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<document id="1781FF9E3DBE89A5FB3DCB6B64BCBC5A" ID-DOI="10.5281/zenodo.203208" ID-GBIF-Dataset="85473c5c-8506-4915-9dd9-0317382ba608" ID-ISSN="1175-5326" ID-Zenodo-Dep="203208" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460297531016" checkinUser="plazi" docAuthor="Kits, Joel H. &amp; Marshall, Stephen A." docDate="2011" docId="5562879BFFD7FFD7EAA4F946B416FCEC" docLanguage="en" docName="zt02863p034.pdf" docOrigin="Zootaxa 2863" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Frutillaria abdita Kits &amp; Marshall, 2011, sp. n." docType="treatment" docVersion="8" lastPageNumber="8" masterDocId="A95BFFE3FFD2FFD0EA33FF81B066FFB0" masterDocTitle="A revision of Frutillaria Richards and Penola Richards (Diptera: Sphaeroceridae: Archiborborinae)" masterLastPageNumber="34" masterPageNumber="1" pageNumber="6" updateTime="1698252249425" updateUser="plazi">
<mods:mods id="F7460F5388751829D97101C6999054FD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E7DD01AE53C8C0F8E994230A6C55E0FF">
<mods:title id="D9EC7ABDF3DE9764799FDB7C3C2046DC">A revision of Frutillaria Richards and Penola Richards (Diptera: Sphaeroceridae: Archiborborinae)</mods:title>
</mods:titleInfo>
<mods:name id="7961F7507C2295019128ACB380FF0E62" type="personal">
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<mods:namePart id="A23BB95CBB0CEBE581FD9F4BA7E7800B">Kits, Joel H.</mods:namePart>
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<mods:name id="DC9DE0C3B72A874FD4B7C43E1B9D149B" type="personal">
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<mods:namePart id="DD3031774C5EC37C66F697A582E8706B">Marshall, Stephen A.</mods:namePart>
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<mods:date id="A1815E160DB22F6613EA2DDF49BEDA84">2011</mods:date>
<mods:detail id="6563D6D520B9C172DFD9CF95699BB0B6" type="volume">
<mods:number id="9B471A1E2942306F6B47C75591461E25">2863</mods:number>
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<mods:start id="AC9AE16CED1EFD81595CDDB9BCE467BF">1</mods:start>
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<mods:classification id="DB6BB5DCEC75C05F81A2D07208A0635D">journal article</mods:classification>
<mods:identifier id="CC0ADBAE70D569F77BE0877D6DBB9B57" type="DOI">10.5281/zenodo.203208</mods:identifier>
<mods:identifier id="4F396C7F8C7E7C822D7D442B06E87275" type="GBIF-Dataset">85473c5c-8506-4915-9dd9-0317382ba608</mods:identifier>
<mods:identifier id="CDAD98D96DFF298686B134D24D0716D9" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="03E8C51CADE28ED1E594AB8CB543D605" type="Zenodo-Dep">203208</mods:identifier>
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<treatment id="5562879BFFD7FFD7EAA4F946B416FCEC" ID-DOI="http://doi.org/10.5281/zenodo.6191597" ID-GBIF-Taxon="119399305" ID-Zenodo-Dep="6191597" LSID="urn:lsid:plazi:treatment:5562879BFFD7FFD7EAA4F946B416FCEC" httpUri="http://treatment.plazi.org/id/5562879BFFD7FFD7EAA4F946B416FCEC" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
<subSubSection id="95D16506FFD7FFD5EAA4F946B1A1F8B2" pageId="5" pageNumber="6" type="nomenclature">
<paragraph id="DD74368DFFD7FFD5EAA4F946B1D5F951" blockId="5.[151,455,1735,1794]" box="[151,435,1735,1761]" pageId="5" pageNumber="6">
<heading id="863C81E1FFD7FFD5EAA4F946B1D5F951" bold="true" box="[151,435,1735,1761]" fontSize="11" level="1" pageId="5" pageNumber="6" reason="1">
<emphasis id="EFBFEA9FFFD7FFD5EAA4F946B1D5F951" bold="true" box="[151,435,1735,1761]" pageId="5" pageNumber="6">
