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<document id="CCB01C9BFBBC07404E18D5DCB7B510CF" ID-DOI="10.3853/j.2201-4349.68.2016.1659" ID-ISSN="2201-4349" ID-Zenodo-Dep="4676914" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617989658085" checkinUser="carolina" docAuthor="Batley, Michael &amp; Popic, Tony J." docDate="2016" docId="5552878F9B76FF9B01CDFD39FE03FE53" docLanguage="en" docName="RecAustMus.68.6.263-268.pdf" docOrigin="Records of the Australian Museum (Rec. Aust. Mus.) 68 (6)" docSource="http://dx.doi.org/10.3853/j.2201-4349.68.2016.1659" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Leioproctus glendae Batley 2016, n. sp." docType="treatment" docUuid="1D4880EC-F83C-4BE8-9A23-5EABE927978B" docUuidSource="ZooBank" docVersion="4" lastPageNumber="267" masterDocId="A96BFFF79B77FF9F012EFF9CFFA2FFFF" masterDocTitle="An Unusual New Leioproctus Species (Hymenoptera: Colletidae)" masterLastPageNumber="268" masterPageNumber="263" pageNumber="264" updateTime="1698904470259" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="7AB13694154EB460BC6683D7F6497431" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="87C46077717FFDF57B093F4A826A0D69">
<mods:title id="7955D2F7B4486CAAB0CC97844DAC50BD">An Unusual New Leioproctus Species (Hymenoptera: Colletidae)</mods:title>
</mods:titleInfo>
<mods:name id="3D09B04F7AA7CB0912B36DFF46F0B1FF" type="personal">
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<mods:namePart id="8896CA3D6BA0380391E8924A7C0104B7">Batley, Michael</mods:namePart>
</mods:name>
<mods:name id="44D134C50E94137BCF54B52C4E9CF720" type="personal">
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<mods:namePart id="CCEBD5CD42B5B3E6B45DEEA8E55E983C">Popic, Tony J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="8B0F02199CA8AA32640B0014DBD38AF0">text</mods:typeOfResource>
<mods:relatedItem id="6F1B2C7A3B99D01BFBC607E322E7D27F" type="host">
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<mods:title id="67C6513B40FB84B405AAF3168F922704">Records of the Australian Museum</mods:title>
</mods:titleInfo>
<mods:part id="395F7161F011C17A324A555BE4D7796D">
<mods:date id="B5D020DB81725774B89481882E984F76">2016</mods:date>
<mods:detail id="48034DF518F793B08694E1D0B3A861F5" type="series">
<mods:title id="2CA6478F578456A9E7831D6CBE43BA37">Rec. Aust. Mus.</mods:title>
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<mods:detail id="DF32076664ABBBA525AA2A451738EA79" type="pubDate">
<mods:number id="4D8DE32D79E7D2CE8FCD1D88FB45755A">2016-11-30</mods:number>
</mods:detail>
<mods:detail id="DCD71D30A61B8988CB26BDFDD10066F6" type="volume">
<mods:number id="511F6388F917FCEB96799543B2AD069C">68</mods:number>
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<mods:detail id="59186B1BA8545A5F85D7FF5914D2B447" type="issue">
<mods:number id="D60679F2073622BCCA0582136F697965">6</mods:number>
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<mods:start id="E9E228DFC74D819D2DA97755D05B2BCB">263</mods:start>
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<mods:url id="D767CF79884AA60AA1E59BA8944E1006">http://dx.doi.org/10.3853/j.2201-4349.68.2016.1659</mods:url>
</mods:location>
<mods:classification id="B4C5F685D4ACA6F4C76279E4A6DEF462">journal article</mods:classification>
<mods:identifier id="A3AA0865E98E7BC88E15DF29CF71E850" type="DOI">10.3853/j.2201-4349.68.2016.1659</mods:identifier>
<mods:identifier id="D4FF128B50257333D917E361C0F47F7E" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="4008229C4F5CF7E9BF653EEDBE029BD7" type="Zenodo-Dep">4676914</mods:identifier>
<mods:identifier id="AFD551ECFBBB59453AD06A18608A1732" type="ZooBank">246FBE46-E573-47C9-84C2-A08322EE92C5</mods:identifier>
</mods:mods>
<treatment id="5552878F9B76FF9B01CDFD39FE03FE53" LSID="urn:lsid:zoobank.org:act:1D4880EC-F83C-4BE8-9A23-5EABE927978B" httpUri="http://treatment.plazi.org/id/5552878F9B76FF9B01CDFD39FE03FE53" lastPageId="4" lastPageNumber="267" pageId="1" pageNumber="264">
<subSubSection id="95E165129B76FF9E01CDFD39FD21FD40" box="[227,643,677,704]" pageId="1" pageNumber="264" type="nomenclature">
<paragraph id="DD4436999B76FF9E01CDFD39FD21FD40" blockId="1.[227,643,677,704]" box="[227,643,677,704]" pageId="1" pageNumber="264">
<heading id="860C81F59B76FF9E01CDFD39FD21FD40" box="[227,643,677,704]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="264" reason="2">
<emphasis id="EF8FEA8B9B76FF9E01CDFD39FD21FD40" bold="true" box="[227,643,677,704]" pageId="1" pageNumber="264">
<taxonomicName id="1AFB4D1A9B76FF9E01CDFD39FD95FD40" authority="Batley" authorityName="Batley" authorityYear="2016" box="[227,567,677,704]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B76FF9E01CDFD39FE7FFD3F" bold="true" box="[227,477,677,704]" italics="true" pageId="1" pageNumber="264">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B76FF9E0311FD39FD21FD40" box="[575,643,677,703]" pageId="1" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="95E165129B76FF9E0157FD49FE8EFC80" pageId="1" pageNumber="264" type="materials_examined">
<paragraph id="DD4436999B76FF9E0157FD49FE8EFC80" blockId="1.[121,749,724,895]" pageId="1" pageNumber="264">
<emphasis id="EF8FEA8B9B76FF9E0157FD49FE87FD13" bold="true" box="[121,293,724,749]" pageId="1" pageNumber="264">Type specimens</emphasis>
.
