294 lines
37 KiB
XML
294 lines
37 KiB
XML
<document id="225DF6E7601E224FA3C6AF34EADFAEDF" ID-ISSN="1536-9307" ID-ZooBank="85D76447-0085-4EDB-BEC0-434DBBA867A2" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638240558011" checkinUser="felipe" docAuthor="Kovařík, František" docDate="2013" docId="5566CB15822EFFAA5298F94CFE93F916" docLanguage="en" docName="Euscorpius.175.1-9.pdf" docOrigin="Euscorpius 175" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Thaicharmus indicus Kovarik 2013, sp. n." docType="treatment" docUuid="4B53DD8B-353D-4F2E-A33B-9EFDFB7FDB21" docUuidSource="ZooBank" docVersion="2" lastPageNumber="8" masterDocId="A95FB36D822CFFA3510BFF90FFC4FF8F" masterDocTitle="A review of Thaicharmus Kovařík, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones: Buthidae)" masterLastPageNumber="9" masterPageNumber="1" pageNumber="1" updateTime="1699434968930" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="15C725971A752A04D580320668F5B568">A review of Thaicharmus Kovařík, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones: Buthidae)</mods:title>
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<mods:name id="50194B38905E211E74E2AF2114441A32" type="personal">
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<mods:roleTerm id="EF0D9763DAEB990DD69F502E4260591D">Author</mods:roleTerm>
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<mods:namePart id="5F922BA5B943A4CE0327BDE8DEF6B03A">Kovařík, František</mods:namePart>
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<mods:typeOfResource id="71EAA2AF20C3B315D514C7E391DD7AF2">text</mods:typeOfResource>
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<mods:title id="87065BA8322BE24EE8FB127BC1912449">Euscorpius</mods:title>
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<mods:part id="309A76ECB2FC86EB438DCD6E322032FE">
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<mods:date id="446E2F650D844AE761743AD9EBA270D7">2013</mods:date>
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<mods:detail id="837457E0C6F4D630DE019CF9DBD20266" type="volume">
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<mods:number id="B72B63F425E4FCC317BEFBE3CD3CD9C6">175</mods:number>
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<subSubSection id="95D52988822EFFA15298F94CFA88F979" box="[915,1356,1756,1782]" pageId="2" pageNumber="1" type="nomenclature">
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<paragraph id="DD707A03822EFFA15298F94CFA88F979" blockId="2.[831,1440,1756,1884]" box="[915,1356,1756,1782]" pageId="2" pageNumber="1">
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<heading id="8638CD6F822EFFA15298F94CFA88F979" box="[915,1356,1756,1782]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1" reason="2">
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<taxonomicName id="1ACF0180822EFFA15298F94CFB3FF979" ID-CoL="55YSX" authority="Kovarik" authorityName="Kovarik" authorityYear="2013" box="[915,1275,1756,1782]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
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<emphasis id="EFBBA611822EFFA15298F94CFB51F979" bold="true" box="[915,1173,1756,1782]" italics="true" pageId="2" pageNumber="1">Thaicharmus indicus</emphasis>
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<emphasis id="EFBBA611822EFFA15596F94EFB3FF979" bold="true" box="[1181,1275,1758,1782]" pageId="2" pageNumber="1">Kovařík</emphasis>
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</taxonomicName>
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<emphasis id="EFBBA611822EFFA155F0F94CFA88F979" bold="true" box="[1275,1356,1756,1782]" pageId="2" pageNumber="1">
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,
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<taxonomicNameLabel id="F4881B6A822EFFA15401F94CFA88F979" box="[1290,1356,1756,1782]" pageId="2" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="95D52988822EFFA1550DF96FFA8AF8D3" pageId="2" pageNumber="1" type="description">
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<paragraph id="DD707A03822EFFA1550DF96FFA64F8B5" blockId="2.[831,1440,1756,1884]" pageId="2" pageNumber="1">
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(
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<figureCitation id="45F46686822EFFA15505F96FFB46F898" box="[1038,1154,1791,1815]" captionStart-0="Figures 1–7" captionStart-1="Figures 8–12" captionStart-2="Figures 13–14" captionStartId-0="3.[192,278,1312,1336]" captionStartId-1="4.[192,278,1344,1368]" captionStartId-2="5.[192,278,1278,1302]" captionTargetBox-0="[203,1436,170,1268]" captionTargetBox-1="[202,1437,194,1312]" captionTargetBox-2="[196,1378,180,1216]" captionTargetId-0="figure-297@3.[192,1440,168,1283]" captionTargetId-1="figure-323@4.[192,1440,192,1315]" captionTargetId-2="figure-361@5.[192,1440,168,1248]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figures 1–7: Thaicharmus indicus sp. n., Figures 1–2, 7: ♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7). Figures 3–6. ♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6)." captionText-1="Figures 8–12: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 9–12. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." captionText-2="Figures 13–14: Thaicharmus indicus sp. n., Figure 13: ♀ (34 mm) holotype, carapace with mesosomal segments I–II, chelicerae, trochanter and part of femur of pedipalp. Figure 14: ♂ (28.5 mm) paratype, telson with fifth and part of fourth metasomal segments." pageId="2" pageNumber="1">Figs. 1–14</figureCitation>
