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<document id="DB8E2761915565B00A0C2A0957BD0622" ID-DOI="10.5281/zenodo.5169575" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169575" ID-ZooBank="89FC75EA-2324-4361-B818-FBA7B7682A00" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628278715662" checkinUser="felipe" docAuthor="Larochelle, Larivière, Marie-Claude, Larochelle &amp; Larivière" docDate="2017" docId="556787D3026DFFAF3FCAFCD4FC3CFCA7" docLanguage="en" docName="InsectaMundi.0594.1-110.pdf" docOrigin="Insecta Mundi 2017 (594)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Oopterus nanus Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière 2017, new species" docType="treatment" docUuid="4D42BAFA-FEA9-4FF3-AD52-A362820DB7F1" docUuidSource="ZooBank" docVersion="5" lastPageNumber="50" masterDocId="A95EFFAB025FFF9C3F0AFFF6FFFFFF9D" masterDocTitle="Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)" masterLastPageNumber="110" masterPageNumber="1" pageNumber="49" updateTime="1699039365375" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="3D7372CC261C69AC89AC5765F15A82CE">Synopsis of the tribe Zolini in New Zealand (Coleoptera: Carabidae)</mods:title>
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<mods:namePart id="96DAA876781CA7848F0558A383BF94C9">Larochelle</mods:namePart>
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<mods:namePart id="A13DEA99FC78B71D9E76830B54672236">Larivière, Marie-Claude</mods:namePart>
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<mods:namePart id="D87CDF2A04B67B49B40A32B7E3EE86F1">Larochelle</mods:namePart>
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<mods:namePart id="C02EDDCBA7832152C46D651E79165C9F">Larivière</mods:namePart>
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<paragraph id="DD7136C5026DFFAE3FCAFCD4FDB9FCA4" blockId="50.[192,582,801,858]" box="[192,582,801,826]" pageId="50" pageNumber="49">
<heading id="863981A9026DFFAE3FCAFCD4FDB9FCA4" bold="true" box="[192,582,801,826]" fontSize="10" level="2" pageId="50" pageNumber="49" reason="3">
<emphasis id="EFBAEAD7026DFFAE3FCAFCD4FDB9FCA4" bold="true" box="[192,582,801,826]" pageId="50" pageNumber="49">
<taxonomicName id="1ACE4D46026DFFAE3FCAFCD4FE66FCA4" ID-CoL="8LY2V" authority="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière, 2017" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[192,409,802,826]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="nanus" status="sp. nov.">
<emphasis id="EFBAEAD7026DFFAE3FCAFCD4FE66FCA4" bold="true" box="[192,409,802,826]" italics="true" pageId="50" pageNumber="49">Oopterus nanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F48957AC026DFFAE3EABFCD4FDB9FCA4" box="[417,582,801,826]" pageId="50" pageNumber="49" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95D4654E026DFFAE3FCAFCB4FC69F8C4" pageId="50" pageNumber="49" type="description">
<paragraph id="DD7136C5026DFFAE3FCAFCB4FE83FCC7" blockId="50.[192,582,801,858]" box="[192,380,833,858]" pageId="50" pageNumber="49">
<figureCitation id="45F52A40026DFFAE3FCAFCB4FEE8FCC4" box="[192,279,833,858]" captionStart="Figures 4750" captionStartId="88.[192,288,1832,1854]" captionTargetBox="[214,1427,204,1816]" captionTargetId="figure-48@88.[192,1435,192,1818]" captionTargetPageId="88" captionText="Figures 4750. Dorsal habitus. 47) Oopterus ocularius. 48) O. marrineri. 49) O. anglemensis new species. 50) O. nanus new species. Scale line = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5169599" httpUri="https://zenodo.org/record/5169599/files/figure.png" pageId="50" pageNumber="49">Fig. 50</figureCitation>
,
