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<document id="76CC4E8B1D4F6F1AAA8A9C00860D2BDE" ID-DOI="10.12782/sd.21.1.071" ID-ISSN="2189-7301" ID-Zenodo-Dep="5737668" ID-ZooBank="8C100709-5B8B-42CB-BF9E-60B6F55F6045" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638226989121" checkinUser="felipe" docAuthor="Prokofiev, Artem M." docDate="2016" docId="554887B6E24A7923FC1643ACFC4CB478" docLanguage="en" docName="SpeciesDiversity.21.1.71-77.pdf" docOrigin="Species Diversity 21" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.7:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="7" docTitle="Anomala pygidialis Kirsch 1876" docType="treatment" docVersion="3" lastPageNumber="72" masterDocId="A971FFCEE24B7922FFA8436AFFFBB730" masterDocTitle="On the Phytophagous Scarabs of the Subfamilies Dynastinae, Rutelinae, and Melolonthinae from the Schouten Islands (Kepulauan Biak), Indonesian Papua (Coleoptera: Scarabaeidae)" masterLastPageNumber="77" masterPageNumber="71" pageNumber="72" updateTime="1699433237484" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="B9D023A2F70C9966BE35EC980D264799">On the Phytophagous Scarabs of the Subfamilies Dynastinae, Rutelinae, and Melolonthinae from the Schouten Islands (Kepulauan Biak), Indonesian Papua (Coleoptera: Scarabaeidae)</mods:title>
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<mods:namePart id="63E3A0A350780221E8B463014C26402B">Prokofiev, Artem M.</mods:namePart>
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<mods:title id="3447802FE406334E68A0AA27D8D03B33">Species Diversity</mods:title>
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<mods:date id="A6D1359DD0A5F17F6EC70F10E2B79C18">2016</mods:date>
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<paragraph id="DD5E36A0E24A7923FC1643ACFAF5B7EE" blockId="1.[958,1294,198,222]" box="[958,1294,198,222]" pageId="1" pageNumber="72">
<heading id="861681CCE24A7923FC1643ACFAF5B7EE" box="[958,1294,198,222]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="72" reason="2">
<taxonomicName id="1AE14D23E24A7923FC1643ACFAF5B7EE" authority="Kirsch, 1876" authorityName="Kirsch" authorityYear="1876" box="[958,1294,198,222]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="EF95EAB2E24A7923FC1643ACFB72B7EE" bold="true" box="[958,1161,198,222]" italics="true" pageId="1" pageNumber="72">Anomala pygidialis</emphasis>
<bibRefCitation id="B9704B51E24A7923FB2743ACFAF5B7EE" author="Kirsch, T." box="[1167,1294,198,222]" pageId="1" pageNumber="72" pagination="135 - 161" refId="ref6160" refString="Kirsch, T. 1876. Beitrag zur Kenntnis der Coleopteren-Fauna von Neu Guinea. Mitteilungen aus dem Koniglichen Zoologischen Museum zu Dresden 2: 135 - 161." type="journal article" year="1876">Kirsch, 1876</bibRefCitation>
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<paragraph id="DD5E36A0E24A7923FC9D426DFB54B62F" blockId="1.[821,1199,263,287]" box="[821,1199,263,287]" pageId="1" pageNumber="72">
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<treatmentCitation id="5C4010B1E24A7923FC9D426DFB50B62F" author="Kirsch, T." box="[821,1195,263,287]" page="141" pageId="1" pageNumber="72" year="1876">
<taxonomicName id="1AE14D23E24A7923FC9D426DFB50B62F" authority="Kirsch, 1876: 141" authorityName="Kirsch" authorityPageNumber="141" authorityYear="1876" box="[821,1195,263,287]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="EF95EAB2E24A7923FC9D426DFC0FB62F" box="[821,1012,263,287]" italics="true" pageId="1" pageNumber="72">Anomala pygidialis</emphasis>
<bibRefCitation id="B9704B51E24A7923FC52426DFB50B62F" author="Kirsch, T." box="[1018,1195,263,287]" pageId="1" pageNumber="72" pagination="135 - 161" refId="ref6160" refString="Kirsch, T. 1876. Beitrag zur Kenntnis der Coleopteren-Fauna von Neu Guinea. Mitteilungen aus dem Koniglichen Zoologischen Museum zu Dresden 2: 135 - 161." type="journal article" year="1876">Kirsch, 1876: 141</bibRefCitation>
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.