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,
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<figureCitation id="45F46686822EFFA15586F96FFB15F898" box="[1165,1233,1791,1815]" captionStart-0="Figures 25–26" captionStart-1="Figures 27–29" captionStart-2="Figure 30" captionStartId-0="7.[192,278,1886,1910]" captionStartId-1="8.[192,278,1883,1907]" captionStartId-2="9.[192,267,983,1007]" captionTargetBox-0="[211,1417,191,1855]" captionTargetBox-1="[214,1405,191,1854]" captionTargetBox-2="[191,1440,191,953]" captionTargetId-0="figure-23@7.[212,1416,192,1854]" captionTargetId-1="figure-23@8.[215,1404,192,1853]" captionTargetId-2="figure-501@9.[192,1439,192,953]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionTargetPageId-2="9" captionText-0="Figures 25–26: Thaicharmus indicus sp. n., ♀ (34 mm) holotype." captionText-1="Figures 27–29: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype." captionText-2="Figure 30: India, Goa State, near Chaudi, type locality of Thaicharmus indicus sp." pageId="2" pageNumber="1">25–30</figureCitation>
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)
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<emphasis id="EFBBA611822EFFA15234F8B2FA64F8B5" bold="true" box="[831,1440,1826,1850]" pageId="2" pageNumber="1">http://zoobank.org/urn:lsid:zoobank.org:act:4B53DD</emphasis>
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</paragraph>
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<paragraph id="DD707A03822EFFA1529AF8D4FA8AF8D3" blockId="2.[831,1440,1756,1884]" box="[913,1358,1860,1884]" pageId="2" pageNumber="1">
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<emphasis id="EFBBA611822EFFA1529AF8D4FA8AF8D3" bold="true" box="[913,1358,1860,1884]" pageId="2" pageNumber="1">8B-353D-4F2E-A33B-9EFDFB7FDB21</emphasis>
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</paragraph>
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</subSubSection>
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<caption id="89B02A8B822FFFA051CBFAB0FE05FAE6" pageId="3" pageNumber="2" startId="3.[192,278,1312,1336]" targetBox="[203,1436,170,1268]" targetPageId="3">
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<paragraph id="DD707A03822FFFA051CBFAB0FE05FAE6" blockId="3.[192,1440,1312,1385]" pageId="3" pageNumber="2">
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<emphasis id="EFBBA611822FFFA051CBFAB0FDADFAB8" bold="true" box="[192,617,1312,1336]" pageId="3" pageNumber="2">
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Figures 1–7:
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<taxonomicName id="1ACF0180822FFFA0505DFAB2FDEFFAB8" authority="Kovařík, 2013" authorityName="Kovařík" authorityYear="2013" box="[342,555,1314,1335]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
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<emphasis id="EFBBA611822FFFA0505DFAB2FDEFFAB8" bold="true" box="[342,555,1314,1335]" italics="true" pageId="3" pageNumber="2">Thaicharmus indicus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="F4881B6A822FFFA05339FAB2FDADFAB8" box="[562,617,1314,1335]" pageId="3" pageNumber="2" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
,
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<emphasis id="EFBBA611822FFFA0537DFAB2FCD4FAB8" bold="true" box="[630,784,1314,1335]" pageId="3" pageNumber="2">Figures 1–2, 7:</emphasis>
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♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7).
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<emphasis id="EFBBA611822FFFA050C5FAAAFDDFFAC0" bold="true" box="[462,539,1338,1359]" pageId="3" pageNumber="2">Figures</emphasis>
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3
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<emphasis id="EFBBA611822FFFA05324FAAAFD89FAC0" bold="true" box="[559,589,1338,1359]" pageId="3" pageNumber="2">–6.</emphasis>
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||
♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6).
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</paragraph>
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</caption>
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<subSubSection id="95D52988822FFFA051CBFA0DFED8F9C1" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="DD707A03822FFFA051CBFA0DFD64FA5C" blockId="3.[192,801,1437,1491]" pageId="3" pageNumber="2">
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<materialsCitation id="6DA7705E822FFFA051CBFA0DFDA4FA3B" box="[192,608,1437,1460]" collectionCode="TYPE, LOCALITY, AND, DEPOSITORY" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
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<collectionCode id="BBDEE2C6822FFFA051CBFA0DFE58FA3B" box="[192,412,1437,1460]" pageId="3" pageNumber="2">TYPE LOCALITY AND</collectionCode>
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<typeStatus id="0274C4A1822FFFA050A9FA31FE13FA3B" box="[418,471,1441,1460]" pageId="3" pageNumber="2">TYPE</typeStatus>
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<collectionCode id="BBDEE2C6822FFFA050D7FA30FDA4FA3B" box="[476,608,1440,1460]" pageId="3" pageNumber="2">DEPOSITORY</collectionCode>
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</materialsCitation>
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.
