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<document id="5C883D5D7A477C356FE925CC9FC9616C" ID-DOI="10.5281/zenodo.5169575" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169575" ID-ZooBank="89FC75EA-2324-4361-B818-FBA7B7682A00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628278715662" checkinUser="felipe" docAuthor="Larochelle, Larivière, Marie-Claude, Larochelle &amp; Larivière" docDate="2017" docId="556787D3027EFFBE3FCAFBF4FAFCFCC7" docLanguage="en" docName="InsectaMundi.0594.1-110.pdf" docOrigin="Insecta Mundi 2017 (594)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Oopterus arthurensis Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière 2017, new species" docType="treatment" docVersion="4" lastPageNumber="33" masterDocId="A95EFFAB025FFF9C3F0AFFF6FFFFFF9D" masterDocTitle="Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="110" masterPageNumber="1" pageNumber="32" updateTime="1699039365375" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="017B3660D69CB9605DD0D42764239C1D">Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="A764D615E982BE2C34189881A16A5CB5">Larochelle</mods:namePart>
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<mods:namePart id="8924366C679EE5B4E71D34767719A42C">Larivière, Marie-Claude</mods:namePart>
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<mods:namePart id="9A2F9E5CB5C082C19408C0E5603009F4">Larochelle</mods:namePart>
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<mods:namePart id="ECE2B9437FCDA5A1C805BF2B73719848">Larivière</mods:namePart>
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<mods:title id="3E0EFCAB58596CB7FA71D90D229853DB">Insecta Mundi</mods:title>
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<mods:date id="4B83951410B4E0A17E1F14DDAFA63AB1">2017</mods:date>
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<mods:number id="0D7F17082D3F061063FD39941E98848F">2017-12-29</mods:number>
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<mods:identifier id="EC5F9CA26E3D0068EDBA0CC9EBE7ECAB" type="ISSN">1942-1354</mods:identifier>
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<paragraph id="DD7136C5027EFFBD3FCAFBF4FD6DFB84" blockId="33.[192,658,1025,1082]" box="[192,658,1025,1050]" pageId="33" pageNumber="32">
<heading id="863981A9027EFFBD3FCAFBF4FD6DFB84" bold="true" box="[192,658,1025,1050]" fontSize="10" level="2" pageId="33" pageNumber="32" reason="3">
<emphasis id="EFBAEAD7027EFFBD3FCAFBF4FD6DFB84" bold="true" box="[192,658,1025,1050]" pageId="33" pageNumber="32">
<taxonomicName id="1ACE4D46027EFFBD3FCAFBF4FE1AFB84" ID-CoL="8LY2M" authority="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière, 2017" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[192,485,1025,1050]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="arthurensis" status="sp. nov.">
<emphasis id="EFBAEAD7027EFFBD3FCAFBF4FE1AFB84" bold="true" box="[192,485,1025,1050]" italics="true" pageId="33" pageNumber="32">Oopterus arthurensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F48957AC027EFFBD3EE7FBF4FD6DFB84" box="[493,658,1025,1050]" pageId="33" pageNumber="32" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027EFFBE3FCAFBD4FADCFF67" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32" type="description">
<paragraph id="DD7136C5027EFFBD3FCAFBD4FEAFFBA7" blockId="33.[192,658,1025,1082]" box="[192,336,1057,1082]" pageId="33" pageNumber="32">