<taxonomicName id="1ACB4D0EFFD7FFD5EAA4F946B10CF951" ID-CoL="8C7DT" box="[151,362,1735,1761]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="abdita" status="sp. nov.">
<emphasis id="EFBFEA9FFFD7FFD5EAA4F946B10CF951" bold="true" box="[151,362,1735,1761]" italics="true" pageId="5" pageNumber="6">Frutillaria abdita</emphasis>
</taxonomicName>
<taxonomicNameLabel id="F48C57E4FFD7FFD5EB42F946B1D5F951" box="[369,435,1735,1761]" pageId="5" pageNumber="6" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="DD74368DFFD7FFD5EAA4F96BB1A1F8B2" blockId="5.[151,455,1735,1794]" box="[151,455,1770,1794]" pageId="5" pageNumber="6">
(
<figureCitation id="45F02A08FFD7FFD5EAACF96BB16DF8B2" box="[159,267,1770,1794]" captionStart="FIGURES 1 – 9" captionStartId="6.[151,264,1676,1699]" captionTargetBox="[162,1428,187,1655]" captionTargetId="figure@6.[151,1436,686,1659]" captionTargetPageId="6" captionText="FIGURES 1 – 9. Frutillaria adults, habitus: (1) F. tenuiforceps, Zapallar; (2) F. abdita paratype, Putú, Chile. Frutillaria male genitalia: F. abdita: (3) cerci and surstylus, posterior view, (4) surstylus, anterior view, (5) phallus, lateral view; F. anticura: (6) postgonite, lateral view, (7) cerci and surstylus, posterior view, (8) surstylus, anterior" httpUri="https://zenodo.org/record/203209/files/figure.png" pageId="5" pageNumber="6">Figs. 2–6</figureCitation>
,
<figureCitation id="45F02A08FFD7FFD5EB24F96BB1EEF8B2" box="[279,392,1770,1794]" captionStart="FIGURES 59 – 67" captionStartId="26.[151,264,1936,1959]" captionTargetBox="[163,1418,783,1907]" captionTargetId="figure@26.[151,1436,769,1915]" captionTargetPageId="26" captionText="FIGURES 59 – 67. Frutillaria female terminalia. F. abdita: (59) tergite 8, epiproct, cerci, (60) sternite 8, hypoproct; F. anticura: (61) tergite 8, epiproct, cerci, (62) sternite 8, hypoproct; F. calceata: (63) tergites 7 – 8, epiproct, cerci, (64) sternites 7 – 8, hypoproct. Spermathecae: (65) F. abdita, (66) F. anticura, (67) F. calceata." httpUri="https://zenodo.org/record/203218/files/figure.png" pageId="5" pageNumber="6">59, 60, 65</figureCitation>
,
<figureCitation id="45F02A08FFD7FFD5EBA7F96BB1D9F8B2" box="[404,447,1770,1794]" captionStart="FIGURES 108 – 115" captionStartId="32.[151,264,1747,1770]" captionTargetBox="[151,1436,591,1724]" captionTargetId="figure@32.[151,1436,591,1725]" captionTargetPageId="32" captionText="FIGURES 108 – 115. Frutillaria and Penola distribution maps. (108) F. abdita, F. f u rc a t a, F. tenuiforceps, F. triangularis; (109) F. anticura, F. chepuensis, F. glabra, P. eudyptidis; (110) F. c o n t u l m o, F. calceata, F. chiloensis, F. propinqua; (111) F. calida; (112) F. edenensis; (113) F. k u s c h e l i, F. r i c ha rds i; (114) F. stenoptera; (115) F. transversa." httpUri="https://zenodo.org/record/203224/files/figure.png" pageId="5" pageNumber="6">108</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="95D16506FFD7FFD7EAA4F8B3B332FD34" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6" type="description">
<paragraph id="DD74368DFFD7FFD5EAA4F8B3B328F8FA" blockId="5.[151,1436,1842,1939]" box="[151,846,1842,1867]" pageId="5" pageNumber="6">
<emphasis id="EFBFEA9FFFD7FFD5EAA4F8B3B14FF8FB" bold="true" box="[151,297,1842,1867]" pageId="5" pageNumber="6">Description.</emphasis>
Length 1.5–1.9 mm (head + thorax 0.8–1.0 mm)
</paragraph>
<paragraph id="DD74368DFFD7FFD5EAF4F8D7B345F822" blockId="5.[151,1436,1842,1939]" pageId="5" pageNumber="6">Occiput brown, face and gena yellow, frons intermediately coloured. Frons with shiny spots lateral to ocellar triangle. Antenna yellow. Clypeus and prementum yellow.</paragraph>