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<materialsCitation id="6DA7705E822FFFA05365FA0DFD59FA5C" collectionCode="FKCP" country="India" location="Goa State" municipality="Chaudi" pageId="3" pageNumber="2" specimenCount="1" typeStatus="holotype">
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<collectingCountry id="A5D83A93822FFFA05365FA0DFD63FA3A" box="[622,679,1437,1461]" name="India" pageId="3" pageNumber="2">India</collectingCountry>
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,
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<location id="D8102CD8822FFFA053B9FA0DFCD9FA3A" LSID="urn:lsid:plazi:treatment:5566CB15822EFFAA5298F94CFE93F916:D8102CD8822FFFA053B9FA0DFCD9FA3A" box="[690,797,1437,1461]" country="India" municipality="Chaudi" name="Goa State" pageId="3" pageNumber="2">Goa State</location>
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||
, near
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||
<collectingMunicipality id="3D14E079822FFFA051FFFA2BFE80FA5C" box="[244,324,1467,1491]" pageId="3" pageNumber="2">Chaudi</collectingMunicipality>
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||
; the author´s collection (
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<collectionCode id="BBDEE2C6822FFFA05347FA2BFD51FA5C" box="[588,661,1467,1491]" pageId="3" pageNumber="2">FKCP</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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<paragraph id="DD707A03822FFFA051CBFA69FED8F9C1" blockId="3.[192,802,1529,1614]" pageId="3" pageNumber="2">
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<materialsCitation id="6DA7705E822FFFA051CBFA69FD62F9A0" collectingDate="2012-11" collectionCode="TYPE, MATERIAL" country="India" location="Goa State" municipality="Chaudi" pageId="3" pageNumber="2" specimenCount="2" specimenCount-female="1" specimenCount-male="1" typeStatus="paratype">
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<collectionCode id="BBDEE2C6822FFFA051CBFA69FEB1F99F" box="[192,373,1529,1552]" pageId="3" pageNumber="2">TYPE MATERIAL</collectionCode>
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.
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<collectingCountry id="A5D83A93822FFFA05081FA69FE07F99E" box="[394,451,1529,1553]" name="India" pageId="3" pageNumber="2">India</collectingCountry>
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,
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<location id="D8102CD8822FFFA050DDFA69FD8EF99E" LSID="urn:lsid:plazi:treatment:5566CB15822EFFAA5298F94CFE93F916:D8102CD8822FFFA050DDFA69FD8EF99E" box="[470,586,1529,1553]" country="India" municipality="Chaudi" name="Goa State" pageId="3" pageNumber="2">Goa State</location>
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, near
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||
<collectingMunicipality id="3D14E079822FFFA05390FA69FD2EF99E" box="[667,746,1529,1553]" pageId="3" pageNumber="2">Chaudi</collectingMunicipality>
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,
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<date id="A9715CC3822FFFA053F5FA69FF3CF9A0" pageId="3" pageNumber="2" value="2012-11">
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<collectingDate id="B935A52B822FFFA053F5FA69FF3CF9A0" pageId="3" pageNumber="2" value="2012-11">XI. 2012</collectingDate>
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</date>
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,
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<specimenCount id="CBC9B18A822FFFA05004F987FEF5F9A0" box="[271,305,1559,1583]" pageId="3" pageNumber="2" type="male">1♂</specimenCount>
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(
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||
<typeStatus id="0274C4A1822FFFA05040F987FE6EF9A0" box="[331,426,1559,1583]" pageId="3" pageNumber="2" type="paratype">paratype</typeStatus>
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)
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||
<specimenCount id="CBC9B18A822FFFA050CFF987FE21F9A0" box="[452,485,1559,1583]" pageId="3" pageNumber="2" type="female">1♀</specimenCount>
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||
(
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<typeStatus id="0274C4A1822FFFA0530BF987FDA7F9A0" box="[512,611,1559,1583]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
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||
), leg
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||
</materialsCitation>
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||
.
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<materialsCitation id="6DA7705E822FFFA053B7F988FCD9F9A0" box="[700,797,1560,1583]" collectionCode="V" country="India" location="Fura" pageId="3" pageNumber="2" specimenCount="1" typeStatus="paratype">
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||
<collectionCode id="BBDEE2C6822FFFA053B7F988FD16F9A0" box="[700,722,1560,1583]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="3" pageNumber="2" type="Museum">V</collectionCode>
|
||
.