<materialsCitation id="6D933CC49B76FF9E001EFD48FE6EFCD8" collectingDate="2012-04-21" collectorName="M. Batley" latitude="-23.738" location="Ethabuka Station" longLatPrecision="74" longitude="138.467" pageId="1" pageNumber="264" specimenCount="1" specimenCount-female="1" stateProvince="Queensland" typeStatus="holotype">
<typeStatus id="0240883B9B76FF9E001EFD48FE33FD13" box="[304,401,724,748]" pageId="1" pageNumber="264" type="holotype">Holotype</typeStatus>
<specimenCount id="CBFDFD109B76FF9E00B8FD49FE08FD13" box="[406,426,725,748]" pageId="1" pageNumber="264" type="female">♀</specimenCount>
,
<location id="D82460429B76FF9E009BFD48FDCAFD13" LSID="urn:lsid:plazi:treatment:5552878F9B76FF9B01CDFD39FE03FE53:D82460429B76FF9E009BFD48FDCAFD13" box="[437,616,724,748]" latitude="-23.738" longLatPrecision="74" longitude="138.467" name="Ethabuka Station" pageId="1" pageNumber="264" stateProvince="Queensland">Ethabuka Station</location>
,
<collectingRegion id="1F3FF87B9B76FF9E035FFD48FD4FFD13" box="[625,749,724,748]" country="Australia" name="Queensland" pageId="1" pageNumber="264">Queensland</collectingRegion>
(
<geoCoordinate id="B8CF505E9B76FF9E01AEFD6EFF46FCF5" box="[128,228,754,778]" degrees="23.738" direction="south" orientation="latitude" pageId="1" pageNumber="264" precision="55" value="-23.738">23.738°S</geoCoordinate>
<geoCoordinate id="B8CF505E9B76FF9E01C7FD6EFEFCFCF5" box="[233,350,754,778]" degrees="138.467" direction="east" orientation="longitude" pageId="1" pageNumber="264" precision="55" value="138.467">138.467°E</geoCoordinate>
),
<date id="A94510599B76FF9E0041FD6EFE51FCF5" box="[367,499,754,778]" pageId="1" pageNumber="264" value="2012-04-21">
<collectingDate id="B901E9B19B76FF9E0041FD6EFE51FCF5" box="[367,499,754,778]" pageId="1" pageNumber="264" value="2012-04-21">21 Apr 2012</collectingDate>
</date>
,
<collectorName id="700E534F9B76FF9E00D2FD6EFDC4FCF5" box="[508,614,754,778]" pageId="1" pageNumber="264">M. Batley</collectorName>
,
<emphasis id="EF8FEA8B9B76FF9E0340FD6EFEA2FCD8" italics="true" pageId="1" pageNumber="264">
ex
<taxonomicName id="1AFB4D1A9B76FF9E03A2FD6EFEA2FCD8" class="Magnoliopsida" family="Goodeniaceae" genus="Scaevola" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="depauperata">Scaevola depauperata</taxonomicName>
</emphasis>
in AM (K363485)
</materialsCitation>
.
<materialsCitation id="6D933CC49B76FF9E00F8FC8CFE8AFC80" collectingDate="2014-04-30" collectingDateMax="2014-05-09" collectingDateMin="2014-04-30" collectorName="M. Batley" latitude="-23.714" location="Ethabuka Station" longLatPrecision="74" longitude="138.453" pageId="1" pageNumber="264" specimenCount="2" specimenCount-male="2" stateProvince="Queensland" typeStatus="paratype">
<typeStatus id="0240883B9B76FF9E00F8FC8CFD9CFCD7" box="[470,574,784,808]" pageId="1" pageNumber="264" type="paratype">Paratypes</typeStatus>
<specimenCount id="CBFDFD109B76FF9E036AFC93FD23FCD8" box="[580,641,783,807]" pageId="1" pageNumber="264" type="male">2 ♂♂</specimenCount>
<location id="D82460429B76FF9E03A6FC93FF68FCBB" LSID="urn:lsid:plazi:treatment:5552878F9B76FF9B01CDFD39FE03FE53:D82460429B76FF9E03A6FC93FF68FCBB" latitude="-23.714" longLatPrecision="74" longitude="138.453" name="Ethabuka Station" pageId="1" pageNumber="264" stateProvince="Queensland">Ethabuka Station</location>
,
<collectingRegion id="1F3FF87B9B76FF9E01F6FCB0FEFBFCBB" box="[216,345,812,836]" country="Australia" name="Queensland" pageId="1" pageNumber="264">Queensland</collectingRegion>
(
<geoCoordinate id="B8CF505E9B76FF9E0042FCB1FE70FCBA" box="[364,466,813,837]" degrees="23.714" direction="south" orientation="latitude" pageId="1" pageNumber="264" precision="55" value="-23.714">23.714°S</geoCoordinate>
<geoCoordinate id="B8CF505E9B76FF9E00F3FCB0FDF4FCBB" box="[477,598,812,836]" degrees="138.453" direction="east" orientation="longitude" pageId="1" pageNumber="264" precision="55" value="138.453">138.453°E</geoCoordinate>
),
<date id="A94510599B76FF9E0342FCB1FEA0FC9D" pageId="1" pageNumber="264" value="2014-04-30" valueMax="2014-05-09" valueMin="2014-04-30">
<collectingDate id="B901E9B19B76FF9E0342FCB1FEA0FC9D" pageId="1" pageNumber="264" value="2014-04-30" valueMax="2014-05-09" valueMin="2014-04-30">30 Apr and 9 May 2014</collectingDate>
</date>
,
<collectorName id="700E534F9B76FF9E0021FCD6FEDCFC9D" box="[271,382,842,866]" pageId="1" pageNumber="264">M. Batley</collectorName>
,
<emphasis id="EF8FEA8B9B76FF9E00A4FCD6FD3EFC9D" box="[394,668,842,866]" italics="true" pageId="1" pageNumber="264">
ex