<caption id="89B46605FFD4FFD6EAA4F90DB394F955" httpUri="https://zenodo.org/record/203209/files/figure.png" pageId="6" pageNumber="7" targetBox="[162,1428,187,1655]" targetPageId="6">
<paragraph id="DD74368DFFD4FFD6EAA4F90DB394F955" blockId="6.[151,1430,1676,1794]" pageId="6" pageNumber="7">
<emphasis id="EFBFEA9FFFD4FFD6EAA4F90DB15CF912" bold="true" box="[151,314,1676,1699]" pageId="6" pageNumber="7">FIGURES 1–9.</emphasis>
<taxonomicName id="1ACB4D0EFFD4FFD6EB73F90CB1CAF913" box="[320,428,1677,1699]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="EFBFEA9FFFD4FFD6EB73F90CB1CAF913" box="[320,428,1677,1699]" italics="true" pageId="6" pageNumber="7">Frutillaria</emphasis>
</taxonomicName>
adults, habitus: (1)
<taxonomicName id="1ACB4D0EFFD4FFD6E845F90FB316F912" authority="Zapallar" authorityName="Zapallar" box="[630,880,1676,1699]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="tenuiforceps">
<emphasis id="EFBFEA9FFFD4FFD6E845F90FB36DF913" box="[630,779,1677,1699]" italics="true" pageId="6" pageNumber="7">F. tenuiforceps</emphasis>
, Zapallar
</taxonomicName>
; (2)
<taxonomicName id="1ACB4D0EFFD4FFD6E9AEF90FB391F913" box="[925,1015,1677,1699]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="abdita">
<emphasis id="EFBFEA9FFFD4FFD6E9AEF90FB391F913" box="[925,1015,1677,1699]" italics="true" pageId="6" pageNumber="7">F. abdita</emphasis>
</taxonomicName>
paratype, Putú, Chile.
<taxonomicName id="1ACB4D0EFFD4FFD6EED3F90CB52AF913" box="[1248,1356,1677,1699]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="genus">
<emphasis id="EFBFEA9FFFD4FFD6EED3F90CB52AF913" box="[1248,1356,1677,1699]" italics="true" pageId="6" pageNumber="7">Frutillaria</emphasis>
</taxonomicName>
male genitalia:
<taxonomicName id="1ACB4D0EFFD4FFD6EAC8F92EB132F974" box="[251,340,1710,1732]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="abdita">
<emphasis id="EFBFEA9FFFD4FFD6EAC8F92EB132F974" box="[251,340,1710,1732]" italics="true" pageId="6" pageNumber="7">F. abdita</emphasis>
</taxonomicName>
: (3) cerci and surstylus, posterior view, (4) surstylus, anterior view, (5) phallus, lateral view;
<taxonomicName id="1ACB4D0EFFD4FFD6EECEF92EB50AF974" box="[1277,1388,1711,1732]" class="Insecta" family="Sphaeroceridae" genus="Frutillaria" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="anticura">
<emphasis id="EFBFEA9FFFD4FFD6EECEF92EB50AF974" box="[1277,1388,1711,1732]" italics="true" pageId="6" pageNumber="7">F. anticura</emphasis>
</taxonomicName>
: (6) postgonite, lateral view, (7) cerci and surstylus, posterior view, (8) surstylus, anterior
</paragraph>
</caption>
<paragraph id="DD74368DFFD4FFD6EAA4F96AB202F8B1" blockId="6.[151,1430,1676,1794]" box="[151,612,1771,1794]" pageId="6" pageNumber="7">view, (9) phallus and postgonite, lateral view.</paragraph>
<paragraph id="DD74368DFFD4FFD6EAF4F8AFB432F866" blockId="6.[151,1436,1838,2006]" pageId="6" pageNumber="7">Thorax black, postalar callus and scutellum reddish. Wing rudiment brown. Mesoscutum and scutellum extensively shiny, some microtomentum near margins. Anepisternum entirely shiny. Katepisternum with a large shiny patch behind fore coxa extending posteriorly as a triangle to near bristle, narrowly separated from posterior shiny patch by a band of microtomentum. Meron and metapleuron shiny with the exception of a strip of microtomentum between posterior thoracic spiracle and base of hind coxa. Laterotergite mostly shiny.</paragraph>
<paragraph id="DD74368DFFD5FFD7EAF4FF19B343FF64" blockId="7.[151,1436,151,860]" pageId="7" pageNumber="8">Legs dark brown, fore coxa, fore femur, trochanters, diffuse area near joints, and tarsi yellow. Mid femur with 2 anterior bristles, mid tibia without posteroventral bristle.</paragraph>