<figureCitation id="45F52A40027EFFBD3FCAFBD4FEE8FBA4" box="[192,279,1057,1082]" captionStart="Figures 3538" captionStartId="85.[192,288,1829,1851]" captionTargetBox="[214,1428,257,1816]" captionTargetId="figure-45@85.[192,1436,192,1819]" captionTargetPageId="85" captionText="Figures 3538. Dorsal habitus. 35) Oopterus arthurensis new species. 36) O. corvinki new species. 37) O. palmai new species. 38) O. monticola new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5169591" httpUri="https://zenodo.org/record/5169591/files/figure.png" pageId="33" pageNumber="32">Fig. 35</figureCitation>
,
<figureCitation id="45F52A40027EFFBD3E2EFBD4FEAFFBA7" box="[292,336,1058,1082]" captionStart="Figures 118123" captionStartId="101.[192,286,1830,1852]" captionTargetBox="[192,1412,192,1818]" captionTargetId="figure-49@101.[192,1412,192,1818]" captionTargetPageId="101" captionText="Figures 118123. Species distribution maps. 118) Maungazolus septempunctatus new species. 119) M. tararuaensis new species. 120) Oopterus anglemensis new species. 121) O. arthurensis new species. 122) O. clivinoides. 123) O. collaris." figureDoi="http://doi.org/10.5281/zenodo.5169631" httpUri="https://zenodo.org/record/5169631/files/figure.png" pageId="33" pageNumber="32">121</figureCitation>
</paragraph>
<paragraph id="DD7136C5027EFFBD3FCAFB94FC5EFB27" blockId="33.[192,1440,1122,1210]" pageId="33" pageNumber="32">
<materialsCitation id="6DA63C98027EFFBD3FCAFB94FC5EFB27" collectionCode="MONZ, NN, CAVE, T" country="New Zealand" location="Mt. Balloon" municipality="Mount Arthur" pageId="33" pageNumber="32" specimenCount="2" typeStatus="holotype">
<typeStatus id="02758867027EFFBD3FCAFB94FECCFBE7" box="[192,307,1122,1146]" pageId="33" pageNumber="32" type="holotype">Holotype</typeStatus>
: female (
<collectionCode id="BBDFAE00027EFFBD3E94FB94FE0BFBE7" box="[414,500,1122,1146]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="33" pageNumber="32" type="Museum">MONZ</collectionCode>
) labeled: “[
<collectingCountry id="A5D97655027EFFBD3D70FB94FCB2FBE7" box="[634,845,1122,1146]" name="New Zealand" pageId="33" pageNumber="32">NEW ZEALAND</collectingCountry>
<collectionCode id="BBDFAE00027EFFBD3C5EFB94FC7DFBE4" box="[852,898,1122,1145]" pageId="33" pageNumber="32">NN</collectionCode>
]
<location id="D811601E027EFFBD3C85FB94FBE0FBE7" LSID="urn:lsid:plazi:treatment:556787D3027EFFBE3FCAFBF4FAFCFCC7:D811601E027EFFBD3C85FB94FBE0FBE7" box="[911,1055,1122,1146]" country="New Zealand" municipality="Mount Arthur" name="Mt. Balloon" pageId="33" pageNumber="32">Mt. Balloon</location>
[= Balloon Hut,
<collectingMunicipality id="3D15ACBF027EFFBD3BE3FB94FA66FBE7" box="[1257,1433,1122,1146]" pageId="33" pageNumber="32">Mount Arthur</collectingMunicipality>
] IN
<collectionCode id="BBDFAE00027EFFBD3FECFB74FECCFB07" box="[230,307,1154,1178]" country="USA" httpUri="http://grbio.org/cool/wvdq-gi6n" name="DOI/NPS, Carlsbad Caverns National Park" pageId="33" pageNumber="32">CAVE</collectionCode>
J.
<collectionCode id="BBDFAE00027EFFBD3E46FB74FE9BFB04" box="[332,356,1154,1177]" name="Tavera, Department of Geology and Geophysics" pageId="33" pageNumber="32">T</collectionCode>
. Salmon 24.I.48 (hand-written) / [female symbol] /
<typeStatus id="02758867027EFFBD3CCFFB74FB9FFB07" box="[965,1120,1154,1178]" pageId="33" pageNumber="32" type="holotype">HOLOTYPE</typeStatus>
[female symbol]
<taxonomicName id="1ACE4D46027EFFBD3A3BFB74FD48FB27" authority="Larochelle &amp; Lariviere, 2017" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="arthurensis">
<emphasis id="EFBAEAD7027EFFBD3A3BFB74FEB1FB27" italics="true" pageId="33" pageNumber="32">Oopterus arthurensis</emphasis>
Larochelle &amp; Larivière, 2017
</taxonomicName>
(red label; typed).”