<taxonomicName id="1AFB4D1A9B76FF9E0085FCD6FD3EFC9D" box="[427,668,842,866]" class="Magnoliopsida" family="Goodeniaceae" genus="Scaevola" kingdom="Plantae" order="Asterales" pageId="1" pageNumber="264" phylum="Tracheophyta" rank="species" species="depauperata">Scaevola depauperata</taxonomicName>
</emphasis>
in AM (K470078–079)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="95E165129B76FF9E005BFC09FEECFB55" pageId="1" pageNumber="264" type="diagnosis">
<paragraph id="DD4436999B76FF9E005BFC09FE52FC50" blockId="1.[373,496,917,943]" box="[373,496,917,943]" pageId="1" pageNumber="264">
<emphasis id="EF8FEA8B9B76FF9E005BFC09FE52FC50" bold="true" box="[373,496,917,943]" pageId="1" pageNumber="264">Diagnosis</emphasis>
</paragraph>
<paragraph id="DD4436999B76FF9E0157FC58FEECFB55" blockId="1.[119,749,964,1194]" pageId="1" pageNumber="264">
Small, moderately hairy bees with black head and mesosoma and orange-brown metasoma. Forewing with two submarginal cells, jugal lobe of the hind wing extending well beyond cu-v (
<figureCitation id="45C02A1C9B76FF9E0025FB80FEFAFBCA" box="[267,344,1052,1077]" captionStart="Figure 1" captionStartId="1.[128,191,589,611]" captionTargetBox="[121,1419,151,581]" captionTargetId="figure-531@1.[121,1420,151,581]" captionTargetPageId="1" captionText="Figure 1. Wing venation of Leioproctus glendae Batley n. sp. male. (a) Hind wing in focus to show jugal lobe; (b) fore wing in focus." figureDoi="http://doi.org/10.5281/zenodo.4676916" httpUri="https://zenodo.org/record/4676916/files/figure.png" pageId="1" pageNumber="264">Fig. 1a</figureCitation>
). Females possess a narrow, flattened fore basitarsus (
<figureCitation id="45C02A1C9B76FF9E0000FBA6FED1FBAD" box="[302,371,1082,1106]" captionStart="Figure 4" captionStartId="2.[253,316,1372,1394]" captionTargetBox="[217,1415,755,1363]" captionTargetId="figure-41@2.[217,1418,755,1363]" captionTargetPageId="2" captionText="Figure 4. Female Leioproctus glendae Batley n. sp. (a) fore tarsus lateral view; (b) front view; and (c) fore tibial spur." figureDoi="http://doi.org/10.5281/zenodo.4676926" httpUri="https://zenodo.org/record/4676926/files/figure.png" pageId="1" pageNumber="264">Fig. 4</figureCitation>
) and mandibles unlike any other
<taxonomicName id="1AFB4D1A9B76FF9E0157FBC4FF55FB90" box="[121,247,1112,1135]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="genus">
<emphasis id="EF8FEA8B9B76FF9E0157FBC4FF55FB90" box="[121,247,1112,1135]" italics="true" pageId="1" pageNumber="264">Leioproctus</emphasis>
</taxonomicName>
species (
<figureCitation id="45C02A1C9B76FF9E0077FBCBFE38FB90" box="[345,410,1111,1135]" captionStart="Figure 3" captionStartId="2.[509,572,653,675]" captionTargetBox="[166,1466,151,642]" captionTargetId="figure-18@2.[166,1466,151,642]" captionTargetPageId="2" captionText="Figure 3. Mandible of Leioproctus glendae Batley n. sp. female." figureDoi="http://doi.org/10.5281/zenodo.4676922" httpUri="https://zenodo.org/record/4676922/files/figure.png" pageId="1" pageNumber="264">Fig. 3</figureCitation>
). Males have similar mandibles and a unique eighth metasomal sternum (
<figureCitation id="45C02A1C9B76FF9E0333FBE8FDC4FB72" box="[541,614,1140,1165]" captionStart="Figure 8" captionStartId="3.[387,450,1944,1966]" captionTargetBox="[388,1155,1571,1933]" captionTargetId="figure-419@3.[388,1156,1571,1933]" captionTargetPageId="3" captionText="Figure 8. Male Leioproctus glendae Batley n. sp. (a) S8; and (b) ventral view of" figureDoi="http://doi.org/10.5281/zenodo.4676940" httpUri="https://zenodo.org/record/4676940/files/figure.png" pageId="1" pageNumber="264">Fig. 8a</figureCitation>
) whose apex is visible externally.
</paragraph>
</subSubSection>
<subSubSection id="95E165129B76FF9C0047FB5CFAB4FD7B" lastPageId="3" lastPageNumber="266" pageId="1" pageNumber="264" type="description">
<paragraph id="DD4436999B76FF9E0047FB5CFE5FFB25" blockId="1.[361,509,1216,1242]" box="[361,509,1216,1242]" pageId="1" pageNumber="264">
<emphasis id="EF8FEA8B9B76FF9E0047FB5CFE5FFB25" bold="true" box="[361,509,1216,1242]" pageId="1" pageNumber="264">Description</emphasis>
</paragraph>
<paragraph id="DD4436999B76FF9C0157FB73FE92FE48" blockId="1.[119,749,1263,1610]" lastBlockId="3.[119,749,151,733]" lastPageId="3" lastPageNumber="266" pageId="1" pageNumber="264">
Female—Head width
<quantity id="1A039B7C9B76FF9E0074FB73FE15FAF8" box="[346,439,1263,1287]" metricMagnitude="-3" metricUnit="m" metricValue="1.83" pageId="1" pageNumber="264" unit="mm" value="1.83">1.83 mm</quantity>
, body length
<quantity id="1A039B7C9B76FF9E0313FB73FD2CFAF8" box="[573,654,1263,1287]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="1" pageNumber="264" unit="mm" value="5.8">5.8 mm</quantity>
. Relative measurements: HW 50, HL 47, UID 33, LID 25, SL 16, CW 26, CL 16, FL 27.