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</paragraph>
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<subSubSection id="95FB652BE24A7923FCE64222FC44B691" pageId="1" pageNumber="72" type="materials_examined">
<paragraph id="DD5E36A0E24A7923FCE64222FC44B691" blockId="1.[821,1432,328,840]" pageId="1" pageNumber="72">
<emphasis id="EF95EAB2E24A7923FCE64222FC55B650" bold="true" box="[846,942,328,352]" pageId="1" pageNumber="72">Material.</emphasis>
<materialsCitation id="6D893CFDE24A7923FC6F4222FABFB650" box="[967,1348,328,352]" country="Indonesia" location="Indonesia" pageId="1" pageNumber="72" specimenCount="4" specimenCount-female="4">
<specimenCount id="CBE7FD29E24A7923FC6F4222FBD1B650" box="[967,1066,328,352]" pageId="1" pageNumber="72" type="female">4 females</specimenCount>
,
<collectingCountry id="A5F67630E24A7923FB9E4222FB67B650" box="[1078,1180,328,352]" name="Indonesia" pageId="1" pageNumber="72">Indonesia</collectingCountry>
,
<taxonomicName id="1AE14D23E24A7923FB0F4223FABFB650" authority=", Yapen I." authorityName="Yapen I." box="[1191,1348,329,352]" class="Insecta" family="Melolonthidae" genus="Apogonia" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="papua">Papua, Yapen I</taxonomicName>
</materialsCitation>
.,
<materialsCitation id="6D893CFDE24A7923FAFD4222FA97B6B1" collectingDate="2012-11-14" country="Indonesia" location="Rosbori village" pageId="1" pageNumber="72" specimenCount="1">
northern coast,
<location id="D83E607BE24A7923FC004203FBBCB6B1" LSID="urn:lsid:plazi:treatment:554887B6E24A7923FC1643ACFC4CB478:D83E607BE24A7923FC004203FBBCB6B1" box="[936,1095,361,385]" country="Indonesia" name="Rosbori village" pageId="1" pageNumber="72">Rosbori village</location>
, at light, 13
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<collectingDate id="B91BE988E24A7923FB7E4203FAB3B6B1" box="[1238,1352,361,385]" pageId="1" pageNumber="72" value="2012-11-14">14.11.2012</collectingDate>
</date>
, A
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<materialsCitation id="6D893CFDE24A7923FAC44203FA6FB6B0" box="[1388,1428,361,385]" collectingDate="2012-11-14" country="Indonesia" isTailEnumeration="true" location="Rosbori village" pageId="1" pageNumber="72" specimenCount="1">. M</materialsCitation>
. Prokofiev leg.
</paragraph>
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<paragraph id="DD5E36A0E24A7923FCE642C0FC4CB478" blockId="1.[821,1432,328,840]" pageId="1" pageNumber="72">
<emphasis id="EF95EAB2E24A7923FCE642C0FC49B6F2" bold="true" box="[846,946,426,450]" pageId="1" pageNumber="72">Remarks.</emphasis>
This species was originally described from “Rubi [?], in the south of Geelvinksbay” and later was redescribed together with specimens from the Vogelkop Peninsula and the adjacent islands (Misool and Waigeo) (
<bibRefCitation id="B9704B51E24A7923FACA4161FC91B574" author="Zorn, C." pageId="1" pageNumber="72" pagination="25 - 71" refId="ref6396" refString="Zorn, C. 2007. Taxonomic revision of the Anomala cuprascens - species group of Sulawesi and the Papuan region: the species with unidentate protibiae (A. chlorotica - subgroup). Arthropod Systematics and Phylogeny. 65: 25 - 71." type="journal article" year="2007">Zorn 2007</bibRefCitation>
). Although I did not collected any males, my female specimens are closer to
<taxonomicName id="1AE14D23E24A7923FB834127FB5CB554" authorityName="Kirsch" authorityYear="1876" box="[1067,1191,588,612]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="EF95EAB2E24A7923FB834127FB5CB554" box="[1067,1191,588,612]" italics="true" pageId="1" pageNumber="72">A. pygidialis</emphasis>
</taxonomicName>
than to two other very similar species (
<taxonomicName id="1AE14D23E24A7923FC714107FB31B5B5" authority="Zorn, 2007" authorityName="Zorn" authorityYear="2007" box="[985,1226,621,645]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="biakensis">