<paragraph id="DD74368DFFD5FFD7EAF4FF5EB3EEFE1C" blockId="7.[151,1436,151,860]" pageId="7" pageNumber="8">
<collectingCountry id="A5DC761DFFD5FFD7EAF4FF5EB167FF47" box="[199,257,223,247]" name="Malta" pageId="7" pageNumber="8">Male</collectingCountry>
abdomen (
<figureCitation id="45F02A08FFD5FFD7EB4DFF5EB18DFF47" box="[382,491,223,247]" captionStart="FIGURES 1 – 9" captionStartId="6.[151,264,1676,1699]" captionTargetBox="[162,1428,187,1655]" captionTargetId="figure@6.[151,1436,686,1659]" captionTargetPageId="6" captionText="FIGURES 1 – 9. Frutillaria adults, habitus: (1) F. tenuiforceps, Zapallar; (2) F. abdita paratype, Putú, Chile. Frutillaria male genitalia: F. abdita: (3) cerci and surstylus, posterior view, (4) surstylus, anterior view, (5) phallus, lateral view; F. anticura: (6) postgonite, lateral view, (7) cerci and surstylus, posterior view, (8) surstylus, anterior" httpUri="https://zenodo.org/record/203209/files/figure.png" pageId="7" pageNumber="8">Figs. 3–6</figureCitation>
): Syntergite 1+2 with some microtomentum in a patch behind scutellum, tergites 3– 5 entirely shiny. Sternites 2–5 entirely shiny. Surstylus with broad basal section and fairly long inner arm, outer corner rounded, not produced. Postgonite bilobed, anterior lobe about ½ length of posterior. Epiphallus barely produced, broadened laterally; preepipallus well-developed, long and thin. Distiphallus with lateral arm produced from ventral sclerite, pointed apically with 2 medial teeth; a large complex, translucent sclerite apically of uncertain homology; ventral sacs clear, not projecting and barely visible.
</paragraph>
<paragraph id="DD74368DFFD5FFD7EAF4FE36B332FD34" blockId="7.[151,1436,151,860]" pageId="7" pageNumber="8">
Female abdomen (
<figureCitation id="45F02A08FFD5FFD7EBA4FE36B22BFE7F" box="[407,589,439,463]" captionStart="FIGURES 59 – 67" captionStartId="26.[151,264,1936,1959]" captionTargetBox="[163,1418,783,1907]" captionTargetId="figure@26.[151,1436,769,1915]" captionTargetPageId="26" captionText="FIGURES 59 – 67. Frutillaria female terminalia. F. abdita: (59) tergite 8, epiproct, cerci, (60) sternite 8, hypoproct; F. anticura: (61) tergite 8, epiproct, cerci, (62) sternite 8, hypoproct; F. calceata: (63) tergites 7 – 8, epiproct, cerci, (64) sternites 7 – 8, hypoproct. Spermathecae: (65) F. abdita, (66) F. anticura, (67) F. calceata." httpUri="https://zenodo.org/record/203218/files/figure.png" pageId="7" pageNumber="8">Figs. 59, 60, 65</figureCitation>
): Syntergite 1+2 with a patch of microtomentum behind scutellum, tergites 3–5 with a band of microtomentum near base. Tergite 6 covered with microtomentum, tergites 7–8 mostly shiny with very little microtomentum apically. Sternites 2–5 shiny, sternite 6 covered with microtomentum, tergite 7 mostly shiny with some microtomentum apically. Tergite 8 complete, epiproct broadly triangular with distinct anterior arms. Sclerites of sternite 8 teardrop-shaped, hypoproct not sclerotized. Spermathecae with basal invagination short, annulated; apical invagination short, subconical, smooth
</paragraph>
</subSubSection>
<subSubSection id="95D16506FFD5FFD7EAF4FD11B3A3FD7C" pageId="7" pageNumber="8" type="materials_examined">
<paragraph id="DD74368DFFD5FFD7EAF4FD11B3A3FD7C" blockId="7.[151,1436,151,860]" pageId="7" pageNumber="8">
<emphasis id="EFBFEA9FFFD5FFD7EAF4FD11B115FD18" bold="true" box="[199,371,655,680]" pageId="7" pageNumber="8">
<typeStatus id="0270882FFFD5FFD7EAF4FD11B166FD18" box="[199,256,656,680]" pageId="7" pageNumber="8">Type</typeStatus>
material.
