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<mods:title id="1B8744FA85B037F00C4FA38143808F24">A review of Thaicharmus Kovařík, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones: Buthidae)</mods:title>
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<heading id="8638CD6F822EFFA1504CF901FD5FF923" box="[327,667,1681,1708]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="1" reason="4">
<taxonomicName id="1ACF0180822EFFA1504CF901FD5FF923" ID-CoL="646HJ" authority="Kovarik, 1995" authorityName="Kovarik" authorityYear="1995" box="[327,667,1681,1708]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EFBBA611822EFFA1504CF901FE2EF924" bold="true" box="[327,490,1681,1707]" italics="true" pageId="2" pageNumber="1">Thaicharmus</emphasis>
<bibRefCitation id="B95E07F2822EFFA150FAF901FD5FF923" author="KOVARIK" box="[497,667,1681,1708]" pageId="2" pageNumber="1" pagination="187 - 207" refId="ref2788" refString="KOVARIK, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae, 59: 187 - 207." type="journal article" year="1995">Kovařík, 1995</bibRefCitation>
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(
<figureCitation id="45F46686822EFFA150B3F93FFDEFF948" box="[440,555,1711,1735]" captionStart-0="Figures 17" captionStart-1="Figures 812" captionStart-2="Figures 1314" captionStart-3="Figures 1524" captionStart-4="Figures 2526" captionStart-5="Figures 2729" captionStart-6="Figure 30" captionStartId-0="3.[192,278,1312,1336]" captionStartId-1="4.[192,278,1344,1368]" captionStartId-2="5.[192,278,1278,1302]" captionStartId-3="6.[192,278,1762,1786]" captionStartId-4="7.[192,278,1886,1910]" captionStartId-5="8.[192,278,1883,1907]" captionStartId-6="9.[192,267,983,1007]" captionTargetBox-0="[203,1436,170,1268]" captionTargetBox-1="[202,1437,194,1312]" captionTargetBox-2="[196,1378,180,1216]" captionTargetBox-3="[262,1368,192,1729]" captionTargetBox-4="[211,1417,191,1855]" captionTargetBox-5="[214,1405,191,1854]" captionTargetBox-6="[191,1440,191,953]" captionTargetId-0="figure-297@3.[192,1440,168,1283]" captionTargetId-1="figure-323@4.[192,1440,192,1315]" captionTargetId-2="figure-361@5.[192,1440,168,1248]" captionTargetId-3="figure-117@6.[258,1368,192,1733]" captionTargetId-4="figure-23@7.[212,1416,192,1854]" captionTargetId-5="figure-23@8.[215,1404,192,1853]" captionTargetId-6="figure-501@9.[192,1439,192,953]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="8" captionTargetPageId-6="9" captionText-0="Figures 17: Thaicharmus indicus sp. n., Figures 12, 7: ♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7). Figures 36. ♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6)." captionText-1="Figures 812: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 912. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." captionText-2="Figures 1314: Thaicharmus indicus sp. n., Figure 13: ♀ (34 mm) holotype, carapace with mesosomal segments III, chelicerae, trochanter and part of femur of pedipalp. Figure 14: ♂ (28.5 mm) paratype, telson with fifth and part of fourth metasomal segments." captionText-3="Figures 1524: Figures 1517: Thaicharmus lowei Kovařík et al., 2007, dorsal (15) and ventral (16) views and live (17), ♀(16 mm) holotype, India, Goa State, Sanguem, FKCP. Figures 1824: Thaicharmus mahunkai Kovařík, 1995. Figures 18, 20, 2324. ♀(29 mm) paratype, Thailand, Phetchaburi Province, Kaeng Krachan, Kaeng Krachan National Park, Reservoir, FKCP, metasoma and telson lateral (18), pectinal area (20), dorsal (23) and ventral (24) views. Figures 19, 2122. ♂(24 mm), Thailand, Kanchanaburi Province, Sai Yok, ZSRO No. 1173 (Universität Rostock, Germany), metasoma and telson lateral (19), dorsal (21) and ventral (22) views." captionText-4="Figures 2526: Thaicharmus indicus sp. n., ♀ (34 mm) holotype." captionText-5="Figures 2729: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype." captionText-6="Figure 30: India, Goa State, near Chaudi, type locality of Thaicharmus indicus sp." pageId="2" pageNumber="1">Figs. 130</figureCitation>