<figureCitation id="45F52A40026DFFAE3E2EFCB4FEBBFCC7" box="[292,324,834,858]" captionStart="Figures 9097" captionStartId="96.[192,288,1830,1852]" captionTargetBox="[248,1407,248,1800]" captionTargetId="figure-62@96.[192,1435,192,1818]" captionTargetPageId="96" captionText="Figures 9097. Aedeagi, lateral. 90) Oopterus trechoides new species. 91) O. sculpturatus. 92) O. taieriensis new species. 93) O. ocularius. 94) O. marrineri. 95) O. anglemensis new species. 96) O. nanus new species. 97) O. garnerae new species." figureDoi="http://doi.org/10.5281/zenodo.5169619" httpUri="https://zenodo.org/record/5169619/files/figure.png" pageId="50" pageNumber="49">96</figureCitation>
,
<figureCitation id="45F52A40026DFFAE3E5AFCB4FE83FCC7" box="[336,380,834,858]" captionStart="Figures 136141" captionStartId="104.[192,288,1829,1851]" captionTargetBox="[192,1411,192,1817]" captionTargetId="figure-54@104.[192,1411,192,1817]" captionTargetPageId="104" captionText="Figures 136141. Species distribution maps. 136) Oopterus nanus new species. 137) O. nunni new species. 138) O. ocularius. 139) O. palmai new species. 140) O. patulus. 141) O. punctatus new species." figureDoi="http://doi.org/10.5281/zenodo.5169637" httpUri="https://zenodo.org/record/5169637/files/figure.png" pageId="50" pageNumber="49">136</figureCitation>
</paragraph>
<paragraph id="DD7136C5026DFFAE3FCAFC74FCADFC67" blockId="50.[192,1440,897,1018]" pageId="50" pageNumber="49">
<materialsCitation id="6DA63C98026DFFAE3FCAFC74FCF8FC47" collectionCode="NZAC" collectorName="J. S. Dugdale" country="New Zealand" elevation="1615" location="NEW ZEALAND" pageId="50" pageNumber="49" specimenCount="2" typeStatus="holotype">
<typeStatus id="02758867026DFFAE3FCAFC74FECCFC07" box="[192,307,898,922]" pageId="50" pageNumber="49" type="holotype">Holotype</typeStatus>
: male (
<collectionCode id="BBDFAE00026DFFAE3E81FC74FE25FC07" box="[395,474,898,922]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="50" pageNumber="49" type="Other institutional type">NZAC</collectionCode>
) labeled: “[
<collectingCountry id="A5D97655026DFFAE3D6EFC74FCC6FC07" box="[612,825,898,922]" name="New Zealand" pageId="50" pageNumber="49">NEW ZEALAND</collectingCountry>
CO] Old Man Ra CO
<quantity id="1A369B20026DFFAE3B47FC77FB5FFC04" box="[1101,1184,897,921]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" pageId="50" pageNumber="49" unit="m" value="1615.0">
<elevation id="56E3D1F6026DFFAE3B47FC77FB5FFC04" box="[1101,1184,897,921]" metricMagnitude="3" metricUnit="m" metricValue="1.6149999999999998" pageId="50" pageNumber="49" unit="m" value="1615.0">1615m</elevation>
</quantity>
26 Feb 74
<collectorName id="703B5313026DFFAE3A27FC74FF0BFC27" pageId="50" pageNumber="49">J.S. Dugdale</collectorName>
(hand-written) / ex Celmisia haastii (hand-written) /
<typeStatus id="02758867026DFFAE3C88FC54FBE2FC27" box="[898,1053,930,954]" pageId="50" pageNumber="49" type="holotype">HOLOTYPE</typeStatus>
[male symbol]
<taxonomicName id="1ACE4D46026DFFAE3BD4FC54FDE0FC47" authority="Larochelle &amp; Lariviere, 2017" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="50" pageNumber="49" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="EFBAEAD7026DFFAE3BD4FC54FA5FFC24" box="[1246,1440,930,953]" italics="true" pageId="50" pageNumber="49">Oopterus nanus</emphasis>
Larochelle &amp; Larivière, 2017
</taxonomicName>
(red label; typed).”
</materialsCitation>
<typeStatus id="02758867026DFFAE3C07FC34FC71FC47" box="[781,910,962,986]" pageId="50" pageNumber="49" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="CBC8FD4C026DFFAE3C92FC34FBD5FC47" box="[920,1066,962,986]" pageId="50" pageNumber="49" type="female">two females</specimenCount>
(LUNZ, NZAC) from the same locality as the
<typeStatus id="02758867026DFFAE3E79FC14FE20FC67" box="[371,479,994,1018]" pageId="50" pageNumber="49" type="holotype">holotype</typeStatus>
, bearing blue
<typeStatus id="02758867026DFFAE3D86FC14FD05FC67" box="[652,762,994,1018]" pageId="50" pageNumber="49" type="paratype">paratype</typeStatus>
labels.