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||
<location id="D8102CD8822FFFA053E2F988FCD9F9A0" LSID="urn:lsid:plazi:treatment:5566CB15822EFFAA5298F94CFE93F916:D8102CD8822FFFA053E2F988FCD9F9A0" box="[745,797,1560,1583]" country="India" name="Fura" pageId="3" pageNumber="2">Fura</location>
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||
</materialsCitation>
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||
, (
|
||
<collectionCode id="BBDEE2C6822FFFA051C3F9A6FED5F9C1" box="[200,273,1590,1614]" pageId="3" pageNumber="2">FKCP</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95D52988822FFFA051CBF9E4FD3FF904" box="[192,763,1651,1676]" pageId="3" pageNumber="2" type="etymology">
|
||
<paragraph id="DD707A03822FFFA051CBF9E4FD3FF904" blockId="3.[192,763,1651,1676]" box="[192,763,1651,1676]" pageId="3" pageNumber="2">ETYMOLOGY. Named after the country of occurrence.</paragraph>
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||
</subSubSection>
|
||
<subSubSection id="95D52988822FFFA051CBF921FAFDF946" pageId="3" pageNumber="2" type="diagnosis">
|
||
<paragraph id="DD707A03822FFFA051CBF921FAFDF946" blockId="3.[192,802,1713,1890]" lastBlockId="3.[831,1441,1437,1737]" pageId="3" pageNumber="2">
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||
DIAGNOSIS. Total length
|
||
<quantity id="1A37D7E6822FFFA050EAF921FDB5F947" box="[481,625,1713,1737]" metricMagnitude="-2" metricUnit="m" metricValue="3.125" metricValueMax="3.4" metricValueMin="2.85" pageId="3" pageNumber="2" unit="mm" value="31.25" valueMax="34.0" valueMin="28.5">28.5–34 mm</quantity>
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||
. Sexual dimorphism manifested in narrower male segments of metasoma and pedipalps. Mesosoma black with brown ornament, metasoma and femur of pedipalp black, chela of pedipalp and telson reddish brown, tibia of legs yellowish brown. Chelicerae yellow, with black reticulation. Carapace without carinae. Metasoma, telson and pedipalps densely hirsute, hairs shorter on pedipalp than on metasoma. First and second metasomal segments almost smooth and punctate, fourth and fifth metasomal segments and telson densely granulated. Movable and fixed fingers of pedipalps bear 10 rows of granules, with internal accessory granules and six distal granules of which three are smaller and form an apical row. Manus to movable finger length ratio 0.7–0.75. Pectinal teeth number
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||
<quantity id="1A37D7E6822FFFA0529CF921FC08F946" box="[919,972,1713,1737]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="3" pageNumber="2" unit="in" value="15.0">15 in</quantity>
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||
female. Sternites without carinae.
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||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95D52988822FFFAA5234F97FFCDBFA46" lastPageId="9" lastPageNumber="8" pageId="3" pageNumber="2" type="description">
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<paragraph id="DD707A03822FFFA75234F97FFDB8FA55" blockId="3.[831,1440,1774,1890]" lastBlockId="4.[192,808,1444,1897]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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||
DESCRIPTION. The adult male is
|
||
<quantity id="1A37D7E6822FFFA05595F97EFB39F889" box="[1182,1277,1774,1798]" metricMagnitude="-2" metricUnit="m" metricValue="2.85" pageId="3" pageNumber="2" unit="mm" value="28.5">28.5 mm</quantity>
|
||
long, the adult female is
|
||
<quantity id="1A37D7E6822FFFA052A4F89DFC3AF8AB" box="[943,1022,1805,1829]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" pageId="3" pageNumber="2" unit="mm" value="34.0">34 mm</quantity>
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||
long. For habitus see
|
||
<figureCitation id="45F46686822FFFA0540AF89DFAA8F8AA" box="[1281,1388,1805,1829]" captionStart="Figures 1–7" captionStartId="3.[192,278,1312,1336]" captionTargetBox="[203,1436,170,1268]" captionTargetId="figure-297@3.[192,1440,168,1283]" captionTargetPageId="3" captionText="Figures 1–7: Thaicharmus indicus sp. n., Figures 1–2, 7: ♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7). Figures 3–6. ♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6)." pageId="3" pageNumber="2">Figs. 1–4</figureCitation>
|
||
. For position and distribution of trichobothria of pedipalps see
|
||
<figureCitation id="45F46686822FFFA0526CF8DAFC1FF8ED" box="[871,987,1866,1890]" captionStart="Figures 8–12" captionStartId="4.[192,278,1344,1368]" captionTargetBox="[202,1437,194,1312]" captionTargetId="figure-323@4.[192,1440,192,1315]" captionTargetPageId="4" captionText="Figures 8–12: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 9–12. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." pageId="3" pageNumber="2">Figs. 9–12</figureCitation>
|
||
. Sexual dimorphism is manifested mainly in narrower male segments of metasoma and pedipalps. Other sexual differences are noted below.
|
||
</paragraph>
|
||
<caption id="89B02A8B8228FFA751CBFAD0FB99FAFF" pageId="4" pageNumber="3" startId="4.[192,278,1344,1368]" targetBox="[202,1437,194,1312]" targetPageId="4">
|
||
<paragraph id="DD707A038228FFA751CBFAD0FB99FAFF" blockId="4.[192,1441,1344,1392]" pageId="4" pageNumber="3">
|
||
<emphasis id="EFBBA6118228FFA751CBFAD0FD40FAD8" bold="true" box="[192,644,1344,1368]" pageId="4" pageNumber="3">
|
||
Figures 8–12:
|
||
<taxonomicName id="1ACF01808228FFA75062FAD2FD84FAD8" authority="Kovařík, 2013" authorityName="Kovařík" authorityYear="2013" box="[361,576,1346,1367]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="EFBBA6118228FFA75062FAD2FD84FAD8" bold="true" box="[361,576,1346,1367]" italics="true" pageId="4" pageNumber="3">Thaicharmus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4881B6A8228FFA75341FAD2FD40FAD8" box="[586,644,1346,1367]" pageId="4" pageNumber="3" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ (28.5 mm) paratype.