)
</paragraph>
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<subSubSection id="95D52988822EFFA151CBF976FE30F8D5" pageId="2" pageNumber="1" type="reference_group">
<paragraph id="DD707A03822EFFA151CBF976FE30F8D5" blockId="2.[192,802,1766,1882]" pageId="2" pageNumber="1">
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<treatmentCitation id="5C6E5C12822EFFA151CBF976FD43F971" author="KOVARIK" box="[192,647,1766,1790]" page="199" pageId="2" pageNumber="1" year="1995">
<taxonomicName id="1ACF0180822EFFA151CBF976FD43F971" ID-CoL="646HJ" authority="Kovarik, 1995: 195 - 199" authorityName="Kovarik" authorityPageNumber="195 - 199" authorityYear="1995" box="[192,647,1766,1790]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EFBBA611822EFFA151CBF976FE89F971" box="[192,333,1766,1790]" italics="true" pageId="2" pageNumber="1">Thaicharmus</emphasis>
<bibRefCitation id="B95E07F2822EFFA15069F976FD43F971" author="KOVARIK" box="[354,647,1766,1790]" pageId="2" pageNumber="1" pagination="187 - 207" refId="ref2788" refString="KOVARIK, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae, 59: 187 - 207." type="journal article" year="1995">Kovařík, 1995: 195199</bibRefCitation>
</taxonomicName>
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, figs. 717;
<treatmentCitation id="5C6E5C12822EFFA151FBF894FE1FF893" author="KOVARIK" box="[240,475,1796,1820]" page="120" pageId="2" pageNumber="1" year="1998">
<bibRefCitation id="B95E07F2822EFFA151FBF894FE1FF893" author="KOVARIK" box="[240,475,1796,1820]" pageId="2" pageNumber="1" refId="ref2833" refString="KOVARIK, F. 1998. Stiri [Scorpiones]. Jihlava (Czech Republic): Publishing House &quot; Madagaskar &quot;, 176 pp. (in Czech)." type="book" year="1998">Kovařík, 1998: 120</bibRefCitation>
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; Fet &amp; Lowe, 2000: 224;
<treatmentCitation id="5C6E5C12822EFFA151FBF8B3FDFEF8B4" author="KOVARIK" box="[240,570,1827,1851]" page="201" pageId="2" pageNumber="1" year="2007">
<bibRefCitation id="B95E07F2822EFFA151FBF8B3FDFEF8B4" author="KOVARIK" box="[240,570,1827,1851]" pageId="2" pageNumber="1" pagination="201 - 209" refId="ref2905" refString="KOVARIK, F., M. E. SOLEGLAD &amp; V. FET. 2007. A new species of scorpion in the &quot; Charmus &quot; group from India (Scorpiones: Buthidae). Boletin de la Sociedad Entomologica Aragonesa, 40: 201 - 209." type="journal article" year="2007">Kovařík et al., 2007: 201209</bibRefCitation>
</treatmentCitation>
, figs. 114;
<treatmentCitation id="5C6E5C12822EFFA153CFF8B3FE8AF8D5" author="KOVARIK" page="18" pageId="2" pageNumber="1" year="2009">
<bibRefCitation id="B95E07F2822EFFA153CFF8B3FE8AF8D5" author="KOVARIK" pageId="2" pageNumber="1" refId="ref2864" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Prague: Clairon Production, 170 pp." type="book" year="2009">Kovařík, 2009: 18</bibRefCitation>
</treatmentCitation>
(fig. 5) and 31.
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</paragraph>
</subSubSection>
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<paragraph id="DD707A03822EFFA15234FCA9FABDFCDE" blockId="2.[831,1401,825,849]" box="[831,1401,825,849]" pageId="2" pageNumber="1">
<materialsCitation id="6DA7705E822EFFA15234FCA9FABDFCDE" box="[831,1401,825,849]" collectionCode="SPECIES" pageId="2" pageNumber="1" specimenCount="1" typeStatus="holotype">
<typeStatus id="0274C4A1822EFFA15234FCA9FCB3FCDF" box="[831,887,825,848]" pageId="2" pageNumber="1">TYPE</typeStatus>
<collectionCode id="BBDEE2C6822EFFA15276FCACFC0BFCDF" box="[893,975,828,848]" pageId="2" pageNumber="1">SPECIES</collectionCode>
.