<emphasis id="EF95EAB2E24A7923FC714107FBAAB5B5" box="[985,1105,621,645]" italics="true" pageId="1" pageNumber="72">A. biakensis</emphasis>
<bibRefCitation id="B9704B51E24A7923FBF04107FB31B5B5" author="Zorn, C." box="[1112,1226,621,645]" pageId="1" pageNumber="72" pagination="25 - 71" refId="ref6396" refString="Zorn, C. 2007. Taxonomic revision of the Anomala cuprascens - species group of Sulawesi and the Papuan region: the species with unidentate protibiae (A. chlorotica - subgroup). Arthropod Systematics and Phylogeny. 65: 25 - 71." type="journal article" year="2007">Zorn, 2007</bibRefCitation>
</taxonomicName>
from Biak I. and
<taxonomicName id="1AE14D23E24A7923FA294107FC03B596" authority="Zorn, 2007" authorityName="Zorn" authorityYear="2007" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="bruggei">
<emphasis id="EF95EAB2E24A7923FA294107FC84B595" italics="true" pageId="1" pageNumber="72">A. bruggei</emphasis>
<bibRefCitation id="B9704B51E24A7923FC2E41E4FC03B596" author="Zorn, C." box="[902,1016,654,678]" pageId="1" pageNumber="72" pagination="25 - 71" refId="ref6396" refString="Zorn, C. 2007. Taxonomic revision of the Anomala cuprascens - species group of Sulawesi and the Papuan region: the species with unidentate protibiae (A. chlorotica - subgroup). Arthropod Systematics and Phylogeny. 65: 25 - 71." type="journal article" year="2007">Zorn, 2007</bibRefCitation>
</taxonomicName>
from Yapen I. and the adjacent coast of New
<collectingCountry id="A5F67630E24A7923FCC241C4FC4DB5F6" box="[874,950,686,710]" name="Guinea-Bissau" pageId="1" pageNumber="72">Guinea</collectingCountry>
) in displaying no any trace of a basal tooth on the protibia and in other minor details (see
<bibRefCitation id="B9704B51E24A7923FB5041A5FA76B5D7" author="Zorn, C." box="[1272,1421,719,743]" pageId="1" pageNumber="72" pagination="25 - 71" refId="ref6396" refString="Zorn, C. 2007. Taxonomic revision of the Anomala cuprascens - species group of Sulawesi and the Papuan region: the species with unidentate protibiae (A. chlorotica - subgroup). Arthropod Systematics and Phylogeny. 65: 25 - 71." type="journal article" year="2007">Zorn 2007: 38</bibRefCitation>
).
<taxonomicName id="1AE14D23E24A7923FC9D4185FC1FB437" authorityName="Zorn" authorityYear="2007" box="[821,996,751,775]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="bruggei">
<emphasis id="EF95EAB2E24A7923FC9D4185FC1FB437" box="[821,996,751,775]" italics="true" pageId="1" pageNumber="72">Anomala bruggei</emphasis>
</taxonomicName>
co-occurs with
<taxonomicName id="1AE14D23E24A7923FB394185FAF4B437" authorityName="Kirsch" authorityYear="1876" box="[1169,1295,751,775]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="72" phylum="Arthropoda" rank="species" species="pygidialis">
<emphasis id="EF95EAB2E24A7923FB394185FAF4B437" box="[1169,1295,751,775]" italics="true" pageId="1" pageNumber="72">A. pygidialis</emphasis>
</taxonomicName>
in area near Sorong city (northwestern coast of New
<collectingCountry id="A5F67630E24A7923FB58407AFAC7B418" box="[1264,1340,784,808]" name="Guinea-Bissau" pageId="1" pageNumber="72">Guinea</collectingCountry>
) as well (
<bibRefCitation id="B9704B51E24A7923FC94405AFC57B478" author="Zorn, C." box="[828,940,816,840]" pageId="1" pageNumber="72" pagination="25 - 71" refId="ref6396" refString="Zorn, C. 2007. Taxonomic revision of the Anomala cuprascens - species group of Sulawesi and the Papuan region: the species with unidentate protibiae (A. chlorotica - subgroup). Arthropod Systematics and Phylogeny. 65: 25 - 71." type="journal article" year="2007">Zorn 2007</bibRefCitation>
).
</paragraph>
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</treatment>
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