|
||
<emphasis id="EFBBA6118228FFA7528DFAD2FC24FAD8" bold="true" box="[902,992,1346,1367]" pageId="4" pageNumber="3">Figure 8</emphasis>
|
||
. Movable finger of pedipalp.
|
||
<emphasis id="EFBBA6118228FFA75418FAD2FA65FAD8" bold="true" box="[1299,1441,1346,1367]" pageId="4" pageNumber="3">Figures 9–12.</emphasis>
|
||
Trichobothrial pattern.
|
||
<emphasis id="EFBBA6118228FFA75094FACAFE6FFAE0" bold="true" box="[415,427,1370,1391]" pageId="4" pageNumber="3">9</emphasis>
|
||
. Chela dorsal.
|
||
<emphasis id="EFBBA6118228FFA75337FACAFD90FAE0" bold="true" box="[572,596,1370,1391]" pageId="4" pageNumber="3">10</emphasis>
|
||
. Patella dorsal.
|
||
<emphasis id="EFBBA6118228FFA753E5FACAFCC2FAE0" bold="true" box="[750,774,1370,1391]" pageId="4" pageNumber="3">11</emphasis>
|
||
. Patella external.
|
||
<emphasis id="EFBBA6118228FFA752B8FACAFC0FFAE0" bold="true" box="[947,971,1370,1391]" pageId="4" pageNumber="3">12</emphasis>
|
||
. Femur dorsal.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD707A038228FFA751CBFA71FD43F93E" blockId="4.[192,808,1444,1897]" pageId="4" pageNumber="3">COLORATION. The base color is black. The mesosoma is black with brown ornament. The femur of pedipalp is black, the patella of pedipalp is variably brownish black, and the chela is reddish brown. The telson is reddish black. The chelicerae are yellow, with black reticulation which is more defined in the anterior part. The legs are black, only the tibiae are yellowish brown.</paragraph>
|
||
<paragraph id="DD707A038228FFA751CBF928FC67F997" blockId="4.[192,808,1444,1897]" lastBlockId="4.[831,1440,1444,1897]" pageId="4" pageNumber="3">
|
||
CARAPACE. The carapace of female lacks carinae and is smooth, but in the anterior/lateral part it has two larger, symmetrically situated elevated areas by a median groove densely granulated (
|
||
<figureCitation id="45F466868228FFA750E1F884FDFDF8A3" box="[490,569,1812,1836]" captionStart="Figures 13–14" captionStartId="5.[192,278,1278,1302]" captionTargetBox="[196,1378,180,1216]" captionTargetId="figure-361@5.[192,1440,168,1248]" captionTargetPageId="5" captionText="Figures 13–14: Thaicharmus indicus sp. n., Figure 13: ♀ (34 mm) holotype, carapace with mesosomal segments I–II, chelicerae, trochanter and part of femur of pedipalp. Figure 14: ♂ (28.5 mm) paratype, telson with fifth and part of fourth metasomal segments." pageId="4" pageNumber="3">Fig. 13</figureCitation>
|
||
). Sparsely granulated is also an area around the median eyes. The male has almost entire carapace densely granulated and its posterior part is bumpy. Present are three well developed and two reduced lateral eyes. The anterior edge exhibits a broad, subtle indentation with a conspicuous epistome medially.
|
||
</paragraph>
|
||
<paragraph id="DD707A038228FFA75234F98FFABCF8A4" blockId="4.[831,1440,1444,1897]" pageId="4" pageNumber="3">
|
||
MESOSOMA. The mesosoma bears one carina and is almost smooth in the female, and is densely granulated in the male. The pectinal tooth count is
|
||
<quantity id="1A37D7E68228FFA755E1F9CCFAE5F9FB" box="[1258,1313,1628,1652]" metricMagnitude="-1" metricUnit="m" metricValue="3.81" pageId="4" pageNumber="3" unit="in" value="15.0">15 in</quantity>
|
||
the female, in the male the pectines are damaged. The marginal tips of pectines extend to three quarters of sternite III in the female. The pectines bear three marginal lamellae and seven middle lamellae. Each lamella bears several white setae, three to five on each fulcrum. Sternites III–VII are smooth, without carinae. Stigmata are short and oval.