<taxonomicName id="1ACF0180822EFFA152D0FCA9FABDFCDE" authority="Kovarik, 1995" authorityName="Kovarik" authorityYear="1995" box="[987,1401,825,849]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="EFBBA611822EFFA152D0FCA9FB1CFCDE" box="[987,1240,825,849]" italics="true" pageId="2" pageNumber="1">Thaicharmus mahunkai</emphasis>
<bibRefCitation id="B95E07F2822EFFA155D4FCA9FABDFCDE" author="KOVARIK" box="[1247,1401,825,849]" pageId="2" pageNumber="1" pagination="187 - 207" refId="ref2788" refString="KOVARIK, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae, 59: 187 - 207." type="journal article" year="1995">Kovařík, 1995</bibRefCitation>
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</paragraph>
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<paragraph id="DD707A03822EFFA15234FCEDFAF5FC3C" blockId="2.[831,1440,892,947]" pageId="2" pageNumber="1">
ETYMOLOGY. Denotes affinity to the genus
<taxonomicName id="1ACF0180822EFFA1541AFCECFAB7FC1B" authorityName="Karsch" authorityYear="1879" box="[1297,1395,892,916]" class="Arachnida" family="Buthidae" genus="Charmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis id="EFBBA611822EFFA1541AFCECFAB7FC1B" box="[1297,1395,892,916]" italics="true" pageId="2" pageNumber="1">Charmus</emphasis>
</taxonomicName>
and the geographic distribution of the
<typeStatus id="0274C4A1822EFFA155A3FC0BFB12FC3C" box="[1192,1238,923,947]" pageId="2" pageNumber="1">type</typeStatus>
species.
</paragraph>
</subSubSection>
<subSubSection id="95D52988822EFFA15234FC4FFAF2FC79" box="[831,1334,990,1014]" pageId="2" pageNumber="1" type="distribution">
<paragraph id="DD707A03822EFFA15234FC4FFAF2FC79" blockId="2.[831,1334,990,1014]" box="[831,1334,990,1014]" pageId="2" pageNumber="1">
DISTRIBUTION.
<collectingCountry id="A5D83A93822EFFA152EEFC4EFB80FC79" box="[997,1092,990,1014]" name="Thailand" pageId="2" pageNumber="1">Thailand</collectingCountry>
and
<collectingCountry id="A5D83A93822EFFA1557CFC4EFB6AFC79" box="[1143,1198,990,1014]" name="India" pageId="2" pageNumber="1">India</collectingCountry>
(Goa State).
</paragraph>
</subSubSection>
<subSubSection id="95D52988822EFFA15234FBB2FBF0F921" pageId="2" pageNumber="1" type="diagnosis">
<paragraph id="DD707A03822EFFA15234FBB2FC40F980" blockId="2.[831,1441,1058,1551]" pageId="2" pageNumber="1">
DIAGNOSIS. Total length
<quantity id="1A37D7E6822EFFA15561FBB2FB2FFBB6" box="[1130,1259,1058,1082]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.4" metricValueMin="1.6" pageId="2" pageNumber="1" unit="mm" value="25.0" valueMax="34.0" valueMin="16.0">1634 mm</quantity>
. Dorsal trichobothria of femur arranged in
<emphasis id="EFBBA611822EFFA15578FBD1FB59FBD6" box="[1139,1181,1089,1113]" italics="true" pageId="2" pageNumber="1">alfa</emphasis>
-configuration (
<figureCitation id="45F46686822EFFA1544FFBD1FA50FBD6" box="[1348,1428,1089,1113]" captionStart="Figures 812" captionStartId="4.[192,278,1344,1368]" captionTargetBox="[202,1437,194,1312]" captionTargetId="figure-323@4.[192,1440,192,1315]" captionTargetPageId="4" captionText="Figures 812: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 912. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." pageId="2" pageNumber="1">Fig. 12</figureCitation>
). Pedipalp patella with trichobothria
<emphasis id="EFBBA611822EFFA155EEFBF1FAC6FBF7" box="[1253,1282,1121,1144]" italics="true" pageId="2" pageNumber="1">est</emphasis>
,
<emphasis id="EFBBA611822EFFA15417FBF1FAFFFBF7" box="[1308,1339,1121,1144]" italics="true" pageId="2" pageNumber="1">em</emphasis>
, and
<emphasis id="EFBBA611822EFFA15486FBF1FA64FBF7" box="[1421,1440,1121,1144]" italics="true" pageId="2" pageNumber="1">et</emphasis>
aligned in a straight line angled distally towards the ventral edge. Pedipalp chela with three
<emphasis id="EFBBA611822EFFA155FBFB0FFACAFB38" box="[1264,1294,1183,1207]" italics="true" pageId="2" pageNumber="1">Eb</emphasis>
trichobothria on manus (
<figureCitation id="45F46686822EFFA152BEFB2EFC33FB59" box="[949,1015,1214,1238]" captionStart="Figures 812" captionStartId="4.[192,278,1344,1368]" captionTargetBox="[202,1437,194,1312]" captionTargetId="figure-323@4.[192,1440,192,1315]" captionTargetPageId="4" captionText="Figures 812: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 912. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." pageId="2" pageNumber="1">Fig. 9</figureCitation>
). Carapace anterior edge with epistome present medially. Third and fourth legs with long tibial spurs. Coxae of legs short, not elongated. Sternum subpentagonal, roughly as wide as long, exhibiting minor horizontal compression. Movable finger of pedipalp longer than manus. Telson vesicle globular, equipped with a small but distinct subaculear tubercle. Movable fingers of pedipalps with 1012 cutting rows of granules. Pedipalps, metasoma and telson densely hirsute. Pectines with enlarged basal middle lamellae and with fulcra.