<emphasis id="EF8FEA8B9B76FF9E0061FAB6FE30FABD" box="[335,402,1322,1346]" italics="true" pageId="1" pageNumber="264">Head:</emphasis>
black, except antenna, labrum, mandible and ventral margin of clypeus orange brown. Clypeus extends well below mandibular articulations giving face an elongate appearance, anterior surface projected in front of eyes, lateral edges strongly convex, epistomal suture indistinct; face with broad, shallow depression from anterior ocellus to ventral margin of clypeus. Mandible with condylar ridge and outer ridge very narrow and strongly projected from outer surface leaving a large, flat outer interspace, below each ridge is a line of long, weakly-branched setae; cap of rutellum thick and greatly elongated into a dagger-like tooth (
<figureCitation id="45C02A1C9B76FF9E0410FD3AFADCFD41" box="[1342,1406,678,702]" captionStart="Figure 3" captionStartId="2.[509,572,653,675]" captionTargetBox="[166,1466,151,642]" captionTargetId="figure-18@2.[166,1466,151,642]" captionTargetPageId="2" captionText="Figure 3. Mandible of Leioproctus glendae Batley n. sp. female." figureDoi="http://doi.org/10.5281/zenodo.4676922" httpUri="https://zenodo.org/record/4676922/files/figure.png" pageId="1" pageNumber="264">Fig. 3</figureCitation>
). Head densely punctate except in medial depression and small areas of vertex adjacent to posterior ocelli. Paraocular areas and frons, except medially, densely covered with long, white, finely-branched hair. Scape short reaching a bit over ½ way to anterior ocellus. Labrum approximately triangular, width
<emphasis id="EF8FEA8B9B76FF9E045DFCA5FA2EFCAF" box="[1395,1420,825,848]" italics="true" pageId="1" pageNumber="264">ca</emphasis>
2.5× length, gently convex, polished, with horizontal carina and stiff setae on the inflexed surface below the carina. Malar space obsolete. Maxillary palpus short (
<emphasis id="EF8FEA8B9B76FF9E0581FC0DFB6AFC57" box="[1199,1224,913,936]" italics="true" pageId="1" pageNumber="264">ca</emphasis>
0.25× head width), labial palpus elongate and flattened, exceeding end of glossa (
<emphasis id="EF8FEA8B9B76FF9E020CFC50FC99FC1C" box="[802,827,972,995]" italics="true" pageId="1" pageNumber="264">ca</emphasis>
0.5× head with, ratio of segments 17:13:12:10 starting from base).
<emphasis id="EF8FEA8B9B76FF9E02B9FC75FBB3FBFF" box="[919,1041,1001,1024]" italics="true" pageId="1" pageNumber="264">Mesosoma:</emphasis>
black with legs orange-brown. Fore basitarsus greatly flattened and elongated (
<emphasis id="EF8FEA8B9B76FF9E05EBFB9AFB7CFBE2" box="[1221,1246,1030,1053]" italics="true" pageId="1" pageNumber="264">ca</emphasis>
¾ as long as tibia and twice as long as remainder of tarsus, excluding claws), bearing widely spaced long hairs (
<figureCitation id="45C02A1C9B76FF9E0550FBDCFB41FBA7" box="[1150,1251,1088,1112]" captionStart="Figure 4" captionStartId="2.[253,316,1372,1394]" captionTargetBox="[217,1415,755,1363]" captionTargetId="figure-41@2.[217,1418,755,1363]" captionTargetPageId="2" captionText="Figure 4. Female Leioproctus glendae Batley n. sp. (a) fore tarsus lateral view; (b) front view; and (c) fore tibial spur." figureDoi="http://doi.org/10.5281/zenodo.4676926" httpUri="https://zenodo.org/record/4676926/files/figure.png" pageId="1" pageNumber="264">Fig. 4a, b</figureCitation>
); fore tibial spur modified (
<figureCitation id="45C02A1C9B76FF9E02ABFBC2FC71FB89" box="[901,979,1118,1142]" captionStart="Figure 4" captionStartId="2.[253,316,1372,1394]" captionTargetBox="[217,1415,755,1363]" captionTargetId="figure-41@2.[217,1418,755,1363]" captionTargetPageId="2" captionText="Figure 4. Female Leioproctus glendae Batley n. sp. (a) fore tarsus lateral view; (b) front view; and (c) fore tibial spur." figureDoi="http://doi.org/10.5281/zenodo.4676926" httpUri="https://zenodo.org/record/4676926/files/figure.png" pageId="1" pageNumber="264">Fig. 4c</figureCitation>
); inner hind tibial spur finely serrate (
<figureCitation id="45C02A1C9B76FF9E044FFBC2FC97FB6C" captionStart="Figure 5" captionStartId="2.[262,325,1892,1914]" captionTargetBox="[166,1464,1484,1880]" captionTargetId="figure-61@2.[166,1465,1484,1880]" captionTargetPageId="2" captionText="Figure 5. Female Leioproctus glendae Batley n. sp. (a) inner hind tibial spur; (b) propodeum; and (c) pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.4676928" httpUri="https://zenodo.org/record/4676928/files/figure.png" pageId="1" pageNumber="264">Fig. 5a</figureCitation>
); claws cleft with inner tooth slightly smaller than outer; hind basitibial plate carinate, well-defined with open cover of minutely-branched hair; scopae formed from combination of long, plumose hair on hind trochanter and basal half of hind femur, openly spaced, weakly-branched hair on outer face of hind tibia and open, erect hair on sterna, widely-branched on
<geoCoordinate id="B8CF505E9B76FF9E0214FAB7FCD1FABC" box="[826,883,1323,1347]" degrees="2, 3" direction="south" orientation="latitude" pageId="1" pageNumber="264" precision="5555" value="-2.3">S2, 3</geoCoordinate>
, simple on
<geoCoordinate id="B8CF505E9B76FF9E02C1FAB7FB8AFABC" box="[1007,1064,1323,1347]" degrees="4, 5" direction="south" orientation="latitude" pageId="1" pageNumber="264" precision="5555" value="-4.5">S4, 5</geoCoordinate>
. Pronotal collar thin, much lower than scutum; metanotum with small tubercle; subhorizontal surface of propodeum about as long as metanotum, rounding smoothly onto vertical surface. Surface polished with close, strong punctures except in propodeal triangle which is weakly transversely striate with large areolae lateromarginally. Most of mesosoma other than propodeum closely covered with moderately long, strongly-branched, white hair; scutellum and posterior half of scutum mostly bare, possibly as a result of wear.