|
||
</paragraph>
|
||
<paragraph id="DD707A038228FFA65234F8A3FD32F93A" blockId="4.[831,1440,1444,1897]" lastBlockId="5.[192,809,1387,1901]" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3">METASOMA AND TELSON. The segments of metasoma have only two dorsal carinae, which are well developed and granulated. The first and second segments are almost smooth and punctate; the third segment is also granulated, more so in the male; and the fourth and fifth segments and the telson are densely granulated. Also the dorsal surfaces of all metasomal segments are granulated in both sexes – less so on the first segment. The entire metasoma and the telson are densely hirsute, less so on the first metasomal segment. The vesicle of telson is somewhat bulbous, with a short and highly curved aculeus. A short, non-granulated but distinct subaculear tooth is present on the posterior aspect of the vesicle.</paragraph>
|
||
<caption id="89B02A8B8229FFA651CBFB6EFE4AFAC9" pageId="5" pageNumber="4" startId="5.[192,278,1278,1302]" targetBox="[196,1378,180,1216]" targetPageId="5">
|
||
<paragraph id="DD707A038229FFA651CBFB6EFE4AFAC9" blockId="5.[192,1440,1278,1350]" pageId="5" pageNumber="4">
|
||
<emphasis id="EFBBA6118229FFA651CBFB6EFD52FA9B" bold="true" box="[192,662,1278,1302]" pageId="5" pageNumber="4">
|
||
Figures 13–14:
|
||
<taxonomicName id="1ACF01808229FFA65073FB6FFD94FA9B" authority="Kovařík, 2013" authorityName="Kovařík" authorityYear="2013" box="[376,592,1279,1300]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="EFBBA6118229FFA65073FB6FFD94FA9B" bold="true" box="[376,592,1279,1300]" italics="true" pageId="5" pageNumber="4">Thaicharmus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4881B6A8229FFA65357FB6FFD52FA9B" box="[604,662,1279,1300]" pageId="5" pageNumber="4" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<emphasis id="EFBBA6118229FFA653ACFB6FFCD2FA9B" bold="true" box="[679,790,1279,1300]" pageId="5" pageNumber="4">Figure 13:</emphasis>
|
||
♀ (34 mm) holotype, carapace with mesosomal segments I–II, chelicerae, trochanter and part of femur of pedipalp.
|
||
<emphasis id="EFBBA6118229FFA653D2FA88FC83FAA2" bold="true" box="[729,839,1304,1325]" pageId="5" pageNumber="4">Figure 14:</emphasis>
|
||
♂ (28.5 mm) paratype, telson with fifth and part of fourth metasomal segments.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD707A038229FFA651CBF92DFEEBF8BF" blockId="5.[192,809,1387,1901]" pageId="5" pageNumber="4">LEGS. Both pedal spurs are present, and a long tibial spur is present on the third and fourth legs. The tarsus has scattered setae on the ventral surface. The coxae are not elongated.</paragraph>
|
||
<paragraph id="DD707A038229FFA651CBF8A7FB56F9F7" blockId="5.[192,809,1387,1901]" lastBlockId="5.[831,1447,1387,1656]" pageId="5" pageNumber="4">PEDIPALPS. The dorsointernal carina of femur is granulated, the ventrointernal carinae are serrate, and the surface is densely granulated. The femur has internal carinae only indicated, and the entire surface is granulated and punctate. The chela lacks carinae, is punctate, and its internal surface is sparsely granulated. All pedipalps are densely hirsute, but the hairs are shorter than on the metasoma. The movable and fixed fingers of pedipalps bear 10 rows of granules, with internal accessory granules and six distal granules of which three are smaller and form an apical row.</paragraph>
|
||
<paragraph id="DD707A038229FFAA5234F90EFEF6FB19" blockId="5.[831,1441,1693,1902]" lastBlockId="9.[192,802,1058,1481]" lastPageId="9" lastPageNumber="8" pageId="5" pageNumber="4">
|
||
MEASUREMENTS IN MM.
|
||
<emphasis id="EFBBA6118229FFA65555F90DFAC6F93A" box="[1118,1282,1693,1717]" italics="true" pageId="5" pageNumber="4">
|
||
Male
|
||
<typeStatus id="0274C4A18229FFA655A8F90EFAC6F93A" box="[1187,1282,1694,1717]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
|
||
</emphasis>
|
||
. Total length 28.5; carapace length 3.5, width 3.4; metasoma and telson length 17.7; first metasomal segment length 2.2, width 1.9; second metasomal segment length 2.6, width 1.8; third metasomal segment length 2.7, width 1.8; fourth metasomal segment length 2.9, width 1.8; fifth metasomal segment length 3.7, width 1.9; telson length 3.6; telson width 1.7; pedipalp femur length 2.6, width 0.9; pedipalp patella length 3.1, width 1.1; chela length 4.6; manus length 2.1; manus width 1; movable finger length 2.8.