</paragraph>
<paragraph id="DD7136C5026DFFAE3FCAFBD7FC69F8C4" blockId="50.[192,1441,1057,1882]" pageId="50" pageNumber="49">
<emphasis id="EFBAEAD7026DFFAE3FCAFBD7FE9CFBA4" bold="true" box="[192,355,1057,1081]" pageId="50" pageNumber="49">Description</emphasis>
. Body length
<quantity id="1A369B20026DFFAE3D07FBD7FD5FFBA4" box="[525,672,1057,1081]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.2" metricValueMin="3.9" pageId="50" pageNumber="49" unit="mm" value="4.55" valueMax="5.2" valueMin="3.9">3.95.2 mm</quantity>
. Head, pronotum, elytra, and abdomen mostly black; abdomen reddish piceous laterally and apically; antennae, palpi, and legs reddish. Microsculpture strong and isodiametric on head, obsolete and moderately transverse on pronotum, feeble and isodiametric on elytra. Iridescence absent. Very shiny, without metallic luster.
<emphasis id="EFBAEAD7026DFFAE3C76FB74FC3AFB07" bold="true" box="[892,965,1154,1178]" pageId="50" pageNumber="49">Head</emphasis>
. Labrum strongly transverse, slightly emarginate anteriorly. Antennae submoniliform, short: segment 1 (scape) moderately long, about 2x longer than its maximum width. Frontal furrows wide, deep, convergent. Eyes moderately convex; two setiferous punctures on inner side of each eye. Tempora moderately inflated (about half as long as eyes). Mentum: medial tooth entire, acute apically, moderately shorter than lateral lobes. Paraglossae membranous, prominent, slightly longer than ligula.
<emphasis id="EFBAEAD7026DFFAE3C16FAD4FC7FFAA7" bold="true" box="[796,896,1314,1338]" pageId="50" pageNumber="49">Thorax</emphasis>
. Pronotum moderately convex, impunctate, wrinkled discally, strongly transverse, widest before middle; apex subtruncate; anterolateral angles poorly developed, rounded; anterior bead incomplete, obsolete medially; sides strongly rounded, not sinuate posteriorly; lateral grooves absent; two setiferous punctures on each side; posterolateral angles subrectangular; posterolateral carinae absent; laterobasal foveae well defined, impunctate, shallow, very wide, rectangular, not prolonged forward, double; posterior bead absent; base emarginate, much narrower than pronotal apex, slightly narrower than elytral base.
<emphasis id="EFBAEAD7026DFFAE3CE2FA14FBD7FA67" bold="true" box="[1000,1064,1506,1530]" pageId="50" pageNumber="49">Legs</emphasis>
. Short.
<emphasis id="EFBAEAD7026DFFAE3B83FA14FB1FFA67" bold="true" box="[1161,1248,1506,1530]" pageId="50" pageNumber="49">Elytra</emphasis>
. Subdepressed, oblong, widest about middle. Basal margin incomplete, reaching about stria 4. Shoulders obtuse. Sides subparallel. Scutellar striole obsolete. Striae complete, shallow, finely punctate; stria 3 with four unusually large setiferous punctures. Recurrent stria long, sharp, directed apically toward stria 5, 6 or 7. Subapical seta present. Intervals depressed. Sutural apices obtuse.
<emphasis id="EFBAEAD7026DFFAE3B74F997FAFEF9E4" bold="true" box="[1150,1281,1633,1657]" pageId="50" pageNumber="49">Abdomen</emphasis>
. Last visible sternum (sternum VII): male with two apical ambulatory setae; female with four apical ambulatory setae.