<emphasis id="EF8FEA8B9B76FF9E0243F9A8FBABF9B4" box="[877,1033,1588,1611]" italics="true" pageId="1" pageNumber="264">Wing venation:</emphasis>
fore wing with two submarginal cells, first recurrent vein entering second submarginal cell; stigma broad,
<emphasis id="EF8FEA8B9B74FF9C01EDFF2AFF7EFF32" box="[195,220,182,205]" italics="true" pageId="3" pageNumber="266">ca</emphasis>
½ as long as costal margin of marginal cell; apex of marginal cell well away from costa; jugal lobe of hind wing reaches well beyond cu-v (similar to that of male,
<figureCitation id="45C02A1C9B74FF9C03EDFF73FF30FEDA" captionStart="Figure 1" captionStartId="1.[128,191,589,611]" captionTargetBox="[121,1419,151,581]" captionTargetId="figure-531@1.[121,1420,151,581]" captionTargetPageId="1" captionText="Figure 1. Wing venation of Leioproctus glendae Batley n. sp. male. (a) Hind wing in focus to show jugal lobe; (b) fore wing in focus." figureDoi="http://doi.org/10.5281/zenodo.4676916" httpUri="https://zenodo.org/record/4676916/files/figure.png" pageId="3" pageNumber="266">Fig. 1a</figureCitation>
).
<emphasis id="EF8FEA8B9B74FF9C0188FE92FEB6FEDA" box="[166,276,270,293]" italics="true" pageId="3" pageNumber="266">Metasoma</emphasis>
orange-brown, slightly darker ventrally, terga translucent apically, strongly and densely punctate with a fine covering of long, adpressed, white, minutely-branched hair; T5,6 with dense, pale prepygidial and pygidial fimbria; pygidial plate weakly emarginate, carinate with medial area acinose (
<figureCitation id="45C02A1C9B74FF9C01F8FE03FE81FE48" box="[214,291,415,439]" captionStart="Figure 5" captionStartId="2.[262,325,1892,1914]" captionTargetBox="[166,1464,1484,1880]" captionTargetId="figure-61@2.[166,1465,1484,1880]" captionTargetPageId="2" captionText="Figure 5. Female Leioproctus glendae Batley n. sp. (a) inner hind tibial spur; (b) propodeum; and (c) pygidial plate." figureDoi="http://doi.org/10.5281/zenodo.4676928" httpUri="https://zenodo.org/record/4676928/files/figure.png" pageId="3" pageNumber="266">Fig. 5c</figureCitation>
).
</paragraph>
<caption id="898466119B76FF9E009CF82FFBF1F836" ID-DOI="http://doi.org/10.5281/zenodo.4676920" ID-Zenodo-Dep="4676920" box="[434,1107,1971,1993]" httpUri="https://zenodo.org/record/4676920/files/figure.png" pageId="1" pageNumber="264" startId="1.[434,497,1971,1993]" targetBox="[122,1421,1663,1957]" targetPageId="1">
<paragraph id="DD4436999B76FF9E009CF82FFBF1F836" blockId="1.[434,1107,1971,1993]" box="[434,1107,1971,1993]" pageId="1" pageNumber="264">
Figure 2. Habitus images of
<taxonomicName id="1AFB4D1A9B76FF9E03EDF828FC6EF836" authority="Batley" authorityName="Batley" authorityYear="2016" box="[707,972,1971,1993]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="1" pageNumber="264" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B76FF9E03EDF828FC25F837" box="[707,903,1971,1993]" italics="true" pageId="1" pageNumber="264">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B76FF9E02FCF82FFBA4F837" box="[978,1030,1971,1992]" pageId="1" pageNumber="264" rank="species">n. sp.</taxonomicNameLabel>
female.
</paragraph>
</caption>
<caption id="898466119B75FF9D00D3FD11FBC7FD5C" ID-DOI="http://doi.org/10.5281/zenodo.4676922" ID-Zenodo-Dep="4676922" box="[509,1125,653,675]" httpUri="https://zenodo.org/record/4676922/files/figure.png" pageId="2" pageNumber="265" startId="2.[509,572,653,675]" targetBox="[166,1466,151,642]" targetPageId="2">
<paragraph id="DD4436999B75FF9D00D3FD11FBC7FD5C" blockId="2.[509,1125,653,675]" box="[509,1125,653,675]" pageId="2" pageNumber="265">
Figure 3. Mandible of
<taxonomicName id="1AFB4D1A9B75FF9D03FBFD12FC7CFD5C" authority="Batley" authorityName="Batley" authorityYear="2016" box="[725,990,653,675]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B75FF9D03FBFD12FC3BFD5D" box="[725,921,653,675]" italics="true" pageId="2" pageNumber="265">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B75FF9D02CAFD12FBBAFD5C" box="[996,1048,654,675]" pageId="2" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
female.