|
||
</paragraph>
|
||
<caption id="89B02A8B822AFFA551CBF972FE5AF8F9" pageId="6" pageNumber="5" startId="6.[192,278,1762,1786]" targetBox="[262,1368,192,1729]" targetPageId="6">
|
||
<paragraph id="DD707A03822AFFA551CBF972FE5AF8F9" blockId="6.[192,1441,1762,1910]" pageId="6" pageNumber="5">
|
||
<emphasis id="EFBBA611822AFFA551CBF972FDCBF976" bold="true" box="[192,527,1762,1786]" pageId="6" pageNumber="5">Figures 15–24: Figures 15–17:</emphasis>
|
||
<taxonomicName id="1ACF0180822AFFA55313F974FC6BF975" authority="Kovarik et al., 2007" authorityName="Kovarik" authorityYear="2007" box="[536,943,1764,1786]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="lowei">
|
||
<emphasis id="EFBBA611822AFFA55313F974FD1EF976" bold="true" box="[536,730,1764,1785]" italics="true" pageId="6" pageNumber="5">Thaicharmus lowei</emphasis>
|
||
<bibRefCitation id="B95E07F2822AFFA553EFF974FC6BF975" author="KOVARIK" box="[740,943,1764,1786]" pageId="6" pageNumber="5" pagination="201 - 209" refId="ref2905" refString="KOVARIK, F., M. E. SOLEGLAD & V. FET. 2007. A new species of scorpion in the " Charmus " group from India (Scorpiones: Buthidae). Boletin de la Sociedad Entomologica Aragonesa, 40: 201 - 209." type="journal article" year="2007">Kovařík et al., 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
, dorsal (15) and ventral (16) views and live (17), ♀(16 mm) holotype, India, Goa State, Sanguem, FKCP.
|
||
<emphasis id="EFBBA611822AFFA553EBF96DFCBCF89D" bold="true" box="[736,888,1789,1810]" pageId="6" pageNumber="5">Figures 18–24:</emphasis>
|
||
<taxonomicName id="1ACF0180822AFFA55275F96DFAC6F89D" authority="Kovarik, 1995" authorityName="Kovarik" authorityYear="1995" box="[894,1282,1788,1810]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="mahunkai">
|
||
<emphasis id="EFBBA611822AFFA55275F96DFBABF89D" bold="true" box="[894,1135,1789,1810]" italics="true" pageId="6" pageNumber="5">Thaicharmus mahunkai</emphasis>
|
||
<bibRefCitation id="B95E07F2822AFFA5557FF96CFAC6F89D" author="KOVARIK" box="[1140,1282,1788,1810]" pageId="6" pageNumber="5" pagination="187 - 207" refId="ref2788" refString="KOVARIK, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae, 59: 187 - 207." type="journal article" year="1995">Kovařík, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="EFBBA611822AFFA55400F96DFF38F8A5" bold="true" pageId="6" pageNumber="5">Figures 18, 20, 23–24</emphasis>
|
||
. ♀(29 mm) paratype, Thailand, Phetchaburi Province, Kaeng Krachan, Kaeng Krachan National Park, Reservoir, FKCP, metasoma and telson lateral (18), pectinal area (20), dorsal (23) and ventral (24) views.
|
||
<emphasis id="EFBBA611822AFFA5551AF8BEFB02F8CC" bold="true" box="[1041,1222,1838,1859]" pageId="6" pageNumber="5">Figures 19, 21–22</emphasis>
|
||
. ♂(24 mm), Thailand, Kanchanaburi Province, Sai Yok, ZSRO No. 1173 (Universität Rostock, Germany), metasoma and telson lateral (19), dorsal (21) and ventral (22) views.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89B02A8B822BFFA451CBF8CEFC93F8FA" box="[192,855,1886,1910]" pageId="7" pageNumber="6" startId="7.[192,278,1886,1910]" targetBox="[211,1417,191,1855]" targetPageId="7">
|
||
<paragraph id="DD707A03822BFFA451CBF8CEFC93F8FA" blockId="7.[192,855,1886,1910]" box="[192,855,1886,1910]" pageId="7" pageNumber="6">
|
||
<emphasis id="EFBBA611822BFFA451CBF8CEFDBBF8FB" bold="true" box="[192,639,1886,1910]" pageId="7" pageNumber="6">
|
||
Figures 25–26:
|
||
<taxonomicName id="1ACF0180822BFFA45064F8CFFD87F8FB" authority="Kovařík, 2013" authorityName="Kovařík" authorityYear="2013" box="[367,579,1887,1908]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="EFBBA611822BFFA45064F8CFFD87F8FB" bold="true" box="[367,579,1887,1908]" italics="true" pageId="7" pageNumber="6">Thaicharmus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4881B6A822BFFA45342F8CFFDBBF8FB" box="[585,639,1887,1908]" pageId="7" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♀ (34 mm) holotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89B02A8B8224FFAB51CBF8CBFCA2F8FC" box="[192,870,1883,1907]" pageId="8" pageNumber="7" startId="8.[192,278,1883,1907]" targetBox="[214,1405,191,1854]" targetPageId="8">
|
||
<paragraph id="DD707A038224FFAB51CBF8CBFCA2F8FC" blockId="8.[192,870,1883,1907]" box="[192,870,1883,1907]" pageId="8" pageNumber="7">
|
||
<emphasis id="EFBBA6118224FFAB51CBF8CBFDBAF8FD" bold="true" box="[192,638,1883,1907]" pageId="8" pageNumber="7">
|
||
Figures 27–29:
|
||
<taxonomicName id="1ACF01808224FFAB5065F8CDFD86F8FD" authority="Kovařík, 2013" authorityName="Kovařík" authorityYear="2013" box="[366,578,1885,1906]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
|
||