</paragraph>
<paragraph id="DD707A03822EFFA15234F9A8FBF0F921" blockId="2.[831,1440,1592,1711]" pageId="2" pageNumber="1">
SUBORDINATE TAXA.
<taxonomicName id="1ACF0180822EFFA15539F9A8FCB0F9E0" authority="Kovarik, 1995" authorityName="Kovarik" authorityYear="1995" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="mahunkai">
<emphasis id="EFBBA611822EFFA15539F9A8FAF1F9DF" box="[1074,1333,1592,1616]" italics="true" pageId="2" pageNumber="1">Thaicharmus mahunkai</emphasis>
<bibRefCitation id="B95E07F2822EFFA15448F9A8FCB0F9E0" author="KOVARIK" pageId="2" pageNumber="1" pagination="187 - 207" refId="ref2788" refString="KOVARIK, F. 1995. Review of Scorpionida from Thailand with descriptions of Thaicharmus mahunkai gen. et sp. n. and Lychas krali sp. n. (Buthidae). Acta Societatis Zoologicae Bohemicae, 59: 187 - 207." type="journal article" year="1995">Kovařík, 1995</bibRefCitation>
</taxonomicName>
(
<figureCitation id="45F46686822EFFA1528DF9C7FBC9F9E0" box="[902,1037,1623,1647]" captionStart="Figures 1524" captionStartId="6.[192,278,1762,1786]" captionTargetBox="[262,1368,192,1729]" captionTargetId="figure-117@6.[258,1368,192,1733]" captionTargetPageId="6" captionText="Figures 1524: Figures 1517: Thaicharmus lowei Kovařík et al., 2007, dorsal (15) and ventral (16) views and live (17), ♀(16 mm) holotype, India, Goa State, Sanguem, FKCP. Figures 1824: Thaicharmus mahunkai Kovařík, 1995. Figures 18, 20, 2324. ♀(29 mm) paratype, Thailand, Phetchaburi Province, Kaeng Krachan, Kaeng Krachan National Park, Reservoir, FKCP, metasoma and telson lateral (18), pectinal area (20), dorsal (23) and ventral (24) views. Figures 19, 2122. ♂(24 mm), Thailand, Kanchanaburi Province, Sai Yok, ZSRO No. 1173 (Universität Rostock, Germany), metasoma and telson lateral (19), dorsal (21) and ventral (22) views." pageId="2" pageNumber="1">Figs. 1824</figureCitation>
);
<taxonomicName id="1ACF0180822EFFA1552FF9C7FA9DF9E0" authority="Kovarik" authorityName="Kovarik, Soleglad et Fet" authorityYear="2007" box="[1060,1369,1623,1647]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="lowei">
<emphasis id="EFBBA611822EFFA1552FF9C7FB31F9E0" box="[1060,1269,1623,1647]" italics="true" pageId="2" pageNumber="1">Thaicharmus lowei</emphasis>
Kovařík
</taxonomicName>
, Sole- glad et Fet, 2007 (
<figureCitation id="45F46686822EFFA15509F9E7FB40F900" box="[1026,1156,1655,1679]" captionStart="Figures 1524" captionStartId="6.[192,278,1762,1786]" captionTargetBox="[262,1368,192,1729]" captionTargetId="figure-117@6.[258,1368,192,1733]" captionTargetPageId="6" captionText="Figures 1524: Figures 1517: Thaicharmus lowei Kovařík et al., 2007, dorsal (15) and ventral (16) views and live (17), ♀(16 mm) holotype, India, Goa State, Sanguem, FKCP. Figures 1824: Thaicharmus mahunkai Kovařík, 1995. Figures 18, 20, 2324. ♀(29 mm) paratype, Thailand, Phetchaburi Province, Kaeng Krachan, Kaeng Krachan National Park, Reservoir, FKCP, metasoma and telson lateral (18), pectinal area (20), dorsal (23) and ventral (24) views. Figures 19, 2122. ♂(24 mm), Thailand, Kanchanaburi Province, Sai Yok, ZSRO No. 1173 (Universität Rostock, Germany), metasoma and telson lateral (19), dorsal (21) and ventral (22) views." pageId="2" pageNumber="1">Figs. 1517</figureCitation>