</materialsCitation>
</paragraph>
<paragraph id="DD7136C5027EFFBE3FCAFB17FADCFF67" blockId="33.[192,1440,1249,1882]" lastBlockId="34.[192,1440,193,250]" lastPageId="34" lastPageNumber="33" pageId="33" pageNumber="32">
<emphasis id="EFBAEAD7027EFFBD3FCAFB17FE9CFB64" bold="true" box="[192,355,1249,1273]" pageId="33" pageNumber="32">Description</emphasis>
. Body length
<quantity id="1A369B20027EFFBD3D06FB14FD94FB64" box="[524,619,1250,1274]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="33" pageNumber="32" unit="mm" value="7.9">7.9 mm</quantity>
. Head, pronotum, elytra, and abdomen mostly rufous; margins and suture of elytra red; antennae, palpi, and legs testaceous. Microsculpture strong, isodiametric on head, very transverse (with microlines) on pronotum and elytra. Iridescence absent on head, very strong on pronotum and elytra. Very shiny, without metallic luster. Forebody narrow in comparison to elytra (as in
<taxonomicName id="1ACE4D46027EFFBD3FD5FA94FE83FAE7" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[223,380,1378,1402]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="trechoides">
<emphasis id="EFBAEAD7027EFFBD3FD5FA94FE83FAE7" box="[223,380,1378,1402]" italics="true" pageId="33" pageNumber="32">O. trechoides</emphasis>
</taxonomicName>
).
<emphasis id="EFBAEAD7027EFFBD3E99FA94FE23FAE7" bold="true" box="[403,476,1378,1402]" pageId="33" pageNumber="32">Head</emphasis>
. Labrum strongly transverse, slightly emarginate anteriorly. Antennae subfiliform (as in
<emphasis id="EFBAEAD7027EFFBD3E43FA74FD6FFA07" box="[329,656,1410,1434]" italics="true" pageId="33" pageNumber="32">
<taxonomicName id="1ACE4D46027EFFBD3E43FA74FE1FFA07" baseAuthorityName="Broun" baseAuthorityYear="1917" box="[329,480,1410,1434]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="ocularius">O. ocularius</taxonomicName>
,
<taxonomicName id="1ACE4D46027EFFBD3EE1FA74FD74FA07" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[491,651,1410,1434]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="trechoides">O. trechoides</taxonomicName>
,
</emphasis>
and
<taxonomicName id="1ACE4D46027EFFBD3DC0FA74FC91FA07" box="[714,878,1410,1434]" class="Insecta" family="Carabidae" genus="Zolus" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="32" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="EFBAEAD7027EFFBD3DC0FA74FCF1FA07" box="[714,782,1410,1434]" italics="true" pageId="33" pageNumber="32">Zolus</emphasis>
species
</taxonomicName>
), very long: segment 1 (scape) very long, about 2.5x longer than its maximum width. Frontal furrows wide, deep, subparallel. Eyes slightly convex; two setiferous punctures on inner side of each eye. Tempora strongly inflated (about as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula.
<emphasis id="EFBAEAD7027EFFBD3C2AF9F4FC7BF987" bold="true" box="[800,900,1538,1562]" pageId="33" pageNumber="32">Thorax</emphasis>
. Pronotum slightly convex, finely punctate across base, wrinkled in basal half, subquadrate, widest before middle; apex subtruncate; anterolateral angles well developed, obtusely rounded; anterior bead incomplete, obsolete medially; sides moderately rounded, moderately sinuate posteriorly; lateral grooves absent; a single setiferous puncture on each side (anteriorly); posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, finely punctate, very deep and wide, oblong, prolonged forward, simple; posterior bead absent; base emarginate, about as wide as pronotal apex, much narrower than elytral base.
<emphasis id="EFBAEAD7027EFFBD3A12F934FAA7F947" bold="true" box="[1304,1368,1730,1754]" pageId="33" pageNumber="32">Legs</emphasis>
. Very long.
<emphasis id="EFBAEAD7027EFFBD3E0EF914FEA4F967" bold="true" box="[260,347,1762,1786]" pageId="33" pageNumber="32">Elytra</emphasis>
. Moderately convex, subovate, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders oblique. Sides strongly rounded. Scutellar striole present. Stria 1 complete, deep; striae 27 incomplete, obsolete; striae 12 finely punctate; stria 3 with a single setiferous puncture (anteriorly). Recurrent stria long, sharp, directed apically toward stria 6. Subapical seta present. Intervals subdepressed. Sutural apices obtusely rounded.