</emphasis>
<typeStatus id="0270882FFFD5FFD7EB4FFD0EB182FD17" box="[380,484,655,679]" pageId="7" pageNumber="8" type="holotype">Holotype</typeStatus>
3:
<collectingCountry id="A5DC761DFFD5FFD7E822FD0EB20EFD17" box="[529,616,655,679]" name="Chile" pageId="7" pageNumber="8">CHILE</collectingCountry>
:
<emphasis id="EFBFEA9FFFD5FFD7E840FD11B2AEFD18" bold="true" box="[627,712,656,680]" pageId="7" pageNumber="8">Maule:</emphasis>
Putú, ~
<quantity id="1A339B68FFD5FFD7E912FD0EB30EFD17" box="[801,872,655,679]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="8" unit="km" value="15.0">15 km</quantity>
N Constitución, along stream,
<date id="A975104DFFD5FFD7EEFBFD0EB539FD17" box="[1224,1375,655,679]" pageId="7" pageNumber="8" value="2006-11-26">26 Nov 2006</date>
, S.A.
<collectingCountry id="A5DC761DFFD5FFD7EAA4FD35B09DFD7C" box="[151,251,692,716]" name="Marshall Islands" pageId="7" pageNumber="8">Marshall</collectingCountry>
(
<collectionCode id="BBDAAE48FFD5FFD7EB3AFD35B102FD7C" box="[265,356,692,716]" httpUri="http://grbio.org/cool/mf2w-ryyx" name="Museo Nacional de Historia Natural, Santiago" pageId="7" pageNumber="8">MNNC</collectionCode>
).
<typeStatus id="0270882FFFD5FFD7EB45FD34B181FD7D" box="[374,487,693,717]" pageId="7" pageNumber="8" type="paratype">Paratypes</typeStatus>
: same data as
<typeStatus id="0270882FFFD5FFD7E8B5FD35B28EFD7C" box="[646,744,692,716]" pageId="7" pageNumber="8" type="holotype">holotype</typeStatus>
(2 3, 8 Ƥ,
<collectionCode id="BBDAAE48FFD5FFD7E95BFD35B3DFFD7C" box="[872,953,692,716]" httpUri="http://grbio.org/cool/yhs7-n61d" name="Ontario Insect Collection, University of Guelph" pageId="7" pageNumber="8">DEBU</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="95D16506FFD5FFD7EAF4FD56B416FCEC" pageId="7" pageNumber="8" type="discussion">
<paragraph id="DD74368DFFD5FFD7EAF4FD56B416FCEC" blockId="7.[151,1436,151,860]" pageId="7" pageNumber="8">
<emphasis id="EFBFEA9FFFD5FFD7EAF4FD56B129FD40" bold="true" box="[199,335,727,752]" pageId="7" pageNumber="8">Comments.</emphasis>
This species was found in a small wooded ravine on the heavily agricultural coastal plain north of Constitución. The habitat patch was extremely small and completely isolated by a large area of pasture from any other intact forest; it was apparently preserved only by virtue of the steep sides which prevented cattle from entering to browse. The species name refers to the hidden and unexpected collection locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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