</paragraph>
</caption>
<caption id="898466119B75FF9D01D3FAC0FAC4FA8D" ID-DOI="http://doi.org/10.5281/zenodo.4676926" ID-Zenodo-Dep="4676926" box="[253,1382,1372,1394]" httpUri="https://zenodo.org/record/4676926/files/figure.png" pageId="2" pageNumber="265" startId="2.[253,316,1372,1394]" targetBox="[217,1415,755,1363]" targetPageId="2">
<paragraph id="DD4436999B75FF9D01D3FAC0FAC4FA8D" blockId="2.[253,1382,1372,1394]" box="[253,1382,1372,1394]" pageId="2" pageNumber="265">
Figure 4. Female
<taxonomicName id="1AFB4D1A9B75FF9D0088FAC1FD0DFA8D" authority="Batley" authorityName="Batley" authorityYear="2016" box="[422,687,1372,1394]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B75FF9D0088FAC1FDC8FA8E" box="[422,618,1372,1394]" italics="true" pageId="2" pageNumber="265">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B75FF9D039BFAC1FD4BFA8D" box="[693,745,1373,1394]" pageId="2" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="EF8FEA8B9B75FF9D03C0FAC0FCA8FA8E" box="[750,778,1372,1393]" italics="true" pageId="2" pageNumber="265">(a)</emphasis>
fore tarsus lateral view;
<emphasis id="EF8FEA8B9B75FF9D02D6FAC0FBB6FA8E" box="[1016,1044,1372,1393]" italics="true" pageId="2" pageNumber="265">(b)</emphasis>
front view; and
<emphasis id="EF8FEA8B9B75FF9D059CFAC0FB6FFA8E" box="[1202,1229,1372,1393]" italics="true" pageId="2" pageNumber="265">(c)</emphasis>
fore tibial spur.
</paragraph>
</caption>
<caption id="898466119B75FF9D0028F8F8FAFCF885" ID-DOI="http://doi.org/10.5281/zenodo.4676928" ID-Zenodo-Dep="4676928" box="[262,1374,1891,1914]" httpUri="https://zenodo.org/record/4676928/files/figure.png" pageId="2" pageNumber="265" startId="2.[262,325,1892,1914]" targetBox="[166,1464,1484,1880]" targetPageId="2">
<paragraph id="DD4436999B75FF9D0028F8F8FAFCF885" blockId="2.[262,1374,1891,1914]" box="[262,1374,1891,1914]" pageId="2" pageNumber="265">
Figure 5. Female
<taxonomicName id="1AFB4D1A9B75FF9D0081F8F8FD1AF885" authority="Batley" authorityName="Batley" authorityYear="2016" box="[431,696,1892,1914]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="265" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B75FF9D0081F8F8FDD1F886" box="[431,627,1892,1913]" italics="true" pageId="2" pageNumber="265">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B75FF9D0390F8F8FD50F886" box="[702,754,1892,1913]" pageId="2" pageNumber="265" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="EF8FEA8B9B75FF9D03D9F8F8FCB1F886" box="[759,787,1892,1913]" italics="true" pageId="2" pageNumber="265">(a)</emphasis>
inner hind tibial spur;
<emphasis id="EF8FEA8B9B75FF9D02C0F8F8FBA9F886" box="[1006,1035,1892,1913]" italics="true" pageId="2" pageNumber="265">(b)</emphasis>
propodeum; and
<emphasis id="EF8FEA8B9B75FF9D059DF8F8FB6CF886" box="[1203,1230,1892,1913]" italics="true" pageId="2" pageNumber="265">(c)</emphasis>
pygidial plate.
</paragraph>
</caption>
<paragraph id="DD4436999B74FF9C01B9FE21FAB4FD7B" blockId="3.[119,749,151,733]" lastBlockId="3.[792,1428,152,645]" pageId="3" pageNumber="266">
Male—Head width 1.74,
<quantity id="1A039B7C9B74FF9C0086FE21FDA8FE2B" box="[424,522,445,469]" metricMagnitude="-3" metricUnit="m" metricValue="1.84" pageId="3" pageNumber="266" unit="mm" value="1.84">1.84 mm</quantity>
, body length 5.5,
<quantity id="1A039B7C9B74FF9C03E5FE21FF05FE0D" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="266" unit="mm" value="5.8">5.8 mm</quantity>
. Relative measurements: HW 50, HL 45, UID 32, LID 24, SL 11, CW 24, CL 18, FL 30 (±1 for both specimens).
<emphasis id="EF8FEA8B9B74FF9C0157FD89FF17FDD2" box="[121,181,533,557]" italics="true" pageId="3" pageNumber="266">Head</emphasis>
: as for female except for following: facial hair somewhat longer and extending onto clypeus and scapes; width labrum
<emphasis id="EF8FEA8B9B74FF9C0028FDCCFEBDFD98" box="[262,287,592,615]" italics="true" pageId="3" pageNumber="266">ca</emphasis>
4× length, apical ½ sharply deflexed, bearing stiff setae; mandible broad basally, tapering evenly to a single tooth, cap of rutellum not thick and extended; labial palpus
<emphasis id="EF8FEA8B9B74FF9C0157FD34FF30FD40" box="[121,146,680,703]" italics="true" pageId="3" pageNumber="266">ca</emphasis>
0.6× head with, ratio of segments 18:16:15:11 starting from base.
<emphasis id="EF8FEA8B9B74FF9C01DEFD5AFEC3FD22" box="[240,353,710,733]" italics="true" pageId="3" pageNumber="266">Mesosoma</emphasis>
as for female except hair a bit longer and very pale brown.
