" box="[760,851,418,441]" pageId="33" pageNumber="32">BMNH</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3C55FE54FC4AFE27" box="[863,949,418,442]" country="New Zealand" httpUri="http://grbio.org/cool/n6z3-qvv0" name="Canterbury Museum" pageId="33" pageNumber="32">CMNZ</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3CCBFE54FBF0FE24" box="[961,1039,418,441]" pageId="33" pageNumber="32">JNNZ</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3B10FE54FB94FE24" box="[1050,1131,418,441]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="33" pageNumber="32">LUNZ</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3B7CFE54FB32FE27" box="[1142,1229,418,442]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="33" pageNumber="32" type="Museum">MONZ</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3BD3FE54FAD4FE27" box="[1241,1323,418,442]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="33" pageNumber="32" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="DD7136C5027EFFBD3FCAFE17FBD4FE64" blockId="33.[192,1067,481,506]" box="[192,1067,481,506]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFE17FDEFFE64" bold="true" box="[192,528,481,505]" pageId="33" pageNumber="32">Geographic distribution</emphasis>
(
<figureCitation id="45F52A40027EFFBD3D2AFE14FD77FE67" box="[544,648,482,506]" captionStart="Figures 130135" captionStartId="103.[192,288,1829,1851]" captionTargetBox="[192,1412,192,1818]" captionTargetId="figure-48@103.[192,1412,192,1818]" captionTargetPageId="103" captionText="Figures 130135. Species distribution maps. 130) Oopterus laeviventris. 131) O. lewisi. 132) O. marrineri. 133) O. marrisi new species. 134) O. mohiensis new species. 135) O. monticola new species." figureDoi="http://doi.org/10.5281/zenodo.5169635" httpUri="https://zenodo.org/record/5169635/files/figure.png" pageId="33" pageNumber="32">Fig. 130</figureCitation>
). South Island:
<collectionCode id="BBDFAE00027EFFBD3C43FE14FC8DFE64" box="[841,882,482,505]" country="Brazil" name="Embrapa Agrobiology Diazothrophic Microbial Culture Collection" pageId="33" pageNumber="32">BR</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3C75FE14FC51FE67" box="[895,942,482,506]" country="0" name="Museo de Cipolleti" pageId="33" pageNumber="32">MC</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3CB0FE14FC17FE64" box="[954,1000,482,505]" pageId="33" pageNumber="32">NN</collectionCode>
,
<collectionCode id="BBDFAE00027EFFBD3CFEFE14FBD8FE64" box="[1012,1063,482,505]" pageId="33" pageNumber="32">WD</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027EFFBD3FCAFDD4FB56FCC7" pageId="33" pageNumber="32" type="biology_ecology">
<paragraph id="DD7136C5027EFFBD3FCAFDD4FDB5FDE7" blockId="33.[192,1440,546,634]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFDD4FED2FDA7" bold="true" box="[192,301,546,570]" pageId="33" pageNumber="32">Ecology</emphasis>
. Lowland, montane, subalpine. Epigean-arboreal. Wet forests (beech, podocarp, broadleaf) and scrublands. Shaded ground. Nocturnal; active at night on mossy logs; hides during the day in moss and leaf litter, as well as under logs.
</paragraph>
<paragraph id="DD7136C5027EFFBD3FCAFD57FDEDFD47" blockId="33.[192,1440,673,730]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFD57FED7FD24" bold="true" box="[192,296,673,697]" pageId="33" pageNumber="32">Biology</emphasis>
. Seasonality: September, NovemberJune. Tenerals: FebruaryMarch (BR). Predacious (based on mouthpart morphology).
</paragraph>
<paragraph id="DD7136C5027EFFBD3FCAFCF7FA7FFC87" blockId="33.[192,1408,769,794]" box="[192,1408,769,794]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFCF7FE5DFC87" bold="true" box="[192,418,769,794]" pageId="33" pageNumber="32">Dispersal power</emphasis>
. Subapterous (incapable of flight). Moderate runner. Occasional climber on logs.
</paragraph>
<paragraph id="DD7136C5027EFFBD3FCAFCB7FB56FCC7" blockId="33.[192,1193,833,858]" box="[192,1193,833,858]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFCB7FE12FCC4" bold="true" box="[192,493,833,857]" pageId="33" pageNumber="32">Collecting techniques</emphasis>
. Pitfall trapping; sifting moss and leaf litter; lifting logs.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027EFFBD3FCAFC77FD58FC27" pageId="33" pageNumber="32" type="reference_group">
<paragraph id="DD7136C5027EFFBD3FCAFC77FD58FC27" blockId="33.[192,1441,897,954]" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFC77FEA7FC04" bold="true" box="[192,344,897,921]" pageId="33" pageNumber="32">References</emphasis>
.
<bibRefCitation id="B95F4B34027EFFBD3E6FFC74FD05FC07" author="Larochelle, A. &amp; M. - C. Lariviere" box="[357,762,898,922]" pageId="33" pageNumber="32" pagination="1 - 281" refId="ref55890" refString="Larochelle, A., and M. - C. Lariviere. 2001. Carabidae (Insecta: Coleoptera): catalogue. Fauna of New Zealand 43: 1 - 281." type="journal article" year="2001">Larochelle and Larivière 2001: 74</bibRefCitation>
(catalogue; biology, dispersal power, ecology, geographic distribution, references), 2016: 20 (list).
</paragraph>
</subSubSection>
</treatment>
</document>