<emphasis id="EFBAEAD7027DFFBE3C98FF37FBEAFF44" bold="true" box="[914,1045,193,217]" pageId="34" pageNumber="33">Abdomen</emphasis>
. Last visible sternum (sternum VII): male unknown; female with four apical ambulatory setae.
<emphasis id="EFBAEAD7027DFFBE3CC5FF17FBABFF64" bold="true" box="[975,1108,225,249]" pageId="34" pageNumber="33">Aedeagus</emphasis>
. Male unknown.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027DFFBE3FCAFED7FB20FEE4" pageId="34" pageNumber="33" type="materials_examined">
<paragraph id="DD7136C5027DFFBE3FCAFED7FD2DFEA7" blockId="34.[192,722,289,314]" box="[192,722,289,314]" pageId="34" pageNumber="33">
<emphasis id="EFBAEAD7027DFFBE3FCAFED7FE3AFEA4" bold="true" box="[192,453,289,313]" pageId="34" pageNumber="33">Material examined</emphasis>
.
<specimenCount id="CBC8FD4C027DFFBE3EDEFED4FDA2FEA7" box="[468,605,290,314]" pageId="34" pageNumber="33" type="generic">1 specimen</specimenCount>
(
<collectionCode id="BBDFAE00027DFFBE3D66FED4FD39FEA7" box="[620,710,290,314]" country="New Zealand" httpUri="http://grbio.org/cool/15kb-kde6" name="Museum of New Zealand Te Papa Tongarewa - Entomology" pageId="34" pageNumber="33" type="Museum">MONZ</collectionCode>
).
</paragraph>
<paragraph id="DD7136C5027DFFBE3FCAFE97FB20FEE4" blockId="34.[192,1247,353,378]" box="[192,1247,353,378]" pageId="34" pageNumber="33">
<emphasis id="EFBAEAD7027DFFBE3FCAFE97FDEFFEE4" bold="true" box="[192,528,353,377]" pageId="34" pageNumber="33">Geographic distribution</emphasis>
(
<figureCitation id="45F52A40027DFFBE3D2AFE94FD78FEE7" box="[544,647,354,378]" captionStart="Figures 118123" captionStartId="101.[192,286,1830,1852]" captionTargetBox="[192,1412,192,1818]" captionTargetId="figure-49@101.[192,1412,192,1818]" captionTargetPageId="101" captionText="Figures 118123. Species distribution maps. 118) Maungazolus septempunctatus new species. 119) M. tararuaensis new species. 120) Oopterus anglemensis new species. 121) O. arthurensis new species. 122) O. clivinoides. 123) O. collaris." figureDoi="http://doi.org/10.5281/zenodo.5169631" httpUri="https://zenodo.org/record/5169631/files/figure.png" pageId="34" pageNumber="33">Fig. 121</figureCitation>
). South Island:
<collectionCode id="BBDFAE00027DFFBE3C43FE94FC8AFEE4" box="[841,885,354,377]" pageId="34" pageNumber="33">NN</collectionCode>
Mount Arthur, Balloon Hut.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027DFFBE3FCAFE54FAC9FDA7" pageId="34" pageNumber="33" type="biology_ecology">
<paragraph id="DD7136C5027DFFBE3FCAFE54FBA8FE24" blockId="34.[192,1111,418,442]" box="[192,1111,418,442]" pageId="34" pageNumber="33">
<emphasis id="EFBAEAD7027DFFBE3FCAFE54FED1FE27" bold="true" box="[192,302,418,442]" pageId="34" pageNumber="33">Ecology</emphasis>
. Subalpine. A beech forest and red tussock area. Found in a cave.
</paragraph>
<paragraph id="DD7136C5027DFFBE3FCAFE17FB7BFE67" blockId="34.[192,1156,481,506]" box="[192,1156,481,506]" pageId="34" pageNumber="33">
<emphasis id="EFBAEAD7027DFFBE3FCAFE17FED6FE64" bold="true" box="[192,297,481,505]" pageId="34" pageNumber="33">Biology</emphasis>
. Seasonality: January. Predacious (based on mouthpart morphology).