<emphasis id="EF8FEA8B9B74FF9C02D9FF04FB34FF50" box="[1015,1174,152,175]" italics="true" pageId="3" pageNumber="266">Wing venation:</emphasis>
as described for female (
<figureCitation id="45C02A1C9B74FF9C020EFF29FCC0FF32" box="[800,866,181,205]" captionStart="Figure 1" captionStartId="1.[128,191,589,611]" captionTargetBox="[121,1419,151,581]" captionTargetId="figure-531@1.[121,1420,151,581]" captionTargetPageId="1" captionText="Figure 1. Wing venation of Leioproctus glendae Batley n. sp. male. (a) Hind wing in focus to show jugal lobe; (b) fore wing in focus." figureDoi="http://doi.org/10.5281/zenodo.4676916" httpUri="https://zenodo.org/record/4676916/files/figure.png" pageId="3" pageNumber="266">Fig. 1</figureCitation>
).
<emphasis id="EF8FEA8B9B74FF9C0250FF2AFC12FF32" box="[894,944,182,205]" italics="true" pageId="3" pageNumber="266">Legs</emphasis>
: dark brown basally, mostly orange-brown distally from apices of femora, outer face of tibiae with dark brown suffusions; claws cleft, teeth of equal size; inner hind tibial spur finely serrate. Lengths of basitarsi ½ length corresponding femora; hind basitibial area small, defined by a fine carina, more easily observed with transmitted light.
<emphasis id="EF8FEA8B9B74FF9C0234FEFAFC28FE82" box="[794,906,358,381]" italics="true" pageId="3" pageNumber="266">Metasoma</emphasis>
orange-brown, slightly darker ventrally, terga translucent apically, strongly and densely punctate with a fine covering of long, adpressed, pale brown, minutely-branched hair. Posterior margin S6 with small medial emargination (
<figureCitation id="45C02A1C9B74FF9C020CFE46FCCDFE0D" box="[802,879,474,498]" captionStart="Figure 8" captionStartId="3.[387,450,1944,1966]" captionTargetBox="[388,1155,1571,1933]" captionTargetId="figure-419@3.[388,1156,1571,1933]" captionTargetPageId="3" captionText="Figure 8. Male Leioproctus glendae Batley n. sp. (a) S8; and (b) ventral view of" figureDoi="http://doi.org/10.5281/zenodo.4676940" httpUri="https://zenodo.org/record/4676940/files/figure.png" pageId="3" pageNumber="266">Fig. 8b</figureCitation>
). Vestiture as in female except somewhat longer and denser.
<emphasis id="EF8FEA8B9B74FF9C0242FE64FC4AFDEF" box="[876,1000,504,528]" italics="true" pageId="3" pageNumber="266">Terminalia:</emphasis>
as shown in
<figureCitation id="45C02A1C9B74FF9C0559FE64FB1EFDF0" box="[1143,1212,504,528]" captionStart="Figure 7" captionStartId="3.[199,262,1506,1528]" captionTargetBox="[122,1421,1131,1493]" captionTargetId="figure-398@3.[122,1421,1131,1493]" captionTargetPageId="3" captionText="Figure 7. Terminalia of Leioproctus glendae Batley n. sp. male (a) genital capsule; (b) apex of penis valves; and (c) S7." figureDoi="http://doi.org/10.5281/zenodo.4676938" httpUri="https://zenodo.org/record/4676938/files/figure.png" pageId="3" pageNumber="266">Figs 7</figureCitation>
,
<figureCitation id="45C02A1C9B74FF9C05E7FE64FB7AFDEF" box="[1225,1240,504,528]" captionStart="Figure 8" captionStartId="3.[387,450,1944,1966]" captionTargetBox="[388,1155,1571,1933]" captionTargetId="figure-419@3.[388,1156,1571,1933]" captionTargetPageId="3" captionText="Figure 8. Male Leioproctus glendae Batley n. sp. (a) S8; and (b) ventral view of" figureDoi="http://doi.org/10.5281/zenodo.4676940" httpUri="https://zenodo.org/record/4676940/files/figure.png" pageId="3" pageNumber="266">8</figureCitation>
; S
<figureCitation id="45C02A1C9B74FF9C05DDFE64FAA3FDEF" box="[1267,1281,504,528]" captionStart="Figure 7" captionStartId="3.[199,262,1506,1528]" captionTargetBox="[122,1421,1131,1493]" captionTargetId="figure-398@3.[122,1421,1131,1493]" captionTargetPageId="3" captionText="Figure 7. Terminalia of Leioproctus glendae Batley n. sp. male (a) genital capsule; (b) apex of penis valves; and (c) S7." figureDoi="http://doi.org/10.5281/zenodo.4676938" httpUri="https://zenodo.org/record/4676938/files/figure.png" pageId="3" pageNumber="266">7</figureCitation>
with simple, greatly reduced posterior lobes and a few simple setae; S8 has a short, bifid apical process, which is exposed externally; genital capsule has strongly hairy gonoforceps and penis valves with broad transparent wings at the apex.
</paragraph>
</subSubSection>
<subSubSection id="95E165129B74FF9C0234FD34FD38F836" pageId="3" pageNumber="266" type="etymology">
<paragraph id="DD4436999B74FF9C0234FD34FAEAFD22" blockId="3.[792,1421,680,733]" pageId="3" pageNumber="266">
<emphasis id="EF8FEA8B9B74FF9C0234FD34FC36FD3F" bold="true" box="[794,916,680,704]" pageId="3" pageNumber="266">Etymology</emphasis>
. The species is named after Prof. Glenda Wardle in recognition of her contributions to desert ecology.