<emphasis id="EFBAEAD7026DFFAE3E1BF957FE69F924" bold="true" box="[273,406,1697,1721]" pageId="50" pageNumber="49">Aedeagus</emphasis>
. Lateral view (
<figureCitation id="45F52A40026DFFAE3D5FF954FD50F927" box="[597,687,1698,1722]" captionStart="Figures 9097" captionStartId="96.[192,288,1830,1852]" captionTargetBox="[248,1407,248,1800]" captionTargetId="figure-62@96.[192,1435,192,1818]" captionTargetPageId="96" captionText="Figures 9097. Aedeagi, lateral. 90) Oopterus trechoides new species. 91) O. sculpturatus. 92) O. taieriensis new species. 93) O. ocularius. 94) O. marrineri. 95) O. anglemensis new species. 96) O. nanus new species. 97) O. garnerae new species." figureDoi="http://doi.org/10.5281/zenodo.5169619" httpUri="https://zenodo.org/record/5169619/files/figure.png" pageId="50" pageNumber="49">Fig. 96</figureCitation>
): strongly arcuate, narrowed in apical half; base moderately biconvex dorsally; middle strongly convex dorsally, slightly convex ventrally, with dorsal membranous area very wide and moderately long; apex slender, strongly concave dorsally, slightly concave ventrally, with extreme tip wide, unusually long. Dorsal view: narrow, asymmetrical (ostium of membranous area deflected to right); apex straight; basal orifice narrow, closed anteriorly, moderately distant from membranous area. Parameres with four to six apical setae.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E026CFFAF3FCAFF37FC51FE87" pageId="51" pageNumber="50" type="materials_examined">
<paragraph id="DD7136C5026CFFAF3FCAFF37FBF4FF47" blockId="51.[192,1035,193,218]" box="[192,1035,193,218]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFF37FE3AFF44" bold="true" box="[192,453,193,217]" pageId="51" pageNumber="50">Material examined</emphasis>
.
<specimenCount id="CBC8FD4C026CFFAF3EDEFF34FD78FF47" box="[468,647,194,218]" pageId="51" pageNumber="50" type="generic">196 specimens</specimenCount>
(
<collectionCode id="BBDFAE00026CFFAF3D9CFF34FD12FF47" box="[662,749,194,218]" country="New Zealand" httpUri="http://grbio.org/cool/76et-m1kt" name="Auckland Institute and Museum" pageId="51" pageNumber="50" type="Museum">AMNZ</collectionCode>
,
<collectionCode id="BBDFAE00026CFFAF3DF2FF34FCB9FF44" box="[760,838,194,217]" pageId="51" pageNumber="50">JNNZ</collectionCode>
,
<collectionCode id="BBDFAE00026CFFAF3C58FF34FC5CFF44" box="[850,931,194,217]" country="New Zealand" httpUri="http://grbio.org/cool/n5zm-bhpt" name="Lincoln University Entomology Research Museum" pageId="51" pageNumber="50">LUNZ</collectionCode>
,
<collectionCode id="BBDFAE00026CFFAF3CA4FF34FBFFFF47" box="[942,1024,194,218]" country="New Zealand" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34070" lsid="urn:lsid:biocol.org:col:34070" name="New Zealand Arthropod Collection" pageId="51" pageNumber="50" type="Other institutional type">NZAC</collectionCode>
).
</paragraph>
<paragraph id="DD7136C5026CFFAF3FCAFEF7FC51FE87" blockId="51.[192,942,257,282]" box="[192,942,257,282]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFEF7FDEFFE84" bold="true" box="[192,528,257,281]" pageId="51" pageNumber="50">Geographic distribution</emphasis>
(
<figureCitation id="45F52A40026CFFAF3D2AFEF4FD77FE87" box="[544,648,258,282]" captionStart="Figures 136141" captionStartId="104.[192,288,1829,1851]" captionTargetBox="[192,1411,192,1817]" captionTargetId="figure-54@104.[192,1411,192,1817]" captionTargetPageId="104" captionText="Figures 136141. Species distribution maps. 136) Oopterus nanus new species. 137) O. nunni new species. 138) O. ocularius. 139) O. palmai new species. 140) O. patulus. 141) O. punctatus new species." figureDoi="http://doi.org/10.5281/zenodo.5169637" httpUri="https://zenodo.org/record/5169637/files/figure.png" pageId="51" pageNumber="50">Fig. 136</figureCitation>
). South Island: CO, OL.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E026CFFAF3FCAFEB4FC65FDC7" pageId="51" pageNumber="50" type="biology_ecology">
<paragraph id="DD7136C5026CFFAF3FCAFEB4FCB0FEE7" blockId="51.[192,1440,322,474]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFEB4FED1FEC7" bold="true" box="[192,302,322,346]" pageId="51" pageNumber="50">Ecology</emphasis>
. Alpine. Epigean. Fellfields; tussock grasslands. Open ground. Nocturnal; hides during the day under stones and in mat plant litter. Gregarious.
</paragraph>
<paragraph id="DD7136C5026CFFAF3FCAFE57FE9EFE47" blockId="51.[192,1440,322,474]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFE57FED6FE24" bold="true" box="[192,297,417,441]" pageId="51" pageNumber="50">Biology</emphasis>
. Seasonality: NovemberApril. Tenerals: JanuaryFebruary. Predacious (based on mouthpart morphology).