</paragraph>
<paragraph id="DD7136C5027DFFBE3FCAFDD7FAC9FDA7" blockId="34.[192,1334,545,570]" box="[192,1334,545,570]" pageId="34" pageNumber="33">
<emphasis id="EFBAEAD7027DFFBE3FCAFDD7FE5DFDA7" bold="true" box="[192,418,545,570]" pageId="34" pageNumber="33">Dispersal power</emphasis>
. Subapterous (incapable of flight). Fast runner (based on leg morphology).
</paragraph>
</subSubSection>
<subSubSection id="95D4654E027DFFBE3FCAFD94FAFCFCC7" pageId="34" pageNumber="33" type="discussion">
<paragraph id="DD7136C5027DFFBE3FCAFD94FAFCFCC7" blockId="34.[192,1440,610,858]" pageId="34" pageNumber="33">
<emphasis id="EFBAEAD7027DFFBE3FCAFD94FEC3FDE7" bold="true" box="[192,316,610,634]" pageId="34" pageNumber="33">Remarks</emphasis>
. This species is named after the
<typeStatus id="02758867027DFFBE3DC3FD94FD01FDE7" box="[713,766,610,634]" pageId="34" pageNumber="33">type</typeStatus>
locality Mount Arthur (NN) and the Latin suffix -
<emphasis id="EFBAEAD7027DFFBE3A56FD94FA66FDE4" box="[1372,1433,610,633]" italics="true" pageId="34" pageNumber="33">ensis</emphasis>
, denoting a place, locality, or country.
<taxonomicName id="1ACE4D46027DFFBE3D84FD74FC6DFD07" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[654,914,642,666]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="arthurensis">
<emphasis id="EFBAEAD7027DFFBE3D84FD74FC6DFD07" box="[654,914,642,666]" italics="true" pageId="34" pageNumber="33">Oopterus arthurensis</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="1ACE4D46027DFFBE3BE9FD74FA95FD07" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[1251,1386,642,666]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="corvinki">
<emphasis id="EFBAEAD7027DFFBE3BE9FD74FA95FD07" box="[1251,1386,642,666]" italics="true" pageId="34" pageNumber="33">O. corvinki</emphasis>
</taxonomicName>
and has the following distinguishing features: body length
<quantity id="1A369B20027DFFBE3C45FD54FC53FD24" box="[847,940,674,698]" metricMagnitude="-3" metricUnit="m" metricValue="7.9" pageId="34" pageNumber="33" unit="mm" value="7.9">7.9 mm</quantity>
; forebody narrow in comparison to elytra; antennae subfiliform, very long; tempora strongly inflated (about as long as eyes); pronotum subquadrate, not cordate, posterolateral angles subrectangular; elytral striae mostly obsolete. The two species are allopatric:
<taxonomicName id="1ACE4D46027DFFBE3E7BFCF4FDDEFC87" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[369,545,770,794]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="arthurensis">
<emphasis id="EFBAEAD7027DFFBE3E7BFCF4FDDEFC87" box="[369,545,770,794]" italics="true" pageId="34" pageNumber="33">O. arthurensis</emphasis>
</taxonomicName>
is restricted to the northwestern South Island (NN, Mount Arthur), while
<taxonomicName id="1ACE4D46027DFFBE3FCAFCD4FEB7FCA7" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[192,328,802,826]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="corvinki">
<emphasis id="EFBAEAD7027DFFBE3FCAFCD4FEB7FCA7" box="[192,328,802,826]" italics="true" pageId="34" pageNumber="33">O. corvinki</emphasis>
</taxonomicName>
occurs in the southwest of the South Island (FD,
<collectingCountry id="A5D97655027DFFBE3CA4FCD4FB93FCA7" box="[942,1132,802,826]" name="Kiribati" pageId="34" pageNumber="33">Gilbert Islands</collectingCountry>
).
<taxonomicName id="1ACE4D46027DFFBE3B8AFCD4FACDFCA7" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[1152,1330,802,826]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="34" pageNumber="33" phylum="Arthropoda" rank="species" species="arthurensis">
<emphasis id="EFBAEAD7027DFFBE3B8AFCD4FACDFCA7" box="[1152,1330,802,826]" italics="true" pageId="34" pageNumber="33">O. arthurensis</emphasis>
</taxonomicName>
, with its darkly colored body and normally developed eyes, is probably not a regular cave-dweller.
</paragraph>
</subSubSection>
</treatment>
</document>