<emphasis id="EFBBA6118224FFAB5065F8CDFD86F8FD" bold="true" box="[366,578,1885,1906]" italics="true" pageId="8" pageNumber="7">Thaicharmus indicus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="F4881B6A8224FFAB5343F8CDFDBAF8FD" box="[584,638,1885,1906]" pageId="8" pageNumber="7" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, ♂ (28.5 mm) paratype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="89B02A8B8225FFAA51CBFC47FC36FC61" box="[192,1010,983,1007]" pageId="9" pageNumber="8" startId="9.[192,267,983,1007]" targetBox="[191,1440,191,953]" targetPageId="9">
|
||
<paragraph id="DD707A038225FFAA51CBFC47FC36FC61" blockId="9.[192,1010,983,1007]" box="[192,1010,983,1007]" pageId="9" pageNumber="8">
|
||
<emphasis id="EFBBA6118225FFAA51CBFC47FEF2FC60" bold="true" box="[192,310,983,1007]" pageId="9" pageNumber="8">Figure 30:</emphasis>
|
||
India, Goa State, near Chaudi, type locality of
|
||
<taxonomicName id="1ACF01808225FFAA53F0FC49FC0BFC61" authorityName="Kovařík" authorityYear="2013" box="[763,975,985,1006]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="indicus">
|
||
<emphasis id="EFBBA6118225FFAA53F0FC49FC0BFC61" bold="true" box="[763,975,985,1006]" italics="true" pageId="9" pageNumber="8">Thaicharmus indicus</emphasis>
|
||
</taxonomicName>
|
||
sp.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DD707A038225FFAA51CBFB0DFCDBFA46" blockId="9.[192,802,1058,1481]" pageId="9" pageNumber="8">
|
||
<emphasis id="EFBBA6118225FFAA51CBFB0DFEB3FB3A" box="[192,375,1181,1205]" italics="true" pageId="9" pageNumber="8">
|
||
Female
|
||
<typeStatus id="0274C4A18225FFAA5010FB0DFEB3FB3A" box="[283,375,1181,1205]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
. Total length 34; carapace length 3.4, width 3.5; metasoma and telson length 18.85; first metasomal segment length 2.3, width 2.4; second metasomal segment length 2.6, width 2.3; third metasomal segment length 2.7, width 2.3; fourth metasomal segment length 3.1, width 1.8; fifth metasomal segment length 3.95, width 2.4; telson length 4.2; telson width 2.2; pedipalp femur length 2.8, width 1.1; pedipalp patella length 3.5, width 1.4; chela length 5.2; manus length 2.25; manus width 1.3; movable finger length 3.2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="95D529888225FFAA51CBFA78FE93F916" pageId="9" pageNumber="8" type="discussion">
|
||
<paragraph id="DD707A038225FFAA51CBFA78FE93F916" blockId="9.[192,802,1512,1689]" pageId="9" pageNumber="8">
|
||
COMMENTS. Regrettably, the male
|
||
<typeStatus id="0274C4A18225FFAA5334FA79FD5FF98E" box="[575,667,1513,1537]" pageId="9" pageNumber="8" type="paratype">paratype</typeStatus>
|
||
died during transport and was partially destroyed (
|
||
<figureCitation id="45F466868225FFAA5356F997FD01F990" box="[605,709,1543,1567]" captionStart="Figures 1–7" captionStartId="3.[192,278,1312,1336]" captionTargetBox="[203,1436,170,1268]" captionTargetId="figure-297@3.[192,1440,168,1283]" captionTargetPageId="3" captionText="Figures 1–7: Thaicharmus indicus sp. n., Figures 1–2, 7: ♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7). Figures 3–6. ♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6)." pageId="9" pageNumber="8">Figs. 1–2</figureCitation>
|
||
), so that only the metasoma and pedipalps could be preserved (
|
||
<figureCitation id="45F466868225FFAA51C3F9D4FEFAF9D3" box="[200,318,1604,1628]" captionStart-0="Figures 1–7" captionStart-1="Figures 8–12" captionStartId-0="3.[192,278,1312,1336]" captionStartId-1="4.[192,278,1344,1368]" captionTargetBox-0="[203,1436,170,1268]" captionTargetBox-1="[202,1437,194,1312]" captionTargetId-0="figure-297@3.[192,1440,168,1283]" captionTargetId-1="figure-323@4.[192,1440,192,1315]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figures 1–7: Thaicharmus indicus sp. n., Figures 1–2, 7: ♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7). Figures 3–6. ♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6)." captionText-1="Figures 8–12: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 9–12. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." pageId="9" pageNumber="8">Figs. 7–12</figureCitation>
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). However, the collector Vladimír Fura photographed the male while still alive at the type locality (
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<figureCitation id="45F466868225FFAA51C3F911FE8EF916" box="[200,330,1665,1689]" captionStart="Figures 27–29" captionStartId="8.[192,278,1883,1907]" captionTargetBox="[214,1405,191,1854]" captionTargetId="figure-23@8.[215,1404,192,1853]" captionTargetPageId="8" captionText="Figures 27–29: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype." pageId="9" pageNumber="8">Figs. 27–29</figureCitation>
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).
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</paragraph>
|
||
</subSubSection>
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</treatment>
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</document> |