</paragraph>
<paragraph id="DD7136C5026CFFAF3FCAFDF7FC0EFD84" blockId="51.[192,1009,513,602]" box="[192,1009,513,538]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFDF7FE5DFD87" bold="true" box="[192,418,513,538]" pageId="51" pageNumber="50">Dispersal power</emphasis>
. Subapterous (incapable of flight). Slow runner.
</paragraph>
<paragraph id="DD7136C5026CFFAF3FCAFDB7FC65FDC7" blockId="51.[192,1009,513,602]" box="[192,922,577,602]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFDB7FE12FDC4" bold="true" box="[192,493,577,601]" pageId="51" pageNumber="50">Collecting techniques</emphasis>
. Lifting stones; sifting plant litter.
</paragraph>
</subSubSection>
<subSubSection id="95D4654E026CFFAF3FCAFD74FC3CFCA7" pageId="51" pageNumber="50" type="discussion">
<paragraph id="DD7136C5026CFFAF3FCAFD74FC3CFCA7" blockId="51.[192,1440,642,826]" pageId="51" pageNumber="50">
<emphasis id="EFBAEAD7026CFFAF3FCAFD74FEC3FD07" bold="true" box="[192,316,642,666]" pageId="51" pageNumber="50">Remarks</emphasis>
. The species name is based on the Latin noun
<taxonomicName id="1ACE4D46026CFFAF3C60FD74FC49FD04" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[874,950,642,665]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="50" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="EFBAEAD7026CFFAF3C60FD74FC49FD04" box="[874,950,642,665]" italics="true" pageId="51" pageNumber="50">nanus</emphasis>
</taxonomicName>
, denoting a dwarf, referring to the small size of the beetle.
<taxonomicName id="1ACE4D46026CFFAF3E90FD54FDA3FD24" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[410,604,674,697]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="50" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="EFBAEAD7026CFFAF3E90FD54FDA3FD24" box="[410,604,674,697]" italics="true" pageId="51" pageNumber="50">Oopterus nanus</emphasis>
</taxonomicName>
is morphologically close to
<taxonomicName id="1ACE4D46026CFFAF3CA1FD54FB99FD27" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[939,1126,674,698]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="50" phylum="Arthropoda" rank="species" species="anglemensis">
<emphasis id="EFBAEAD7026CFFAF3CA1FD54FB99FD27" box="[939,1126,674,698]" italics="true" pageId="51" pageNumber="50">O. anglemensis</emphasis>
</taxonomicName>
. In addition to diagnostic characters of the male genitalia,
<taxonomicName id="1ACE4D46026CFFAF3D43FD34FD49FD44" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[585,694,706,729]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="50" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="EFBAEAD7026CFFAF3D43FD34FD49FD44" box="[585,694,706,729]" italics="true" pageId="51" pageNumber="50">O. nanus</emphasis>
</taxonomicName>
has the following distinguishing features: body length
<quantity id="1A369B20026CFFAF3A42FD37FF0DFD64" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.2" metricValueMin="3.9" pageId="51" pageNumber="50" unit="mm" value="4.55" valueMax="5.2" valueMin="3.9">3.95.2 mm</quantity>
; pronotum not cordate, laterobasal foveae well defined, rectangular; elytra subdepressed, oblong, sides subparallel. The two species are allopatric:
<taxonomicName id="1ACE4D46026CFFAF3C19FCF4FC7DFC84" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[787,898,770,793]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="50" phylum="Arthropoda" rank="species" species="nanus">
<emphasis id="EFBAEAD7026CFFAF3C19FCF4FC7DFC84" box="[787,898,770,793]" italics="true" pageId="51" pageNumber="50">O. nanus</emphasis>
</taxonomicName>
occurs in southern areas of the South Island (CO, OL), while
<taxonomicName id="1ACE4D46026CFFAF3E82FCD4FDBCFCA7" authorityName="Larochelle &amp; Larivière &amp; Larochelle &amp; Larivière" authorityYear="2017" box="[392,579,802,826]" class="Insecta" family="Carabidae" genus="Oopterus" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="50" phylum="Arthropoda" rank="species" species="anglemensis">
<emphasis id="EFBAEAD7026CFFAF3E82FCD4FDBCFCA7" box="[392,579,802,826]" italics="true" pageId="51" pageNumber="50">O. anglemensis</emphasis>
</taxonomicName>
is restricted to Stewart Island.
</paragraph>
</subSubSection>
</treatment>
</document>