);
<taxonomicName id="1ACF0180822EFFA15592F9E7FABDF900" authority="Kovařík, 2013" authorityName="Kovařík" authorityYear="2013" box="[1177,1401,1655,1679]" class="Arachnida" family="Buthidae" genus="Thaicharmus" kingdom="Animalia" order="Scorpiones" pageId="2" pageNumber="1" phylum="Arthropoda" rank="species" species="indicus" status="sp. nov.">
<emphasis id="EFBBA611822EFFA15592F9E7FABDF900" box="[1177,1401,1655,1679]" italics="true" pageId="2" pageNumber="1">Thaicharmus indicus</emphasis>
</taxonomicName>
<emphasis id="EFBBA611822EFFA1548AF9E7FC90F920" bold="true" pageId="2" pageNumber="1">
<taxonomicNameLabel id="F4881B6A822EFFA1548AF9E7FC90F920" pageId="2" pageNumber="1" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="45F46686822EFFA15268F906FC12F921" box="[867,982,1686,1710]" captionStart-0="Figures 17" captionStart-1="Figures 812" captionStart-2="Figures 1314" captionStartId-0="3.[192,278,1312,1336]" captionStartId-1="4.[192,278,1344,1368]" captionStartId-2="5.[192,278,1278,1302]" captionTargetBox-0="[203,1436,170,1268]" captionTargetBox-1="[202,1437,194,1312]" captionTargetBox-2="[196,1378,180,1216]" captionTargetId-0="figure-297@3.[192,1440,168,1283]" captionTargetId-1="figure-323@4.[192,1440,192,1315]" captionTargetId-2="figure-361@5.[192,1440,168,1248]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="Figures 17: Thaicharmus indicus sp. n., Figures 12, 7: ♂ (28.5 mm) paratype, dorsal (1) and ventral (2) views, metasoma and telson lateral view (7). Figures 36. ♀ (34 mm) holotype, dorsal (3) and ventral (4) views, pectinal area (5) and metasoma and telson lateral view (6)." captionText-1="Figures 812: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype. Figure 8. Movable finger of pedipalp. Figures 912. Trichobothrial pattern. 9. Chela dorsal. 10. Patella dorsal. 11. Patella external. 12. Femur dorsal." captionText-2="Figures 1314: Thaicharmus indicus sp. n., Figure 13: ♀ (34 mm) holotype, carapace with mesosomal segments III, chelicerae, trochanter and part of femur of pedipalp. Figure 14: ♂ (28.5 mm) paratype, telson with fifth and part of fourth metasomal segments." pageId="2" pageNumber="1">Figs. 114</figureCitation>
,
<figureCitation id="45F46686822EFFA152E9F906FBE2F921" box="[994,1062,1686,1710]" captionStart-0="Figures 2526" captionStart-1="Figures 2729" captionStartId-0="7.[192,278,1886,1910]" captionStartId-1="8.[192,278,1883,1907]" captionTargetBox-0="[211,1417,191,1855]" captionTargetBox-1="[214,1405,191,1854]" captionTargetId-0="figure-23@7.[212,1416,192,1854]" captionTargetId-1="figure-23@8.[215,1404,192,1853]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 2526: Thaicharmus indicus sp. n., ♀ (34 mm) holotype." captionText-1="Figures 2729: Thaicharmus indicus sp. n., ♂ (28.5 mm) paratype." pageId="2" pageNumber="1">2529</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>