</paragraph>
<caption id="898466119B74FF9C0092FBAFFBE9FBB6" ID-DOI="http://doi.org/10.5281/zenodo.4676932" ID-Zenodo-Dep="4676932" box="[444,1099,1075,1097]" httpUri="https://zenodo.org/record/4676932/files/figure.png" pageId="3" pageNumber="266" startId="3.[444,507,1075,1097]" targetBox="[145,1420,790,1061]" targetPageId="3">
<paragraph id="DD4436999B74FF9C0092FBAFFBE9FBB6" blockId="3.[444,1099,1075,1097]" box="[444,1099,1075,1097]" pageId="3" pageNumber="266">
Figure 6. Habitus images of
<taxonomicName id="1AFB4D1A9B74FF9C03E0FBA8FC74FBB6" authority="Batley" authorityName="Batley" authorityYear="2016" box="[718,982,1075,1097]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B74FF9C03E0FBA8FC33FBB7" box="[718,913,1075,1097]" italics="true" pageId="3" pageNumber="266">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B74FF9C02F2FBA8FBB2FBB6" box="[988,1040,1076,1097]" pageId="3" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
male.
</paragraph>
</caption>
<caption id="898466119B74FF9C01E9FA7EFAE2FA07" ID-DOI="http://doi.org/10.5281/zenodo.4676938" ID-Zenodo-Dep="4676938" box="[199,1344,1506,1528]" httpUri="https://zenodo.org/record/4676938/files/figure.png" pageId="3" pageNumber="266" startId="3.[199,262,1506,1528]" targetBox="[122,1421,1131,1493]" targetPageId="3">
<paragraph id="DD4436999B74FF9C01E9FA7EFAE2FA07" blockId="3.[199,1344,1506,1528]" box="[199,1344,1506,1528]" pageId="3" pageNumber="266">
Figure 7. Terminalia of
<taxonomicName id="1AFB4D1A9B74FF9C0085FA7FFD11FA07" authority="Batley" authorityName="Batley" authorityYear="2016" box="[427,691,1506,1528]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B74FF9C0085FA7FFDCCFA08" box="[427,622,1506,1528]" italics="true" pageId="3" pageNumber="266">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B74FF9C0397FA7FFD4FFA07" box="[697,749,1507,1528]" pageId="3" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
male
<emphasis id="EF8FEA8B9B74FF9C0209FA7FFCE1FA07" box="[807,835,1507,1528]" italics="true" pageId="3" pageNumber="266">(a)</emphasis>
genital capsule;
<emphasis id="EF8FEA8B9B74FF9C02CBFA7EFBA3FA08" box="[997,1025,1506,1527]" italics="true" pageId="3" pageNumber="266">(b)</emphasis>
apex of penis valves; and
<emphasis id="EF8FEA8B9B74FF9C05D1FA7FFAB8FA07" box="[1279,1306,1507,1528]" italics="true" pageId="3" pageNumber="266">(c)</emphasis>
S7.
</paragraph>
</caption>
<caption id="898466119B74FF9C00ADF804FB21F851" ID-DOI="http://doi.org/10.5281/zenodo.4676940" ID-Zenodo-Dep="4676940" box="[387,1155,1944,1966]" httpUri="https://zenodo.org/record/4676940/files/figure.png" pageId="3" pageNumber="266" startId="3.[387,450,1944,1966]" targetBox="[388,1155,1571,1933]" targetPageId="3">
<paragraph id="DD4436999B74FF9C00ADF804FB21F851" blockId="3.[387,1155,1944,1993]" box="[387,1155,1944,1966]" pageId="3" pageNumber="266">
Figure 8. Male
<taxonomicName id="1AFB4D1A9B74FF9C0339F805FC82F851" authority="Batley" authorityName="Batley" authorityYear="2016" box="[535,800,1944,1966]" class="Insecta" family="Colletidae" genus="Leioproctus" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="266" phylum="Arthropoda" rank="species" species="glendae" status="sp. nov.">
<emphasis id="EF8FEA8B9B74FF9C0339F805FD79F852" box="[535,731,1944,1966]" italics="true" pageId="3" pageNumber="266">Leioproctus glendae</emphasis>
Batley
</taxonomicName>
<taxonomicNameLabel id="F4BC57F09B74FF9C0208F805FCF8F851" box="[806,858,1945,1966]" pageId="3" pageNumber="266" rank="species">n. sp.</taxonomicNameLabel>
<emphasis id="EF8FEA8B9B74FF9C0271F805FCD9F851" box="[863,891,1945,1966]" italics="true" pageId="3" pageNumber="266">(a)</emphasis>
S8; and
<emphasis id="EF8FEA8B9B74FF9C02FEF804FC4EF852" box="[976,1004,1944,1965]" italics="true" pageId="3" pageNumber="266">(b)</emphasis>
ventral view of
</paragraph>
</caption>
<paragraph id="DD4436999B74FF9C00ADF82FFD38F836" blockId="3.[387,1155,1944,1993]" box="[387,666,1971,1993]" pageId="3" pageNumber="266">metasoma; apex S8 arrowed.</paragraph>
</subSubSection>
<subSubSection id="95E165129B73FF9B0088FF0BFE03FE53" pageId="4" pageNumber="267" type="discussion">
<paragraph id="DD4436999B73FF9B0088FF0BFDB8FF4E" blockId="4.[422,538,151,177]" box="[422,538,151,177]" pageId="4" pageNumber="267">
<emphasis id="EF8FEA8B9B73FF9B0088FF0BFDB8FF4E" bold="true" box="[422,538,151,177]" pageId="4" pageNumber="267">Remarks</emphasis>
</paragraph>
<paragraph id="DD4436999B73FF9B0188FF5AFE03FE53" blockId="4.[166,795,198,428]" pageId="4" pageNumber="267">
Although the sexes were collected 2 years apart, both were found on the same flower at sites separated by about
<quantity id="1A039B7C9B73FF9B03F3FF78FCB4FF03" box="[733,790,228,252]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="267" unit="km" value="3.0">3 km</quantity>
, each time when the areas were recovering from recent fire. Given the many morphological similarities, but especially the similar mandibles there can be little doubt that they belong to the same species. During the second collection event, flowers were examined for evidence of bud damage, but none was observed.
</paragraph>
</subSubSection>
</treatment>
</document>
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